Abstract
The two species Hilliardiella elaeagnoides and H. oligocephala were confused with each other in the past. Examination of herbarium specimens proved that they are two distinct and separate species. H. oligocephala var. besteri is newly described.
References
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Beentje, H. (2016) The Kew plant glossary. 2<sup>nd</sup> Edition. Royal Botanic Gardens, Kew, 184 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Candolle, A.P., De (1836) Compositae. Prodromus Systematis Naturalis Regni Vegetabilis 5. Treuttel & Würtz, Paris, 706 pp. https://doi.org/10.5962/bhl.title.286</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Drège, J.F. (1843) Zwei pflanzengeographische Documente, nebst einer Einleitung von Dr. E. Meyer. Regensburg, Leipzig, 230 pp. https://doi.org/10.5962/bhl.title.87612 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Glen, H.F. & Germishuizen, G. (2010) Botanical exploration of southern Africa. Edition 2. Strelitzia 26. South African National Biodiversity Institute, Pretoria, 489 pp. https://doi.org/10.5962/bhl.title.145034</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Harvey, W.H. (1865) III. Vernonia, Schreb. In: Harvey, W.H. & Sonder, O.W. (Eds.) Flora capensis 3. Hodges, Smith and Co., Dublin. pp. 48–53. https://doi.org/10.5962/bhl.title.821</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hilliard, O.M. (1977) Compositae in Natal. University of Natal Press, Pietermaritzburg, 659 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Holmgren, P.K., Holmgren, N.H. & Barnett, L.C. (1990) Index Herbariorum 1: the herbaria of the world. Regnum Vegetabile Ed. 8. New York Botanical Garden, New York, 704 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Jeffrey, C. & Beentje, H.J. (2000) Vernonieae. Flora of Tropical East Africa Compositae (Part 1): 108–285.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">JSTOR Plants science (2022) Global Plants. Available from: https://plants.jstor.org/stable/pdf/10.5555/al.ap.specimen.g00464319 (accessed March 2022); https://plants.jstor.org/stable/pdf/10.5555/al.ap.specimen.g00464306 (accessed March 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Kuntze, C.E.O. (1891) Compositae. Revisio generum plantarum. Vol. 1, Pars 2. Charles Klincksieck, Paris / Arthur Felix, Leipzig, 634 pp. https://doi.org/10.5962/bhl.title.327</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Leistner, O.A. & Morris, J.W. (1976) Southern African place names. Annals of the Cape Provincial Museums 12. Cape Provincial Museums, Albany Museum, Grahamstown, 565 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">McNeill, J. (2014) Holotype specimens and type citations: General issues. Taxon 63 (5): 1112–1113. https://doi.org/10.12705/635.7</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Montgomery, F.H. (1977) Seeds and fruits of plants of eastern Canada and northeastern United States. University of Toronto Press, Toronto, 232 pp. [based on Taxon 11: 145–156 (1962).] https://doi.org/10.3138/9781487583156</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pope, G.V. (1992) Tribe 3. Vernonieae Cass. Flora zambesiaca 6 (1): 56–189.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Robinson, H. (1999) Revisions in paleotropical Vernonieae (Asteraceae). Proceedings of the Biological Society of Washington 112 (1): 220–247.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Robinson, H., Skvarla, J.J. & Funk, V.A. (2016) Vernonieae (Asteraceae) of southern Africa: A generic disposition of the species and a study of their pollen<span style="color: #1f1f1f;">. </span>PhytoKeys 60: 49–126. https://doi.org/10.3897/phytokeys.60.6734</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schultz Bipontinus, C.H. (1844) Enumeratio Compositarum a cl. Dr. Krauss annis 1838—40 in Capite bonae spei et ad portum Natalensem lectarum. Flora 39 oder Allgemeine botanische Zeitung II/27 (1): 667–682.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Stearn, W.T. (1966) Botanical Latin. Hafner Publishing Company, New York, 566 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Swelankomo, N., Manning, J.C. & Magee, A.R. (2016) The genus Hilliardiella (Asteraceae: Vernonieae) in southern Africa. South African Journal of Botany 106: 41–59. https://doi.org/10.1016/j.sajb.2016.05.010</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Van Wyk, A.E. & Klopper, R.R. (2021) Hilliardiella oligocephala, the correct name for H. elaeagnoides (Asteraceae: Vernonieae). Phytotaxa 513 (2): 166–168. https://doi.org/10.11646/phytotaxa.513.2.8</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Walpers, G.G. (W.G.) (1843) Repertorium botanices systematicae. Tomus 2, Supplementum 1. Hofmeister, Lipsiae, 1029 pp. https://doi.org/10.5962/bhl.title.7553</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wild, H. (1978) The Compositae of the Flora Zambesiaca area 8—Vernonieae (Vernonia). Kirkia 11: 31–127.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wood, J.M. (1904) Natal plants. Volume 4 (2). Bennett & Davis, Durban. https://doi.org/10.5962/bhl.title.18042</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Candolle, A.P., De (1836) Compositae. Prodromus Systematis Naturalis Regni Vegetabilis 5. Treuttel & Würtz, Paris, 706 pp. https://doi.org/10.5962/bhl.title.286</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Drège, J.F. (1843) Zwei pflanzengeographische Documente, nebst einer Einleitung von Dr. E. Meyer. Regensburg, Leipzig, 230 pp. https://doi.org/10.5962/bhl.title.87612 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Glen, H.F. & Germishuizen, G. (2010) Botanical exploration of southern Africa. Edition 2. Strelitzia 26. South African National Biodiversity Institute, Pretoria, 489 pp. https://doi.org/10.5962/bhl.title.145034</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Harvey, W.H. (1865) III. Vernonia, Schreb. In: Harvey, W.H. & Sonder, O.W. (Eds.) Flora capensis 3. Hodges, Smith and Co., Dublin. pp. 48–53. https://doi.org/10.5962/bhl.title.821</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hilliard, O.M. (1977) Compositae in Natal. University of Natal Press, Pietermaritzburg, 659 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Holmgren, P.K., Holmgren, N.H. & Barnett, L.C. (1990) Index Herbariorum 1: the herbaria of the world. Regnum Vegetabile Ed. 8. New York Botanical Garden, New York, 704 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Jeffrey, C. & Beentje, H.J. (2000) Vernonieae. Flora of Tropical East Africa Compositae (Part 1): 108–285.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">JSTOR Plants science (2022) Global Plants. Available from: https://plants.jstor.org/stable/pdf/10.5555/al.ap.specimen.g00464319 (accessed March 2022); https://plants.jstor.org/stable/pdf/10.5555/al.ap.specimen.g00464306 (accessed March 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Kuntze, C.E.O. (1891) Compositae. Revisio generum plantarum. Vol. 1, Pars 2. Charles Klincksieck, Paris / Arthur Felix, Leipzig, 634 pp. https://doi.org/10.5962/bhl.title.327</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Leistner, O.A. & Morris, J.W. (1976) Southern African place names. Annals of the Cape Provincial Museums 12. Cape Provincial Museums, Albany Museum, Grahamstown, 565 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">McNeill, J. (2014) Holotype specimens and type citations: General issues. Taxon 63 (5): 1112–1113. https://doi.org/10.12705/635.7</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Montgomery, F.H. (1977) Seeds and fruits of plants of eastern Canada and northeastern United States. University of Toronto Press, Toronto, 232 pp. [based on Taxon 11: 145–156 (1962).] https://doi.org/10.3138/9781487583156</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pope, G.V. (1992) Tribe 3. Vernonieae Cass. Flora zambesiaca 6 (1): 56–189.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Robinson, H. (1999) Revisions in paleotropical Vernonieae (Asteraceae). Proceedings of the Biological Society of Washington 112 (1): 220–247.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Robinson, H., Skvarla, J.J. & Funk, V.A. (2016) Vernonieae (Asteraceae) of southern Africa: A generic disposition of the species and a study of their pollen<span style="color: #1f1f1f;">. </span>PhytoKeys 60: 49–126. https://doi.org/10.3897/phytokeys.60.6734</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schultz Bipontinus, C.H. (1844) Enumeratio Compositarum a cl. Dr. Krauss annis 1838—40 in Capite bonae spei et ad portum Natalensem lectarum. Flora 39 oder Allgemeine botanische Zeitung II/27 (1): 667–682.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Stearn, W.T. (1966) Botanical Latin. Hafner Publishing Company, New York, 566 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Swelankomo, N., Manning, J.C. & Magee, A.R. (2016) The genus Hilliardiella (Asteraceae: Vernonieae) in southern Africa. South African Journal of Botany 106: 41–59. https://doi.org/10.1016/j.sajb.2016.05.010</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Van Wyk, A.E. & Klopper, R.R. (2021) Hilliardiella oligocephala, the correct name for H. elaeagnoides (Asteraceae: Vernonieae). Phytotaxa 513 (2): 166–168. https://doi.org/10.11646/phytotaxa.513.2.8</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Walpers, G.G. (W.G.) (1843) Repertorium botanices systematicae. Tomus 2, Supplementum 1. Hofmeister, Lipsiae, 1029 pp. https://doi.org/10.5962/bhl.title.7553</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wild, H. (1978) The Compositae of the Flora Zambesiaca area 8—Vernonieae (Vernonia). Kirkia 11: 31–127.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wood, J.M. (1904) Natal plants. Volume 4 (2). Bennett & Davis, Durban. https://doi.org/10.5962/bhl.title.18042</span></span></span>
