QIN, XIAO-WEI, Spice and Beverage Research Institute, CATAS, Wanning 571533, Hainan, China, Ministry of Agriculture Key Laboratory of Genetic Resources Utilization of Spice and Beverage Crops, Wanning 571533, Hainan, China, Hainan Provincial Key Laboratory of Genetic Improvement and Quality Regulation for Tropical Spice and Beverage Crops, Wanning 571533, Hainan, China, China