Abstract
A new species of Cortinarius sect. Calochroi, C. pseudocisticola from European deciduous forests, is described on the basis of morphological and genetic features. Several collections of the species from the Czech Republic, Germany, Hungary and Italy were studied. Illustrations of fresh basidiomes in situ and of the main macro- and micromorphological features are provided. Its morphological characters are compared to those of the molecularly closest related and most similar species.
References
<p>Ballarà, J., Mahiques, R. & Garrido-Benavent, I. (2019) Alguns cortinaris interessants del catàleg iberoinsular (I). <em>The Journal of the J.E.C.</em> 21: 55–69.</p>
<p>Bellanger, J.M. (2015) Les cortinaires calochroïdes: une mise au point taxinomique. <em>Documents Mycologiques </em>34: 3–34.</p>
<p>Bidaud, A., Moënne-Loccoz, P. & Reumaux, P. (2001) <em>Atlas des Cortinaires</em> 11: 527–626.</p>
<p>Borovička, J., Bušek, B., Mikšík, M., Dvořák, D., Jeppesen, T.S., Dima, B., Albert, L. & Frøslev, T.G. (2015) <em>Cortinarius prodigiosus</em> – a new species of the subgenus <em>Phlegmacium</em> from Central Europe. <em>Mycological</em> <em>Progress</em> 14: 29. https://doi.org/10.1007/s11557-015-1051-1</p>
<p>Borovička, J., Braeuer, S., Žigová, A., Gryndler, M., Dima, B., Goessler, W., Frøslev, T.G., Kameník, J. & Kärcher, R. (2017) Resurrection of <em>Cortinarius</em> <em>coalescens</em>: taxonomy, chemistry, and ecology. <em>Mycological Progress</em> 16 (9): 927–939. https://doi.org/10.1007/s11557-017-1331-z</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (1989) <em>Cortinarius: Flora Photographica. </em>Vol. I (Swedish version).<em> Cortinarius</em> HB, Sweden.</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (1992) <em>Cortinarius: Flora Photographica. </em>Vol. II (Swedish version). <em>Cortinarius </em>HB, Sweden.</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (1994) <em>Cortinarius: Flora Photographica. </em>Vol. III (Swedish version). <em>Cortinarius</em> HB, Sweden.</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (1998) <em>Cortinarius: Flora Photographica. </em>Vol. IV (Swedish version). <em>Cortinarius</em> HB, Sweden.</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (2012) <em>Cortinarius:</em> <em>Flora Photographica. </em>Vol. V (Swedish version). <em>Cortinarius</em> HB, Sweden.</p>
<p>Brandrud, T.E., Schmidt-Stohn, G. & Dima, B. (2019) <em>Cortinarius</em> <em>hildegardiae</em> and <em>C. mariekristinae</em> spp. nov., two new species in the phlegmacioid clade Humolentes (sect. <em>Calochroi</em> s. l.). <em>Sydowia</em> 71: 115–127.</p>
<p>Castresana, J. (2000) Selection of conserved blocks from multiple alignments for their use in phylogenetic analysis. <em>Molecular Biology and Evolution </em>17 (4): 540–552. https://doi.org/10.1093/oxfordjournals.molbev.a026334</p>
<p>Chevassut, G. & Henry, R. (1975) Six cortinaires méditerranéens nouveaux du chêne vert dans le Bas-Languedoc. <em>Documents Mycologiques</em> 5 (20): 23–36.</p>
<p>Clericuzio, M., Dovana, F., Bellanger, J.M., Brandrud, T.E., Dima, B., Frøslev, T.G., Boccardo, F., Jeppesen, T.S. & Vizzini, A. (2017) <em>Cortinarius parasuaveolens</em> (= <em>C. pseudogracilior</em>): new data and a synonym of a very poorly known species of section <em>Calochroi</em>. <em>Sydowia</em> 69: 215–228.</p>
<p>Dovana, F., Contu, M., Angeli, P., Brandi, A. & Mucciarelli, M. (2017) <em>Leucoagaricus ariminensis sp. nov.</em>, a lilac species from Italy. <em>Mycotaxon </em>132: 205–216.</p>
<p>Dovana, F., Boccardo, F., Clericuzio, M. & Vizzini, A. (2020) <em>Cortinarius lentus </em>(Agaricales, Cortinariaceae)<em>, </em>a new species in section <em>Calochroi. Phytotaxa </em>447 (1): 31–41. https://doi.org/10.11646/phytotaxa.447.1.3</p>
<p>Doyle, J.J. & Doyle, J.L. (1987) A rapid DNA isolation procedure for small quantities of fresh leaf material. <em>Phytochemical Bulletin </em>19: 11–15.</p>
<p>Felsenstein, J. (1985) Confidence limits on phylogenies: an approach using the bootstrap. <em>Evolution</em> 39: 783–791. https://doi.org/10.1111/j.1558-5646.1985.tb00420.x</p>
<p>Fries, E. (1838) <em>Epicrisis Systematis Mycologici.</em> Typographia Academica, 610 pp.</p>
<p>Frøslev, T.G., Jeppesen, T.S. & Brandrud, T.E. (2006a) New specs and combinations in <em>Cortinarius </em>subgenus <em>Phlegmacium </em>section <em>Calochroi</em>. <em>Mycotaxon</em> 97: 367–377.</p>
<p>Frøslev, T.G., Jeppesen, T.S. & Læssøe, T. (2006b) Seven new calochroid and fulvoid species of <em>Cortinarius</em>. <em>Mycological Research </em>110: 1148–1160. https://doi.org/10.1016/j.mycres.2006.05.012</p>
<p>Frøslev, T.G., Jeppesen, T.S., Læssøe, T. & Kjøller, R. (2007) Molecular phylogenetics and delimitation of species in <em>Cortinarius </em>section <em>Calochroi </em>(Basidiomycota, Agaricales) in Europe. <em>Molecular Phylogenetics and Evolution </em>44: 217– 227. https://doi.org/10.1016/j.ympev.2006.11.013</p>
<p>Frøslev, T.G., Brandrud, T.E. & Dima, B. (2017) <em>Cortinarius stjernegaardii</em> and <em>C. kristinae</em> (Basidiomycota, Agaricales), two new European species with a mainly northern distribution. <em>Mycological Progress</em> 16: 145–153. https://doi.org/10.1007/s11557-016-1264-y</p>
<p>Gardes, M. & Bruns, T.D. (1993) ITS primers with enhanced specificity for basidiomycetes application to the identification of mycorrhizae and rusts. <em>Molecular Ecology </em>2: 113–118. https://doi.org/10.1111/j.1365-294X.1993.tb00005.x</p>
<p>Garnica, S., Weiß, M., Oertel, B., Ammirati, J. & Oberwinkler, F. (2009) Phylogenetic relationships in <em>Cortinarius</em>, section <em>Calochroi</em>, inferred from nuclear DNA sequences. <em>BMC Evolutionary Biology </em>9 (1): 1. https://doi.org/10.1186/1471-2148-9-1</p>
<p>Garnica, S., Schön, M.E., Abarenkov, K., Riess, K., Liimatainen, K., Niskanen, T., Dima, B., Soop, K., Frøslev, T.G., Jeppesen, T.S., Peintner, U., Kuhnert-Finkernagel, R., Brandrud, T.E., Saar, G., Oertel, B. & Ammirati, J.F. (2016) Determining threshold values for barcoding fungi: lessons from <em>Cortinarius </em>(Basidiomycota), a highly diverse and widespread ectomycorrhizal genus. <em>FEMS Microbiology Ecology </em>92 (4): fiw045. https://doi.org/10.1093/femsec/fiw045</p>
<p>Gray, S.F. (1821) <em>A natural arrangement of British plants</em>. United Kingdom, London: Baldwin, Cradock and Joy, 824 pp.</p>
<p>He, M.Q., Zhao, R.L., Hyde, K.D., Begerow, D., Kemler, M., Yurkov, A., McKenzie, E.H.C., Raspé, O., Kakishima, M., Sánchez-Ramírez, S., Vellinga, E.C., Halling, R., Papp, V., Zmitrovich, I.V., Buyck, B., Ertz, D., Wijayawardene, N.N., Cui, B.K., Schoutteten, N., Liu, X.Z., Li, T.H., Yao, Y.J., Zhu, X.Y., Liu, A.Q., Li, G.J., Zhang, M.Z., Ling, Z.L., Cao, B., Antonín, V., Boekhout, T., Da Silva, B.D.B., De Crop, E., Decock, C., Dima, B., Dutta, A.K., Fell, J.W., Geml, J., Ghobad-Nejhad, M., Giachini, A.J., Gibertoni, T.B., Gorjón, S.P., Haelewaters, D., He, S.H., Hodkinson, B.P., Horak, E., Hoshino, T., Justo, A., Lim, Y.W., Menolli Jr, N., Mešić, A., Moncalvo, J.M., Mueller, G.M., Nagy, L.G., Nilsson, R.H., Noordeloos, M., Nuytinck, J., Orihara, T., Ratchadawan, C., Rajchenberg, M., Silva-Filho, A.G.S., Sulzbacher, M.A., Tkalčec, Z., Valenzuela, R., Verbeken, A., Vizzini, A., Wartchow, F., Wei, T.Z., Weiß, M., Zhao, C.L. & Kirk, P.M. (2019) Notes, outline and divergence times of Basidiomycota<strong>. </strong><em>Fungal Diversity</em> 99: 105–367. https://doi.org/10.1007/s13225-019-00435-4</p>
<p>Henry, R. (1952) Les Scauri. <em>Bulletin de la Société Mycologique de France</em> 67 (3): 225–322.</p>
<p>Kalichman, J., Kirk, P.M. & Matheny, P.B. (2020) A compendium of generic names of agarics and Agaricales. <em>Taxon</em> 69 (3): 425–447. https://doi.org/10.1002/tax.12240</p>
<p>Katoh, K. & Toh, H. (2008) Recent developments in the MAFFT multiple sequence alignment program. <em>Briefings in Bioinformatics </em>9: 286–298. https://doi.org/10.1093/bib/bbn013</p>
<p>Kearse, M., Moir, R., Wilson, A., Stones-Havas, S., Cheung, M., Sturrock, S., Buxton, S., Cooper, A., Markowitz, S., Duran, C., Thierer, T., Ashton, B., Meintjes, P. & Drummond, A. (2012) Geneious Basic: an integrated and extendable desktop software platform for the organisation and analysis of sequence data. <em>Bioinformatics </em>28: 1647–1649. https://doi.org/10.1093/bioinformatics/bts199</p>
<p>Kirk, P.M., Cannon, P.F., Minter, D.W. & Stalpers, J.A. (2008) <em>Ainsworth & Bisby’s dictionary of the fungi. 10th Edition</em>. CABI, Europe, UK, 492 pp. https://doi.org/10.1079/9780851998268.0000</p>
<p>Lanfear, R., Frandsen, P.B., Wright, A.M., Senfeld, T. & Calcott, B. (2017) PartitionFinder 2: new methods for selecting partitioned models of evolution for molecular and morphological hylogenetic analyses. <em>Molecular Biology and Evolution </em>34: 772–773. https://doi.org/10.1093/molbev/msw260</p>
<p>Liimatainen, K., Niskanen, T., Dima, B., Kytövuori, I., Ammirati, J.F. & Frøslev, T.G. (2014) The largest type study of Agaricales species to date: bringing identification and nomenclature of <em>Phlegmacium </em>(<em>Cortinarius</em>) into the DNA era. <em>Persoonia </em>33: 98–140. https://doi.org/10.3767/003158514X684681</p>
<p>Mahiques, R., Ballarà, J., Salom, J.C., Bellanger, J.M. & Garrido-Benavent, I. (2018) Morphogenetic diversity of the ectomycorrhizal genus <em>Cortinarius </em>section <em>Calochroi </em>in the Iberian Peninsula. <em>Mycological Progress </em>17: 815–831. https://doi.org/10.1007/s11557-018-1394-5</p>
<p>Melot, J. (1989) Novitates - Combinationes et taxa nova. <em>Documents Mycologiques</em> 20 (77): 93–100.</p>
<p>Miller, M.A., Pfeiffer, W. & Schwartz, T. (2010) Creating the CIPRES Science Gateway for inference of large phylogenetic trees. <em>In: Proceedings of the Gateway Computing Environments Workshop (GCE)</em>. 14 Nov. 2010, New Orleans, LA, pp. 1–8. https://doi.org/10.1109/GCE.2010.5676129</p>
<p>Moser, M. (1960) <em>Die Gattung Phlegmacium (Sleimköpfe) Die Pilze Mitteleuropas</em>, Band IV. Julius Klinkhardt, Bad Heilbrunn. 441 pp.</p>
<p>Moser, M. (1967) <em>Die Röhrlinge und Blätterpilze in Gams, W.: Kleine Kryptogamenflora</em>. Band II, b/2. 3 Aufl. G. Fischer. Stuttgart. 443 pp.</p>
<p>Moser, M. & Horak, E. (1975) <em>Cortinarius </em>Fr. und nahe verwandte Gattungen in Südamerika. <em>Nova hedwigia</em>. Beiefte 51–52. pp. 464–465.</p>
<p>Muñoz Sánchez, J.A. (2020) “<em>Cortinarius cisticola</em> Frøslev & T.S. Jeppesen” Available from: https://micologica-barakaldo.org/wp-content/uploads/2020/12/Cortinarius-cisticola.pdf (accessed 30 August 2021)</p>
<p>Munsell, C. (1994) <em>Soil, color charts</em>, revised edition. Macbeth Division of Kollmorgen Instruments Corporation, New York (USA).</p>
<p>Ortega, A., Suárez-Santiago, V.N. & Reyes, J.D. (2008) Morphological and ITS identification of <em>Cortinarius </em>species (section <em>Calochroi</em>) collected in Mediterranean <em>Quercus </em>woodlands. <em>Fungal Diversity </em>29: 73–88.</p>
<p>Papp, V. & Dima, B. (2018) New systematic position of <em>Aurantiporus</em> <em>alborubescens</em> (Meruliaceae, Basidiomycota), a threatened old-growth forest polypore. <em>Mycological</em> <em>Progress</em> 17: 319–332. https://doi.org/10.1007/s11557-017-1356-3</p>
<p>Persoon, C<em>.</em>H. (1801) <em>Synopsis methodica fungorum.</em> Pars prima & secunda. Goetingae.</p>
<p>Pouzar, Z. (1983) Taxonomic and nomenclatural notes on some families of larger fungi. <em>Česká Mykologie </em>37 (3): 172–176.</p>
<p>Rambaut, A. & Drummond, A.J. (2009) <em>Tracer version 1.5</em>. Available from: http://tree.bio.ed.ac.uk/software/tracer/ (accessed 17 February 2021)</p>
<p>Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. <em>Systematic Biology </em>61: 539–542. https://doi.org/10.1093/sysbio/sys029</p>
<p>Soop, K., Dima, B., Cooper, J.A., Duckchul, P. & Oertel, B. (2019) A phylogenetic approach to a global supraspecific taxonomy of <em>Cortinarius</em> (Agaricales) with an emphasis on the southern mycota. <em>Persoonia</em> 42: 261–290. https://doi.org/10.3767/persoonia.2019.42.10</p>
<p>Stamatakis, A. (2006) RAxML-VI-HPC: maximum likelihood-based phylogenetic analyses with thousands of taxa and mixed models. <em>Bioinformatics </em>22: 2688–2690. https://doi.org/10.1093/bioinformatics/btl446</p>
<p>Vizzini, A., Clericuzio, M., Boccardo, F. & Ercole, E. (2012) A new <em>Cortinarius </em>of section <em>Calochroi </em>(Basidiomycota, Agaricomycetes) from Mediterranean <em>Quercus </em>woodlands (Italy). <em>Mycologia </em>104 (6): 1502–1509. https://doi.org/10.3852/12-072</p>
<p>White, T.J., Bruns, T., Lee, S.S. & Taylor, J. (1990) Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. <em>In</em>: Innis, M.A., Gelfand, D.H., Sninsky, J.J. & White, T.J. (Eds.) <em>PCR Protocols: A Guide to Methods and Applications. </em>Academic Press, New York, pp. 315–322. https://doi.org/10.1016/B978-0-12-372180-8.50042-1</p>
<p>Bellanger, J.M. (2015) Les cortinaires calochroïdes: une mise au point taxinomique. <em>Documents Mycologiques </em>34: 3–34.</p>
<p>Bidaud, A., Moënne-Loccoz, P. & Reumaux, P. (2001) <em>Atlas des Cortinaires</em> 11: 527–626.</p>
<p>Borovička, J., Bušek, B., Mikšík, M., Dvořák, D., Jeppesen, T.S., Dima, B., Albert, L. & Frøslev, T.G. (2015) <em>Cortinarius prodigiosus</em> – a new species of the subgenus <em>Phlegmacium</em> from Central Europe. <em>Mycological</em> <em>Progress</em> 14: 29. https://doi.org/10.1007/s11557-015-1051-1</p>
<p>Borovička, J., Braeuer, S., Žigová, A., Gryndler, M., Dima, B., Goessler, W., Frøslev, T.G., Kameník, J. & Kärcher, R. (2017) Resurrection of <em>Cortinarius</em> <em>coalescens</em>: taxonomy, chemistry, and ecology. <em>Mycological Progress</em> 16 (9): 927–939. https://doi.org/10.1007/s11557-017-1331-z</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (1989) <em>Cortinarius: Flora Photographica. </em>Vol. I (Swedish version).<em> Cortinarius</em> HB, Sweden.</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (1992) <em>Cortinarius: Flora Photographica. </em>Vol. II (Swedish version). <em>Cortinarius </em>HB, Sweden.</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (1994) <em>Cortinarius: Flora Photographica. </em>Vol. III (Swedish version). <em>Cortinarius</em> HB, Sweden.</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (1998) <em>Cortinarius: Flora Photographica. </em>Vol. IV (Swedish version). <em>Cortinarius</em> HB, Sweden.</p>
<p>Brandrud, T.E., Lindström, H., Marklund, H., Melot, J. & Muskos, S. (2012) <em>Cortinarius:</em> <em>Flora Photographica. </em>Vol. V (Swedish version). <em>Cortinarius</em> HB, Sweden.</p>
<p>Brandrud, T.E., Schmidt-Stohn, G. & Dima, B. (2019) <em>Cortinarius</em> <em>hildegardiae</em> and <em>C. mariekristinae</em> spp. nov., two new species in the phlegmacioid clade Humolentes (sect. <em>Calochroi</em> s. l.). <em>Sydowia</em> 71: 115–127.</p>
<p>Castresana, J. (2000) Selection of conserved blocks from multiple alignments for their use in phylogenetic analysis. <em>Molecular Biology and Evolution </em>17 (4): 540–552. https://doi.org/10.1093/oxfordjournals.molbev.a026334</p>
<p>Chevassut, G. & Henry, R. (1975) Six cortinaires méditerranéens nouveaux du chêne vert dans le Bas-Languedoc. <em>Documents Mycologiques</em> 5 (20): 23–36.</p>
<p>Clericuzio, M., Dovana, F., Bellanger, J.M., Brandrud, T.E., Dima, B., Frøslev, T.G., Boccardo, F., Jeppesen, T.S. & Vizzini, A. (2017) <em>Cortinarius parasuaveolens</em> (= <em>C. pseudogracilior</em>): new data and a synonym of a very poorly known species of section <em>Calochroi</em>. <em>Sydowia</em> 69: 215–228.</p>
<p>Dovana, F., Contu, M., Angeli, P., Brandi, A. & Mucciarelli, M. (2017) <em>Leucoagaricus ariminensis sp. nov.</em>, a lilac species from Italy. <em>Mycotaxon </em>132: 205–216.</p>
<p>Dovana, F., Boccardo, F., Clericuzio, M. & Vizzini, A. (2020) <em>Cortinarius lentus </em>(Agaricales, Cortinariaceae)<em>, </em>a new species in section <em>Calochroi. Phytotaxa </em>447 (1): 31–41. https://doi.org/10.11646/phytotaxa.447.1.3</p>
<p>Doyle, J.J. & Doyle, J.L. (1987) A rapid DNA isolation procedure for small quantities of fresh leaf material. <em>Phytochemical Bulletin </em>19: 11–15.</p>
<p>Felsenstein, J. (1985) Confidence limits on phylogenies: an approach using the bootstrap. <em>Evolution</em> 39: 783–791. https://doi.org/10.1111/j.1558-5646.1985.tb00420.x</p>
<p>Fries, E. (1838) <em>Epicrisis Systematis Mycologici.</em> Typographia Academica, 610 pp.</p>
<p>Frøslev, T.G., Jeppesen, T.S. & Brandrud, T.E. (2006a) New specs and combinations in <em>Cortinarius </em>subgenus <em>Phlegmacium </em>section <em>Calochroi</em>. <em>Mycotaxon</em> 97: 367–377.</p>
<p>Frøslev, T.G., Jeppesen, T.S. & Læssøe, T. (2006b) Seven new calochroid and fulvoid species of <em>Cortinarius</em>. <em>Mycological Research </em>110: 1148–1160. https://doi.org/10.1016/j.mycres.2006.05.012</p>
<p>Frøslev, T.G., Jeppesen, T.S., Læssøe, T. & Kjøller, R. (2007) Molecular phylogenetics and delimitation of species in <em>Cortinarius </em>section <em>Calochroi </em>(Basidiomycota, Agaricales) in Europe. <em>Molecular Phylogenetics and Evolution </em>44: 217– 227. https://doi.org/10.1016/j.ympev.2006.11.013</p>
<p>Frøslev, T.G., Brandrud, T.E. & Dima, B. (2017) <em>Cortinarius stjernegaardii</em> and <em>C. kristinae</em> (Basidiomycota, Agaricales), two new European species with a mainly northern distribution. <em>Mycological Progress</em> 16: 145–153. https://doi.org/10.1007/s11557-016-1264-y</p>
<p>Gardes, M. & Bruns, T.D. (1993) ITS primers with enhanced specificity for basidiomycetes application to the identification of mycorrhizae and rusts. <em>Molecular Ecology </em>2: 113–118. https://doi.org/10.1111/j.1365-294X.1993.tb00005.x</p>
<p>Garnica, S., Weiß, M., Oertel, B., Ammirati, J. & Oberwinkler, F. (2009) Phylogenetic relationships in <em>Cortinarius</em>, section <em>Calochroi</em>, inferred from nuclear DNA sequences. <em>BMC Evolutionary Biology </em>9 (1): 1. https://doi.org/10.1186/1471-2148-9-1</p>
<p>Garnica, S., Schön, M.E., Abarenkov, K., Riess, K., Liimatainen, K., Niskanen, T., Dima, B., Soop, K., Frøslev, T.G., Jeppesen, T.S., Peintner, U., Kuhnert-Finkernagel, R., Brandrud, T.E., Saar, G., Oertel, B. & Ammirati, J.F. (2016) Determining threshold values for barcoding fungi: lessons from <em>Cortinarius </em>(Basidiomycota), a highly diverse and widespread ectomycorrhizal genus. <em>FEMS Microbiology Ecology </em>92 (4): fiw045. https://doi.org/10.1093/femsec/fiw045</p>
<p>Gray, S.F. (1821) <em>A natural arrangement of British plants</em>. United Kingdom, London: Baldwin, Cradock and Joy, 824 pp.</p>
<p>He, M.Q., Zhao, R.L., Hyde, K.D., Begerow, D., Kemler, M., Yurkov, A., McKenzie, E.H.C., Raspé, O., Kakishima, M., Sánchez-Ramírez, S., Vellinga, E.C., Halling, R., Papp, V., Zmitrovich, I.V., Buyck, B., Ertz, D., Wijayawardene, N.N., Cui, B.K., Schoutteten, N., Liu, X.Z., Li, T.H., Yao, Y.J., Zhu, X.Y., Liu, A.Q., Li, G.J., Zhang, M.Z., Ling, Z.L., Cao, B., Antonín, V., Boekhout, T., Da Silva, B.D.B., De Crop, E., Decock, C., Dima, B., Dutta, A.K., Fell, J.W., Geml, J., Ghobad-Nejhad, M., Giachini, A.J., Gibertoni, T.B., Gorjón, S.P., Haelewaters, D., He, S.H., Hodkinson, B.P., Horak, E., Hoshino, T., Justo, A., Lim, Y.W., Menolli Jr, N., Mešić, A., Moncalvo, J.M., Mueller, G.M., Nagy, L.G., Nilsson, R.H., Noordeloos, M., Nuytinck, J., Orihara, T., Ratchadawan, C., Rajchenberg, M., Silva-Filho, A.G.S., Sulzbacher, M.A., Tkalčec, Z., Valenzuela, R., Verbeken, A., Vizzini, A., Wartchow, F., Wei, T.Z., Weiß, M., Zhao, C.L. & Kirk, P.M. (2019) Notes, outline and divergence times of Basidiomycota<strong>. </strong><em>Fungal Diversity</em> 99: 105–367. https://doi.org/10.1007/s13225-019-00435-4</p>
<p>Henry, R. (1952) Les Scauri. <em>Bulletin de la Société Mycologique de France</em> 67 (3): 225–322.</p>
<p>Kalichman, J., Kirk, P.M. & Matheny, P.B. (2020) A compendium of generic names of agarics and Agaricales. <em>Taxon</em> 69 (3): 425–447. https://doi.org/10.1002/tax.12240</p>
<p>Katoh, K. & Toh, H. (2008) Recent developments in the MAFFT multiple sequence alignment program. <em>Briefings in Bioinformatics </em>9: 286–298. https://doi.org/10.1093/bib/bbn013</p>
<p>Kearse, M., Moir, R., Wilson, A., Stones-Havas, S., Cheung, M., Sturrock, S., Buxton, S., Cooper, A., Markowitz, S., Duran, C., Thierer, T., Ashton, B., Meintjes, P. & Drummond, A. (2012) Geneious Basic: an integrated and extendable desktop software platform for the organisation and analysis of sequence data. <em>Bioinformatics </em>28: 1647–1649. https://doi.org/10.1093/bioinformatics/bts199</p>
<p>Kirk, P.M., Cannon, P.F., Minter, D.W. & Stalpers, J.A. (2008) <em>Ainsworth & Bisby’s dictionary of the fungi. 10th Edition</em>. CABI, Europe, UK, 492 pp. https://doi.org/10.1079/9780851998268.0000</p>
<p>Lanfear, R., Frandsen, P.B., Wright, A.M., Senfeld, T. & Calcott, B. (2017) PartitionFinder 2: new methods for selecting partitioned models of evolution for molecular and morphological hylogenetic analyses. <em>Molecular Biology and Evolution </em>34: 772–773. https://doi.org/10.1093/molbev/msw260</p>
<p>Liimatainen, K., Niskanen, T., Dima, B., Kytövuori, I., Ammirati, J.F. & Frøslev, T.G. (2014) The largest type study of Agaricales species to date: bringing identification and nomenclature of <em>Phlegmacium </em>(<em>Cortinarius</em>) into the DNA era. <em>Persoonia </em>33: 98–140. https://doi.org/10.3767/003158514X684681</p>
<p>Mahiques, R., Ballarà, J., Salom, J.C., Bellanger, J.M. & Garrido-Benavent, I. (2018) Morphogenetic diversity of the ectomycorrhizal genus <em>Cortinarius </em>section <em>Calochroi </em>in the Iberian Peninsula. <em>Mycological Progress </em>17: 815–831. https://doi.org/10.1007/s11557-018-1394-5</p>
<p>Melot, J. (1989) Novitates - Combinationes et taxa nova. <em>Documents Mycologiques</em> 20 (77): 93–100.</p>
<p>Miller, M.A., Pfeiffer, W. & Schwartz, T. (2010) Creating the CIPRES Science Gateway for inference of large phylogenetic trees. <em>In: Proceedings of the Gateway Computing Environments Workshop (GCE)</em>. 14 Nov. 2010, New Orleans, LA, pp. 1–8. https://doi.org/10.1109/GCE.2010.5676129</p>
<p>Moser, M. (1960) <em>Die Gattung Phlegmacium (Sleimköpfe) Die Pilze Mitteleuropas</em>, Band IV. Julius Klinkhardt, Bad Heilbrunn. 441 pp.</p>
<p>Moser, M. (1967) <em>Die Röhrlinge und Blätterpilze in Gams, W.: Kleine Kryptogamenflora</em>. Band II, b/2. 3 Aufl. G. Fischer. Stuttgart. 443 pp.</p>
<p>Moser, M. & Horak, E. (1975) <em>Cortinarius </em>Fr. und nahe verwandte Gattungen in Südamerika. <em>Nova hedwigia</em>. Beiefte 51–52. pp. 464–465.</p>
<p>Muñoz Sánchez, J.A. (2020) “<em>Cortinarius cisticola</em> Frøslev & T.S. Jeppesen” Available from: https://micologica-barakaldo.org/wp-content/uploads/2020/12/Cortinarius-cisticola.pdf (accessed 30 August 2021)</p>
<p>Munsell, C. (1994) <em>Soil, color charts</em>, revised edition. Macbeth Division of Kollmorgen Instruments Corporation, New York (USA).</p>
<p>Ortega, A., Suárez-Santiago, V.N. & Reyes, J.D. (2008) Morphological and ITS identification of <em>Cortinarius </em>species (section <em>Calochroi</em>) collected in Mediterranean <em>Quercus </em>woodlands. <em>Fungal Diversity </em>29: 73–88.</p>
<p>Papp, V. & Dima, B. (2018) New systematic position of <em>Aurantiporus</em> <em>alborubescens</em> (Meruliaceae, Basidiomycota), a threatened old-growth forest polypore. <em>Mycological</em> <em>Progress</em> 17: 319–332. https://doi.org/10.1007/s11557-017-1356-3</p>
<p>Persoon, C<em>.</em>H. (1801) <em>Synopsis methodica fungorum.</em> Pars prima & secunda. Goetingae.</p>
<p>Pouzar, Z. (1983) Taxonomic and nomenclatural notes on some families of larger fungi. <em>Česká Mykologie </em>37 (3): 172–176.</p>
<p>Rambaut, A. & Drummond, A.J. (2009) <em>Tracer version 1.5</em>. Available from: http://tree.bio.ed.ac.uk/software/tracer/ (accessed 17 February 2021)</p>
<p>Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. <em>Systematic Biology </em>61: 539–542. https://doi.org/10.1093/sysbio/sys029</p>
<p>Soop, K., Dima, B., Cooper, J.A., Duckchul, P. & Oertel, B. (2019) A phylogenetic approach to a global supraspecific taxonomy of <em>Cortinarius</em> (Agaricales) with an emphasis on the southern mycota. <em>Persoonia</em> 42: 261–290. https://doi.org/10.3767/persoonia.2019.42.10</p>
<p>Stamatakis, A. (2006) RAxML-VI-HPC: maximum likelihood-based phylogenetic analyses with thousands of taxa and mixed models. <em>Bioinformatics </em>22: 2688–2690. https://doi.org/10.1093/bioinformatics/btl446</p>
<p>Vizzini, A., Clericuzio, M., Boccardo, F. & Ercole, E. (2012) A new <em>Cortinarius </em>of section <em>Calochroi </em>(Basidiomycota, Agaricomycetes) from Mediterranean <em>Quercus </em>woodlands (Italy). <em>Mycologia </em>104 (6): 1502–1509. https://doi.org/10.3852/12-072</p>
<p>White, T.J., Bruns, T., Lee, S.S. & Taylor, J. (1990) Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. <em>In</em>: Innis, M.A., Gelfand, D.H., Sninsky, J.J. & White, T.J. (Eds.) <em>PCR Protocols: A Guide to Methods and Applications. </em>Academic Press, New York, pp. 315–322. https://doi.org/10.1016/B978-0-12-372180-8.50042-1</p>