Abstract
The two species Hilliardiella elaeagnoides and H. oligocephala were confused with each other in the past. Examination of herbarium specimens proved that they are two distinct and separate species. H. oligocephala var. besteri is newly described.
References
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<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hilliard, O.M. (1977) <em>Compositae in Natal</em>. University of Natal Press, Pietermaritzburg, 659 pp.</span></span></span></p>
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<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">JSTOR Plants science (2022) Global Plants. Available from: https://plants.jstor.org/stable/pdf/10.5555/al.ap.specimen.g00464319 (accessed March 2022); https://plants.jstor.org/stable/pdf/10.5555/al.ap.specimen.g00464306 (accessed March 2022).</span></span></span></p>
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<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Leistner, O.A. & Morris, J.W. (1976) Southern African place names. <em>Annals of the Cape Provincial Museums</em> 12. Cape Provincial Museums, Albany Museum, Grahamstown, 565 pp.</span></span></span></p>
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<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Montgomery, F.H. (1977) <em>Seeds and fruits of plants of eastern Canada and northeastern United States</em>. University of Toronto Press, Toronto, 232 pp. [based on <em>Taxon</em> 11: 145–156 (1962).] https://doi.org/10.3138/9781487583156</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pope, G.V. (1992) Tribe 3. Vernonieae Cass. <em>Flora zambesiaca</em> 6 (1): 56–189.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Robinson, H. (1999) Revisions in paleotropical Vernonieae (Asteraceae). <em>Proceedings of the Biological Society of Washington</em> 112 (1): 220–247.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Robinson, H., Skvarla, J.J. & Funk, V.A. (2016) Vernonieae (Asteraceae) of southern Africa: A generic disposition of the species and a study of their pollen<span style="color: #1f1f1f;">. </span><em>PhytoKeys</em> 60: 49–126. https://doi.org/10.3897/phytokeys.60.6734</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schultz Bipontinus, C.H. (1844) Enumeratio Compositarum a cl. Dr. Krauss annis 1838—40 in Capite bonae spei et ad portum Natalensem lectarum. <em>Flora</em> 39 oder <em>Allgemeine botanische Zeitung </em>II/27 (1): 667–682.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Stearn, W.T. (1966) <em>Botanical Latin</em>. Hafner Publishing Company, New York, 566 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Swelankomo, N., Manning, J.C. & Magee, A.R. (2016) The genus <em>Hilliardiella</em> (Asteraceae: Vernonieae) in southern Africa. <em>South African Journal of Botany</em> 106: 41–59. https://doi.org/10.1016/j.sajb.2016.05.010</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Van Wyk, A.E. & Klopper, R.R. (2021) <em>Hilliardiella oligocephala</em>, the correct name for <em>H.</em> <em>elaeagnoides</em> (Asteraceae: Vernonieae). <em>Phytotaxa</em> 513 (2): 166–168. https://doi.org/10.11646/phytotaxa.513.2.8</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Walpers, G.G. (W.G.) (1843) <em>Repertorium botanices systematicae</em>. Tomus 2, Supplementum 1. Hofmeister, Lipsiae, 1029 pp. https://doi.org/10.5962/bhl.title.7553</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wild, H. (1978) The Compositae of the Flora Zambesiaca area 8—Vernonieae (<em>Vernonia</em>). <em>Kirkia</em> 11: 31–127.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wood, J.M. (1904) <em>Natal plants</em>. Volume 4 (2). Bennett & Davis, Durban. https://doi.org/10.5962/bhl.title.18042</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Candolle, A.P., De (1836) Compositae. <em>Prodromus Systematis Naturalis Regni Vegetabilis</em> 5. Treuttel & Würtz, Paris, 706 pp. https://doi.org/10.5962/bhl.title.286</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Drège, J.F. (1843) <em>Zwei pflanzengeographische Documente, nebst einer Einleitung von Dr. E. Meyer</em>. Regensburg, Leipzig, 230 pp. https://doi.org/10.5962/bhl.title.87612 </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Glen, H.F. & Germishuizen, G. (2010) <em>Botanical exploration of southern Africa</em>. Edition 2. <em>Strelitzia</em> 26. South African National Biodiversity Institute, Pretoria, 489 pp. https://doi.org/10.5962/bhl.title.145034</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Harvey, W.H. (1865) III. <em>Vernonia</em>, Schreb. <em>In</em>: Harvey, W.H. & Sonder, O.W. (Eds.) <em>Flora capensis</em> 3. Hodges, Smith and Co., Dublin. pp. 48–53. https://doi.org/10.5962/bhl.title.821</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hilliard, O.M. (1977) <em>Compositae in Natal</em>. University of Natal Press, Pietermaritzburg, 659 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Holmgren, P.K., Holmgren, N.H. & Barnett, L.C. (1990) Index Herbariorum 1: the herbaria of the world. <em>Regnum Vegetabile</em> Ed. 8. New York Botanical Garden, New York, 704 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Jeffrey, C. & Beentje, H.J. (2000) Vernonieae. <em>Flora of Tropical East Africa</em> Compositae (Part 1): 108–285.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">JSTOR Plants science (2022) Global Plants. Available from: https://plants.jstor.org/stable/pdf/10.5555/al.ap.specimen.g00464319 (accessed March 2022); https://plants.jstor.org/stable/pdf/10.5555/al.ap.specimen.g00464306 (accessed March 2022).</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Kuntze, C.E.O. (1891) Compositae. <em>Revisio generum plantarum</em>. Vol. 1, Pars 2. Charles Klincksieck, Paris / Arthur Felix, Leipzig, 634 pp. https://doi.org/10.5962/bhl.title.327</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Leistner, O.A. & Morris, J.W. (1976) Southern African place names. <em>Annals of the Cape Provincial Museums</em> 12. Cape Provincial Museums, Albany Museum, Grahamstown, 565 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">McNeill, J. (2014) Holotype specimens and type citations: General issues. <em>Taxon</em> 63 (5): 1112–1113. https://doi.org/10.12705/635.7</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Montgomery, F.H. (1977) <em>Seeds and fruits of plants of eastern Canada and northeastern United States</em>. University of Toronto Press, Toronto, 232 pp. [based on <em>Taxon</em> 11: 145–156 (1962).] https://doi.org/10.3138/9781487583156</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pope, G.V. (1992) Tribe 3. Vernonieae Cass. <em>Flora zambesiaca</em> 6 (1): 56–189.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Robinson, H. (1999) Revisions in paleotropical Vernonieae (Asteraceae). <em>Proceedings of the Biological Society of Washington</em> 112 (1): 220–247.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Robinson, H., Skvarla, J.J. & Funk, V.A. (2016) Vernonieae (Asteraceae) of southern Africa: A generic disposition of the species and a study of their pollen<span style="color: #1f1f1f;">. </span><em>PhytoKeys</em> 60: 49–126. https://doi.org/10.3897/phytokeys.60.6734</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schultz Bipontinus, C.H. (1844) Enumeratio Compositarum a cl. Dr. Krauss annis 1838—40 in Capite bonae spei et ad portum Natalensem lectarum. <em>Flora</em> 39 oder <em>Allgemeine botanische Zeitung </em>II/27 (1): 667–682.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Stearn, W.T. (1966) <em>Botanical Latin</em>. Hafner Publishing Company, New York, 566 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Swelankomo, N., Manning, J.C. & Magee, A.R. (2016) The genus <em>Hilliardiella</em> (Asteraceae: Vernonieae) in southern Africa. <em>South African Journal of Botany</em> 106: 41–59. https://doi.org/10.1016/j.sajb.2016.05.010</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Van Wyk, A.E. & Klopper, R.R. (2021) <em>Hilliardiella oligocephala</em>, the correct name for <em>H.</em> <em>elaeagnoides</em> (Asteraceae: Vernonieae). <em>Phytotaxa</em> 513 (2): 166–168. https://doi.org/10.11646/phytotaxa.513.2.8</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Walpers, G.G. (W.G.) (1843) <em>Repertorium botanices systematicae</em>. Tomus 2, Supplementum 1. Hofmeister, Lipsiae, 1029 pp. https://doi.org/10.5962/bhl.title.7553</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wild, H. (1978) The Compositae of the Flora Zambesiaca area 8—Vernonieae (<em>Vernonia</em>). <em>Kirkia</em> 11: 31–127.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wood, J.M. (1904) <em>Natal plants</em>. Volume 4 (2). Bennett & Davis, Durban. https://doi.org/10.5962/bhl.title.18042</span></span></span></p>