HAN, LI-HONG, Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, Yunnan, China. Key Laboratory of Yunnan Province Universities of the Diversity and Ecological Adaptive Evolution for Animals and Plants on Yungui Plateau, Qujing Normal University, Qujing 655011, Yunnan, China University of Chinese Academy of Sciences, Beijing 100039, China, China