EL MOKNI, RIDHA, University of Carthage, Laboratory of Botany and Plant Ecology (SNA-214), Department of Life Sciences, Faculty of Sciences of Bizerte, Jarzouna, 7021, Bizerte, Tunisia. University of Jendouba, Laboratory of Silvo-Pastoral Resources, Silvo-Pastoral Institute of Tabarka, BP. 345, 8110-Tabarka, Tunisia. University of Monastir, Laboratory of Botany and Plant Biology, Faculty of Pharmacy of Monastir, Avenue Avicenna, 5000-Monastir, Tunisia., Tunisia