Skip to main content Skip to main navigation menu Skip to site footer
Responsive image
Issue: Vol. 658 No. 1: 2 Jul. 2024
Type: Article
Published: 2024-07-02
DOI: 10.11646/phytotaxa.658.1.1
Page range: 1-49
Abstract views: 134
PDF downloaded: 4

New orchids in the flora of Vietnam VIII (Orchidaceae: Epidendroideae, tribes Malaxideae, Neottieae and Podochileae)

Komarov Botanical Institute of the Russian Academy of Sciences, 2, Prof. Popov Street, 197022, St. Petersburg, Russia
##plugins.generic.jatsParser.article.authorBio##
×

LEONID V. AVERYANOV

 

 

Institute of Applied Technology, Thu Dau Mot University, No. 6, Tran Van On Street, Phu Hoa Ward, Thu Dau Mot City, Binh Duong Province, Vietnam
Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85, Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam
Department of Higher Plants, Biological Faculty, M.V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234, Moscow, Russia. Joint Russian-Vietnamese Tropical Scientific and Technological Center, 63, Nguyen Van Huyen Street, Cau Giay District, 122000, Hanoi, Vietnam.
Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoang Quoc Viet Road, Cau Giay District, Hanoi, Vietnam. Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoang Quoc Viet Road, Cau Giay District, Hanoi, Vietnam.
Komarov Botanical Institute of the Russian Academy of Sciences, 2, Prof. Popov Street, 197022, St. Petersburg, Russia
Komarov Botanical Institute of the Russian Academy of Sciences, 2, Prof. Popov Street, 197022, St. Petersburg, Russia
Komarov Botanical Institute of the Russian Academy of Sciences, 2, Prof. Popov Street, 197022, St. Petersburg, Russia
Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam
Tay Nguyen Institute for Scientific Research, Vietnam Academy of Science and Technology, Da Lat, Vietnam
endemism nature conservation new national records new species nomenclatural combinations plant diversity plant taxonomy taxonomical history Monocots

Abstract

The paper presents new data on the orchid diversity of Vietnam obtained mainly in the course of the fieldwork in 2020–2023, as well as from the investigation of the collections stored at the Herbaria of Russia and Vietnam. It includes descriptions of four species (Appendicula spicata, Dendrobium khanhii, D. vietii, Listera anthropophora) and one variety (Eria dacrydium var. cornuta) new to science, as well as reports of 18 species new to the flora of Vietnam (Bulbophyllum conchiferum, B. gibbosum, B. lasiochilum, B. malleolabrum, B. medusae, B. mucronatum, B. reclusum, B. siamense, B. sutepense, Dendrobium deltatum, D. densiflorum, D. lampongense, D. polyanthum, Oberonia maxima, Phreatia sulcata, Podochilus lucescens, Sunipia grandiflora and S. rimannii). In addition, a new name, Dendrobium nobile var. luteoalbum, and a new nomenclatural combination, Eria chienii, are proposed, and lectotypes for the names Bulbophyllum mucronatum, B. sutepense, Dendrobium deltatum and D. nobile var. luteoalbum are designated. For each of the studied species, relevant synonyms, studied specimens, ecology, phenology, distribution, conservation status assessment according to the IUCN Red List Categories and Criteria, brief taxonomic notes, and illustrations are provided. For the newly described taxa, morphological descriptions and notes on etymology are included.

References

  1. Aiton, W.T. (1813) Hortus Kewensis, or, A catalogue of the plants cultivated in the Royal Botanic Garden at Kew. Vol. 5. Longman, London, 568 pp. https://doi.org/10.5962/bhl.title.105339
  2. Ansari, R. & Balakrishnan, N.P. (1990) A revision of the Indian species of Oberonia (Orchidaceae). Orchid Monographs 4 (1): 1–43.
  3. Augustine, J., Kumar, Y. & Sharma, J. (2001) Orchids of India – II: Biodiversity and status of Bulbophyllum Thou. Daya Publishing House, Delhi, 113 pp.
  4. Aung, Y.L., Mu, A.T., Aung, M.H., Liu, Q. & Jin, X.H. (2020) An annotated checklist of Myanmar orchid flora. In: Jin, X.-H., Xia, N.-H. & Tan, Y.-H. (Eds.) Plant diversity of Southeast Asia-II. PhytoKeys 138: 49–112. https://doi.org/10.3897/phytokeys.138.36144
  5. Averyanov, L.V. (1988) Preliminary list of the Vietnamese orchids. Part 1 (genera A–L). Botanical Institute of the Academy of Sciences of the USSR, Leningrad, 247 pp. [in Russian]
  6. Averyanov, L.V. (1990) Orchidaceae. In: Tzvelev, N.N. (Ed.) Vascular plants synopsis of Vietnamese flora. Vol. 1. Nauka, Leningrad, 20 pp. 20–185.
  7. Averyanov, L.V. (1994) Identification guide to Vietnamese orchids (Orchidaceae Juss.). World and Family, St. Petersburg, 432 pp. [in Russian]
  8. Averyanov, L.V. (1999) New species of orchids from Vietnam. Lindleyana 14 (4): 214–224.
  9. Averyanov, L.V. (2006) Rare species of orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 9 (3): 48–89.
  10. Averyanov, L.V. (2010) Species of orchids (Orchidaceae) newly recorded to the flora of Vietnam. Taiwania 55 (1): 1–7. https://doi.org/10.6165/tai.2010.55(1).1
  11. Averyanov, L.V. (2013) New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos. Turczaninowia 16 (4): 26–46. https://doi.org/10.14258/turczaninowia.16.4.7
  12. Averyanov, L.V. & Averyanova, A.L. (2002) Rare species of orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 5 (4): 49–108.
  13. Averyanov, L.V. & Averyanova, A.L. (2003) Updated checklist of the orchids of Vietnam. Vietnam National University Publishing House, Hanoi, 102 pp. [in English and Vietnamese]
  14. Averyanov, L.V. & Gruß, O. (2017) Neue und seltene Orchideen aus Kambodscha und Laos. Orchideen Zauber 2017 (6): 26–39.
  15. Averyanov, L.V., Loc, P.K., Hiep, N.T. & Harder, D.K. (2003) Phytogeographic review of Vietnam and adjacent areas of Eastern Indochina. Komarovia 3: 1–83.
  16. Averyanov, L.V., Nguyen, K.S., Maisak, T.V., Konstantinov, E.L., Nguyen, T.H. & Bounphanmy, S. (2016) New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos. Turczaninowia 19 (3): 5–58. https://doi.org/10.14258/turczaninowia.19.3.1
  17. Averyanov, L.V., Averyanova, A.L., Nguyen, K.S., Orlov, N.L., Maisak, T.V. & Nguyen, H.T. (2018a) New and rare orchid species (Orchidaceae) in the flora of Cambodia and Laos. Novitates Systematicae Plantarum Vascularium 49: 24–41. https://doi.org/10.31111/novitates/2018.49.24
  18. Averyanov, L.V., Nong, V.D., Nguyen, H.T., Nuraliev, M.S., Maisak, T.V. & Nguyen, C.A. (2018b) New species of Bulbophyllum (Orchidaceae) in the flora of Vietnam. Phytotaxa 369 (1): 1–14. https://doi.org/10.11646/phytotaxa.369.1.1
  19. Averyanov, L.V., Nguyen, V.C., Nguyen, K.S., Maisak, T.V. & Truong, B.V. (2019a) New orchids (Orchidaceae) in the flora of Vietnam I. Epidendroideae. Taiwania 64 (2): 176–188. https://doi.org/10.6165/tai.2019.64.176
  20. Averyanov, L.V., Truong, B.V., Nguyen, V.C., Nguyen, K.S. & Maisak, T.V. (2019b) New orchids (Orchidaceae) in the flora of Vietnam II. Vandeae. Taiwania 64 (3): 285–298. https://doi.org/10.6165/tai.2019.64.285
  21. Averyanov, L.V., Truong, B.V., Nguyen, V.C., Maisak, T.V., Dinh, Q.D., Nuraliev, M.S., Nguyen, K.S. & Chu, V.T. (2020) New orchids in the flora of Vietnam III (Collabieae, Malaxideae, Nervilieae and Orchideae). Taiwania 65 (4): 478‒492. https://doi.org/10.6165/tai.2020.65.478
  22. Averyanov, L., Dat, P.T.T., Truong, B.V., Orlov, N., Maisak, T., Nguyen, T.H. & Dinh, Q.D. (2021) Studies of Bulbophyllum (Orchidaceae) in eastern Indochina Ⅳ. New species in the flora of Laos and Vietnam. Phytotaxa 514 (3): 187–204. https://doi.org/10.11646/phytotaxa.514.3.1
  23. Averyanov, L.V., Thai, T.H., Truong, B.V., Nguyen, V.C., Nguyen, T.H., Maisak, T.V., Nguyen, K.S. & Nguyen, V.K. (2022a) New orchids in the flora of Vietnam IV (Orchidaceae, Aeridinae). Phytotaxa 555 (2): 113–135. https://doi.org/10.11646/phytotaxa.555.2.1
  24. Averyanov, L.V., Thai, T.H., Nguyen, V.C., Truong, B.V., Maisak, T.V., Doan, N.T. & Nguyen, K.S. (2022b) New orchids in the flora of Vietnam V (Orchidaceae, Aeridinae). Phytotaxa 572 (1): 43–60. https://doi.org/10.11646/phytotaxa.572.1.3
  25. Averyanov, L.V., Nguyen, V.C., Truong, B.V., Nguyen, K.S., Nguyen, C.H., Maisak, T.V., Doan, N.T., Nguyen, T.H., Pham, V.T., Thanh, P.T., Thai, T.H., Nguyen, V.K. & Trinh, N.B. (2023a) New orchids in the flora of Vietnam VI (Orchidaceae, tribes Arethuseae, Cymbidieae, Diurideae, Epidendreae, Vandeae, and Vanilleae). Phytotaxa 597 (2): 87–110. https://doi.org/10.11646/phytotaxa.597.2.1
  26. Averyanov, L.V., Nguyen, V.C., Truong, B.V., Nguyen, K.S., Nuraliev, M.S., Nguyen, C.H., Ormerod, P.A., Maisak, T.V., Diep, D.Q., Lyskov, D.F. & Nong, D.V. (2023b) New orchids in the flora of Vietnam VII (Orchidaceae: tribes Cypripedieae, Cranichideae, Orchideae, and Collabieae). Phytotaxa 619 (4): 255–276. https://doi.org/10.11646/phytotaxa.619.4.1
  27. Backer, C.A. & Bakhuizen van den Brink, R.C. (1968) Flora of Java 3. N.V.P. Noordhoff, Groningen, 761 pp.
  28. Banerji, M.L. (1996) Orchids of Nepal. Bishen Singh Mahendra Pal Singh, Dehradun, 135 pp.
  29. Beentje, H. (2016) The Kew Plant Glossary: an illustrated dictionary of plant terms (ed. 2). Royal Botanic Gardens, Kew, London, 184 pp.
  30. Blume, C.L. (1825) Bijdragen tot de flora van Nederlandsch Indië. Ter Lands Drukkerij, Batavia, 1169 pp. https://doi.org/10.5962/bhl.title.395
  31. Blume, C.L. (1828) Flora Javae nec non insularum adjacentium. Vol. 1. J. Frank, Bruxellis, 196 pp. https://doi.org/10.5962/bhl.title.48445
  32. Bose, T.K. & Bhattacharjee, S.K. (1980) Orchids of India. Naya Prokash, Calcutta, 538 pp.
  33. Bunpha, K. & Pedersen, H.Æ. (2022) 104. Oberonia. In: Chayamarit, K. & Balslev, H. (Eds.) Flora of Thailand 12 (3). The Forest Herbarium, Royal Forest Department, Bangkok, pp. 854–908.
  34. Bunpha, K.-O., Sridith, K. & Peresen, H.Æ. (2019) A taxonomic revision of Oberonia (Orchidaceae: Epidendroideae: Malaxideae) in Thailand. Phytotaxa 420 (2): 102–156. https://doi.org/10.11646/phytotaxa.420.2.2
  35. Carr, C.E. (1932) Some Malayan orchids, III. Gardens’ Bulletin. Straits Settlements. Singapore 7 (1): 1–59.
  36. Chantanaorrapint, S. (2022) 89. Appendicula. In: Chayamarit, K. & Balslev, H. (Eds.) Flora of Thailand 12 (3). The Forest Herbarium, Royal Forest Department, Bangkok, pp. 676–689.
  37. Chase, M.W., Cameron, K.M., Freudenstein, J.V., Pridgeon, A.M., Salazar, G., van den Berg, C. & Schuiteman, A. (2015) An updated classification of Orchidaceae. Botanical Journal of the Linnean Society 177 (2): 151–174. https://doi.org/10.1111/boj.12234
  38. Chen, S.-C. (1981) Three new species of Listera (Orchidaceae) from China. Kew Bulletin 35 (4): 759–762. https://doi.org/10.2307/4110171
  39. Chen, S., Tsi, Z. & Luo, Y. (1999) Native orchids of China in colour. Science Press, Beijing, New York, 416 pp.
  40. Chen, S.C., Liu, Z.J., Zhu, G.H., Lang, K.Y., Ji, Z.H., Luo, Y.B., Jin, X.H., Cribb, P.J., Wood, J.J., Gale, S.W., Ormerod, P., Vermeulen, J.J., Wood, H.P., Clayton, D. & Bell, A. (2009) Orchidaceae. In: Wu, Z.Y., Raven, P.H. & Hong, D. (Eds.) Flora of China, vol. 25. Science Press, Beijing & Missouri Botanical Garden Press, St. Louis, pp. 211–235.
  41. Clayton, D. (2017) Charles Parish – Plant hunter and botanical artist in Burma. Ray Society, in association with Royal Botanic Gardens, Kew, London, 278 pp.
  42. Comber, J.B. (1990) Orchids of Java. Bentham-Moxon Trust, Royal Botanic Gardens, Kew, 407 pp.
  43. Comber, J.B. (2001) Orchids of Sumatra. The Royal Botanic Gardens, Kew, 1026 pp.
  44. Coode, M.J.E., Dransfield, J., Forman, L.L., Kirkup, D.W. & Said, I.M. (Eds.) (1996) A checklist of the flowering plants and gymnosperms of Brunei Darussalam. Ministry of Industry and Primary Resources, Bandar Seri Begawan, 477 pp.
  45. Cribb, P.J. & Ng, Y.P. (2005) Tribe Podochileae. In: Pridgeon, A.M., Cribb, P.J., Chase, M.W. & Rasmussen, F.N. (Eds.) Genera Orchidacearum 4, Epidendroideae (Part one). Oxford University Press, Oxford, pp. 529–596.
  46. Díaz-Morales, M., Pupulin, P. & Strigari, S. (2019) Bulbophyllum medusae. Orchids (October 2019): 735–737.
  47. Downie, D.C. (1925) Contributions to the flora of Siam. Additamentum XVI. Bulletin of Miscellaneous Information (Royal Botanic Gardens, Kew) 1925 (9): 367–394. https://doi.org/10.2307/4107477
  48. Duong, H.D. & Averyanov, L.V. (1989) A new variety – Dendrobium nobile var. alboluteum from the south Vietnam region. Botanical Journal (Leningrad) 74 (7): 1039–1040.
  49. Fischer, C.E.C. (1928) Flora of the Presidency of Madras. Part 8. West, Newman and Adlard, London, pp. 1347–1532.
  50. Fu, L.-F., Monro, A., Do, T.V., Nuraliev, M.S., Averyanov, L.V., Wen, F., Xin, Z.-B., Maisak, T.V., Kuznetsov, A.N., Kuznetsova, S.P., Nguyen, K.S. & Wei, Y.-G. (2019) Checklist to the Elatostema (Urticaceae) of Vietnam including 19 new records, ten new combinations, two new names and four new synonyms. PeerJ 7: e6188. http://doi.org/10.7717/peerj.6188
  51. Gagnepain, F. (1950) Orchidacées nouvelles d’Indochine. Bulletin du Muséum National d’Histoire Naturelle, sér. 2, 21 (6): 737–743.
  52. Gagnepain, F. & Guillaumin, A. (1932–1934) Orchidacées. In: Gagnepain, F. (Ed.) Flore générale de l’Indo-Chine 6. Masson et Cie., Paris, pp. 142–647.
  53. Gale, S.W. (2022) 89. Podochilus. In: Chayamarit, K. & Balslev, H. (Eds.) Flora of Thailand 12 (3). The Forest Herbarium, Royal Forest Department, Bangkok, pp. 665–971.
  54. Gale, S., Kumar, P. & Phaxaysombath, T. (2018) A guide to orchids of Laos. Natural History Publications, Kota Kinabalu and Kadoorie Farm & Botanic Garden, Hong Kong, 204 pp.
  55. Garay, L.A., Hamer, F. & Siegerist, E.S. (1992) Beiträge zu kultivierten Bulbophyllum-Arten. Die Orchidee 43: 139–141.
  56. Geiger, D.L. (2016) Studies in Oberonia 1: Oberonia maxima and its new synonyms. Orchids 85 (5): 390–395.
  57. Griffith, W. (1851) Notulae ad plantas asiaticas. 3. Bishop’s College Press, Calcutta, 636 pp. https://doi.org/10.5962/bhl.title.70352
  58. Guillaumin, A. (1930) Espèces et localités nouvelles d'Orchidées-Vandées d'Indo-Chine. Bulletin de la Société Botanique de France 77 (3): 326–340. https://doi.org/10.1080/00378941.1930.10833732
  59. Guillaumin, A. (1955) Plantes nouvelles, rares ou critiques des serres du Muséum (Notules sur quelque Orchidées d’Indochine, IX). Bulletin du Muséum National d’Histoire Naturelle, sér. 2, 26 (6): 691–694.
  60. Guillaumin, A. (1961) Plantes nouvelles, rares ou critiques des serres du Muséum (Notules sur quelques Orchidees d'lndochine, XXV). Bulletin du Muséum National d’Histoire Naturelle, sér. 2, 32 (6): 562–565.
  61. Guillaumin, A. (1963) Plantes nouvelles, rares ou critiques des serres du Muséum (Notules sur quelques Orchidées s’Indochine, XXXI). Bulletin du Muséum National d’Histoire Naturelle, sér. 2, 35 (2): 204–206.
  62. Gyeltshen, C., Gyeltshen, N. & Higgins, W.E. (2021) A century of new orchid records in Bhutan, ed 2. National Biodiversity Centre, Thimphu, 210 pp.
  63. Ho, P.-H. (1972) Cay-Co Mien-Nam Viet-Nam. 2. ed., 2, Nam Quoc-Te Phat-Trien Sach, Saigon, 1139 pp.
  64. Ho, P.-H. (1993) An illustrated flora of Vietnam. 3 (2). Mekong Printing, Montreal, 1176 pp.
  65. Ho, P.-H. (2000) An illustrated flora of Vietnam. 3. Nha Xuat Ban Tre, Ho Chi Minh City, 1020 pp.
  66. Holttum, R.E. (1964) A revised flora of Malaya. Vol. 1. Orchids of Malaya. 3rd ed. Government Printing Office, Singapore, 759 pp.
  67. Hooker, J.D. (1888–1890) Order CXLVIII Orchideae. In: Hooker, J.D. (Ed.) The flora of British India 5. Reeve, London, pp. 667–910. https://doi.org/10.5962/bhl.title.678
  68. Hop, T. (1998) The orchids of Vietnam. Nha Xuat Ban Nong Nghiep, Ho Chi Minh City, 703 pp.
  69. Huang, M.-Z., Yang, G.-S., Lan, S.-R. & Yin, J.-M. (2020) Bulbophyllum lingii, a new species (Malaxideae, Epidedroideae, Orchidaceae) from Hainan, China. Phytotaxa 452 (2): 185–188. https://doi.org/10.11646/phytotaxa.452.2.8
  70. Hunt, P.F. (1971) Notes on asiatic orchids: VI. Kew Bulletin 26 (1): 171–185. https://doi.org/10.2307/4117342
  71. IUCN Standards and Petitions Committee (2024) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. Prepared by the Standards and Petitions Committee. Available from: https://nc.iucnredlist.org/redlist/content/attachment_files/RedListGuidelines.pdf (accessed 20 February 2024).
  72. Jalal, J.S. & Jayanthi, J. (2015) An annotated checklist of the orchids of Western Himalaya, India. Lankesteriana 15 (1): 7–50. https://doi.org/10.15517/lank.v15i1.18521
  73. Jin, X., Li, J. & Ye, D. (2023) Atlas of Chinese native orchids. Springer, Singapore, 787 pp. https://doi.org/10.1007/978-981-99-4853-6
  74. Kaliamoorthy, S. & Saravanan, T.S. (2019) Additions to the orchid flora of Peninsular India. Rheedea 29 (4): 319–322. https://dx.doi.org/10.22244/rheedea.2019.29.4.05
  75. Kerr, A.D. (1969) On a collection of orchids from Laos. Natural History Bulletin of the Siam Society 23: 185–211.
  76. Li, J.W., Ye, D.P., Liu, Q. & Yin, J.T. (2013) Two new records of orchid from China. Plant Diversity and Resources 35 (2): 128–130. https://doi.org/10.7677/ynzwyj201312105
  77. Lindley, J. (1825) Eria stellata. Starry Eria. Edward’s Botanical Register 11: tab. 904.
  78. Lindley, J. (1830–1840) The genera and species of orchidaceous plants. James Ridgway and Sons, London, 554 pp. https://doi.org/10.5962/bhl.title.499
  79. Lindley, J. (1842) Cirropetalum medusae. Edwards’s Botanical Register 28: tab. 12.
  80. Lindley, J. (1845) Dendrobium anosmum. Edwards’s Botanical Register 31: 32.
  81. Lindley, J. (1847a) Bulbophyllum lobbii. Edwards’s Botanical Register 33: sub pl. 29.
  82. Lindley, J. (1847b) Dendrobium cretaceum. Chalk-white Dendrobe. Edwards’s Botanical Register 33: tab. 62.
  83. Lindley, J. (1858) Contributions to the orchidology of India. – No. II. Journal of the Proceedings of the Linnean Society of London. Botany 3 (9): 1–63. https://doi.org/10.1111/j.1095-8339.1858.tb02055.x
  84. Lindley, J. (1898) Dendrobium nobile Lindl. varietates. Lindenia 14: 45–48. https://doi.org/10.5962/bhl.title.64391
  85. Lindsay, S., Middleton, D.J., Ho, B.C., Chong, K.Y., Turner, I.M., Ali, Ibrahim, Alonso-García, M., Ang, W.F., Ashton, P.S., Athen, P., Atkins, S., Bazilah Ibrahim, Beentje, H.J., Boo, C.M., Boyce, P.C., Bramley, G.L.C., Buerki, S., Callmander, M.W., Chantanaorrapint, S., Cheek, M., Chen, C.-W., Chen, J., Chen, L.M.J., Chew, P.T., Chong, R., Choo, L.M., Chung, R.C.K., Coode, M.J.E., Chua, S.C., Cicuzza, D., de Kok, R.P.J., Davison, G.W.H., de Wilde, W.J.J.O., Duistermaat, H., Dubéarnès, A., Duyfjes, B.E.E., Ellis, L.T., Esser, H.-J., Gajurel, P.R., Gale, S.W., Ganesan, S.K., Gardner, E.M., Geiger, D.L., Harwood, R.K., Hassan Ibrahim, He, S., Henderson, A., Hovenkamp, P.H., Hughes, M., Zaki Jamil, Jebb, M.H.P., Johnson, D.M., Kartonegoro, A., Kiew, R., Knapp, S., Koh, S.L., Kurzweil, H., Lee, S., Leong, P.K.F., Leong-Škorničková, J., Levin, G.A., Liew, D.C.H., Lim, R.C.J., Lim, W.H., Loo, A.H.B., Low, Y.W., Lua, H.K., Lum, S., Mabberley, D.J., Mahyuni, R., Maslin, B., Murray, N.A., Neo, L., Ng, X.Y., Ngo, K.M., Niissalo, M.A., Ong, P.T., Pannell, C.M., Phang, A., Prance, G.T., Promma, C., Puglisi, C., Rodda, M.L., Rubasinghe, S.C.K., Saunders, R.M.K., Savinov, I.A., Saw, L.G., Schuiteman, A., Seah, W.W., Simpson, D.A., Strijk, J.S., Sukkharak, P., Sugumaran, M., Syahida-Emiza, S., Tan, J.P.C., Taylor, N.P., Teo, Y.K.L., Thomas, D.C., Trias-Blasi, A., Utteridge, T., van Welzen, P.C., Veldkamp, J.F., Vermeulen, J., Wang, R., Wilkie, P., Wei, Y.-M., Wong, S.Y., Wong, K.M., Yaakub, S., Yam, T.W., Yang, S., Yao, T.L., Ye, W., Yee, A.T.K., Yeo, C.K., Yeoh, Y.S., Yong, C., Yong, K.T., Zerega, N.J.C., Zhu, R.-L. & Er, K.B.H. (2022) Flora of Singapore: Checklist and bibliography. Gardens’ Bulletin Singapore 74 (Suppl. 1): 3–860. https://doi.org/10.26492/gbs74(suppl.1).2022-01
  86. Liu, Q. & Gao, J.Y. (2016) New orchids record in the flora of China. Current Science 110 (11): 2064–2066.
  87. Low, H. (1863) Dendrobium parishii. Proceedings of the Royal Horticultural Society, London 3: 281.
  88. Newman, M., Ketphanh, S., Svengsuksa, B., Thomas, P., Sengdala, K., Lamxay, V. & Armstrong, K. (2007) A checklist of the vascular plants of Lao PDR. Royal Botanical Garden Edinburgh, Edinburgh, 394 pp.
  89. Ng, Y.P., Schuiteman, A., Pedersen, H.Æ., Petersen, G., Watthana, S., Seberg, O., Pridgeon, A.M., Cribb, P.J. & Chase, M.W. (2018) Phylogenetics and systematics of Eria and related genera (Orchidaceae: Podochileae). Botanical Journal of the Linnean Society 186 (2): 179–201. https://doi.org/10.1093/botlinnean/box088
  90. Nguyen, V.C., Averyanov, L.V., Nguyen, V.K., Pham, T.T.D., Maisak, T.V. & Truong, B.V. (2020) Cylindrolobus chienii (Orchidaceae), a new species from Central Highlands of Vietnam. Taiwania 65 (3): 272‒276. https://doi.org/10.6165/tai.2020.65.272
  91. Nuraliev, M.S., Schori, M., Tagane, S., Kuznetsov, A.N., Kuznetsova, S.P. & Utteridge, T.M. (2022) Citronella suaveolens, a new generic record for Vietnam, with a key to Vietnamese Cardiopteridaceae. Phytotaxa 532 (1): 67–77. https://doi.org/10.11646/phytotaxa.532.1.5
  92. O’Byrne, P. (2001) A to Z of South East Asian orchid species. Orchid Society of South East Asia, Singapore, 168 pp.
  93. Ormerod, P. (2004) Orchidaceous additions to the floras of China and Taiwan (II). Taiwania 49 (2): 95–101. https://doi.org/10.6165/tai.2004.49(2).95
  94. Ormerod, P. (2014) A synopsis of Eria Lindl. section Cylindrolobus (Blume) Lindl. (Orchidaceae: Eriinae) in Malesia. Harvard Papers in Botany 19 (1): 77–95. https://doi.org/10.3100/hpib.v19iss1.2014.n6
  95. Ormerod, P., Kurzweil, H. & Watthana, S. (2021) Annotated list of Orchidaceae for Myanmar. Phytotaxa 481 (1): 1–262. https://doi.org/10.11646/phytotaxa.481.1.1
  96. Pearce, N.R. & Cribb, P.J. (2002) The orchids of Bhutan. The Charlesworth Group, Huddersfield, 643 pp.
  97. Phonepaseuth, P., Souvannakhoummane, K., Xayyasith, S., Keovongsy, C., Soulida, K. & Souladeth, P. (2023) Two new distribution records of Dendrobium (Orchidaceae), with an updated checklist of Dendrobium for Laos. Thai Forest Bulletin, Botany 51 (1): 24–35. https://doi.org/10.20531/tfb.2023.51.1.04
  98. POWO (2024) Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet. Available from: http://www.plantsoftheworldonline.org/ (Retrieved 23 March 2024).
  99. Raskoti, B.B. (2009) The orchids of Nepal. Bhakta Bahadur Raskoti and Rita Ale, Kathmandu, 252 pp.
  100. Reichenbach, H.G. (1861) Subclassis VIII. Gynandrae Endl. Annales Botanices Systematicae (Walpers) 6: 167–933. https://doi.org/10.5962/bhl.title.7556
  101. Reichenbach, H.G. (1867) Bulbophyllum siamense, Rchb.f. Gardeners’ Chronicle 1867: 572.
  102. Reichenbach, H.G. (1874) Enumeration of the orchids collected by the rev. E.C. Parish in the neighborhood of Moulmein, with descriptions of the new species. Transactions of the Linnean Society of London 30 (1): 133–155. https://doi.org/10.1111/j.1096-3642.1874.tb00004.x
  103. Reichenbach, H.G. (1882) Acrochaene Rimannii n.sp. Gardeners’ Chronicle 17: 796.
  104. Retzius, A.J. (1791) Observationes botanicae: sex fasciculis comprehensae 6. Siegfried Lebrecht Crusium, Lipsiae, 67 pp. https://doi.org/10.5962/bhl.title.11760
  105. Ridley, H.N. (1920) New and rare species of Malayan plants. Journal of the Straits Branch of the Royal Asiatic Society 82: 167–204.
  106. Ridley, H.N. (1924) The flora of the Malay Peninsula. IV. Monocotyledones. L. Reeve & Co., Ltd, London; Reprint, 1967 by A. Asher & Co., Amsterdam, and L. Reeve & Co., Brook Nr. Ashford, 383 pp.
  107. Rokaya, M.B., Raskoti, B.B., Timsina, B. & Münzbergová, Z. (2013) An annotated checklist of the orchids of Nepal. Nordic Journal of Botany 31 (5): 511–550. https://doi.org/10.1111/j.1756-1051.2013.01230.x
  108. Rolfe, A.R. (1908) New orchids: decade 32. Bulletin of Miscellaneous Information (Royal Botanic Gardens, Kew) 1908 (9): 412–416. https://doi.org/10.2307/4113217
  109. Roxburgh, W. (1795) Plants of the coast of Coromandel. W. Bulmer and Co., London, 100 pp. https://doi.org/10.5962/bhl.title.467
  110. Saunders, W. (1872) Tribe Malaxideae. Genus Bulbophyllum, P.Th. Refugium Botanicum 2: tab. 116.
  111. Schlechter, R. (1914) Die Orchideen: Ihre Beschreibung, Kultur und Züchtung. P. Parey, Berlin, 837 pp. https://doi.org/10.5962/bhl.title.46623
  112. Schuiteman, A., Bonnet, P., Svengsuksa, B. & Barthélémy, D. (2008) An annotated checklist of the Orchidaceae of Laos. Nordic Journal of Botany 26 (5–6): 257–316. https://doi.org/10.1111/j.1756-1051.2008.00265.x
  113. Schuiteman, A., Ryan, C. & Nut, M. (2015) New records of Orchidaceae from Cambodia I. Cambodian Journal of Natural History 2015 (2): 131–138.
  114. Schuiteman, A., Ryan, C., Nut, M., Nay, S. & Att, S. (2016) New records of Orchidaceae from Cambodia III. Cambodian Journal Natural History 2016 (2): 84–89.
  115. Schuiteman, A., Nay, S. & Att, S. (2022) New records of Orchidaceae from Cambodia V. Cambodian Journal of Natural History 2022 (2): 127–143.
  116. Seidenfaden, G. (1968) The genus Oberonia Lindl. in mainland Asia. Dansk Botanisk Arkiv 25 (3): 1–125.
  117. Seidenfaden, G. (1969) Notes on the genus Ione. Botanisk Tidsskrift 64 (2–3): 205–238.
  118. Seidenfaden, G. (1971) Contributions to the orchid flora of Thailand III. Botanisk Tidsskrift 66 (4): 303–356.
  119. Seidenfaden, G. (1972a) An enumeration of Laotian orchids. Bulletin du Muséum National d’Histoire Naturelle, sér. 3, 71, Botanique 5: 101–152.
  120. Seidenfaden, G. (1972b) Contributions to the orchid flora of Thailand IV. Botanisk Tidsskrift 67 (1–2): 76–127.
  121. Seidenfaden, G. (1973a) Notes on Cirrhopetalum Lindl. Dansk Botanisk Arkiv 29 (1): 1–260.
  122. Seidenfaden, G. (1973b) Contributions to the orchid flora of Thailand V. Botanisk Tidsskrift 68 (1): 41–95.
  123. Seidenfaden, G. (1975a) Contributions to a revision of the orchid flora of Cambodia, Laos and Vietnam. Kai Olsen, Fredensborg, 117 pp.
  124. Seidenfaden, G. (1975b) Contributions to the orchid flora of Thailand VI. Botanisk Tidsskrift 70 (1): 64–97.
  125. Seidenfaden, G. (1976) Contributions to the orchid flora of Thailand VII. Botanisk Tidsskrift 71 (1–2): 1–30.
  126. Seidenfaden, G. (1978) Orchid genera in Thailand VII. Oberonia Lindl. & Malaxis Sol. ex Sw. Dansk Botanisk Arkiv 33 (1): 1–94.
  127. Seidenfaden, G. (1979) Orchid genera in Thailand VIII. Bulbophyllum. Dansk Botanisk Arkiv 33 (3): 1–240.
  128. Seidenfaden, G. (1980) A new species of Sunipia (Orchidaceae) from Thailand. Natural History Bulletin of the Siam Society 28: 1–8.
  129. Seidenfaden, G. (1982) Contributions to the orchid flora of Thailand X. Nordic Journal of Botany 2 (3): 193–218. https://doi.org/10.1111/j.1756-1051.1982.tb01182.x
  130. Seidenfaden, G. (1985) Orchid genera in Thailand XII. Dendrobium Sw. Opera Botanica 83: 1–295.
  131. Seidenfaden, G. (1986a) A collection of orchids from Malaya. Nordic Journal of Botany 6 (2): 157–181. https://doi.org/10.1111/j.1756-1051.1986.tb00872.x
  132. Seidenfaden, G. (1986b) Orchid genera in Thailand XIII. Thirty-three epidendroid genera. Opera Botanica 89: 1–216.
  133. Seidenfaden, G. (1992) The orchids of Indochina. Opera Botanica 114: 1–502.
  134. Seidenfaden, G. (1995a) Contributions to the orchids of Thailand XII. Opera Botanica 124: 1–90.
  135. Seidenfaden, G. (1995b) The Descriptiones Epidendrorum of J.G. König 1791. Olsen & Olsen, Fredensborg, 61 pp.
  136. Seidenfaden, G. (1997) Contributions to the orchid flora of Thailand XIII. Olsen & Olsen, Fredensborg, 64 pp.
  137. Seidenfaden, G. & Smitinand, T. (1959–1965) The orchids of Thailand. A preliminary list. Siam Society, Bangkok, 870 pp.
  138. Seidenfaden, G. & Wood, J.J. (1992) The orchids of Peninsular Malaysia and Singapore. Olsen & Olsen, Fredensborg, 779 pp.
  139. Simpson, M.G. (2019) Plant systematics. 3 ed. Elsevier, Amsterdam etc., 761 pp. https://doi.org/10.1016/B978-0-12-812628-8.50001-8
  140. Smith, J.J. (1905) Die orchideen von Java. E.J. Brill, Leiden, 672 pp. https://doi.org/10.5962/bhl.title.86938
  141. Smith, J.J. (1908) Neue Orchideen des malaiischen Archipels, II. Bulletin du Département de l’Agriculture aux Indes Néerlandaises 15: 1–29.
  142. Smith, J.J. (1932) Orchidaceae novae Malayenses XIV. Bulletin du Jardin Botanique de Buitenzorg, sér. 3, 12: 105–150.
  143. Stapf, O. (1894) On the flora of Mount Kinabalu, in north Borneo. Transactions of the Linnean Society of London, Botany 4: 69–263. https://doi.org/10.1111/j.1095-8339.1894.tb00044.x
  144. Swartz, O. (1799) Dendrobium Sw. Nova Acta Regiae Societatis Scientiarum Upsaliensis. Ser. 2, 6: 82–85.
  145. Tanaka, N., Yukawa, T., Htwe, K.M., Koyama, T. & Murata, J. (2011) New or noteworthy plant collections from Myanmar (7): fourteen additional species of Orchidaceae. Acta Phytotaxonomica et Geobotanica 61 (3): 161–165. https://doi.org/10.18942/apg.KJ00007062775
  146. Tanaka, N., Yukawa, T., Htwe, K.M. & Murata, J. (2015) An orchid checklist of Mt. Popa, central Myanmar. Bulletin of the National Museum of Nature and Science, ser. B, 41 (2): 69–89.
  147. Teoh, E.S. (2021) Orchid species from Himalaya and Southeast Asia 1 (A–E). Springer, Cham, 504 pp. https://doi.org/10.1007/978-3-030-58872-4
  148. Thiers, B. (2023) [continuously updated] Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 20 February 2024).
  149. Thouars du Petit, A. (1822) Histoire particulière des plantes Orchidées recueillies sur les trois îles australes d’Afrique, de France, de Bourbon et de Madagascar. L’auteur, Paris, 32 pp. https://doi.org/10.5962/bhl.title.492
  150. Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashütten. https://doi.org/10.12705/Code.2018
  151. Vermeulen, J.J. (1991) Orchids of Borneo 2: Bulbophyllum. Royal Botanic Gardens, Kew, 342 pp.
  152. Vermeulen, J.J. & O'Byrne, P. (2003) New species and new records of Southeast Asian Bulbophyllum (Orchidaceae). Gardens’ Bulletin Singapore 55 (1): 125–162.
  153. Vermeulen, J.J., Schuiteman, A. & de Vogel, E.F. (2014) Nomenclatural changes in Bulbophyllum (Orchidaceae; Epidendroideae). Phytotaxa 166 (2): 101–113. https://doi.org/10.11646/phytotaxa.166.2.1
  154. Vermeulen, J., O’Byrne, P. & Lamb, A. (2015) Bulbophyllum of Borneo. Natural History Publications (Borneo), Kota Kinabalu, 728 pp.
  155. Vietnam Administrative Atlas (2015) Vietnam Publishing House of Natural Resources, Environment and Cartography, Hanoi, 131 pp.
  156. Wallich, N. (1830) Dendrobium densiflorum. Plantae Asiaticae Rariores 1: 34. https://doi.org/10.5962/bhl.title.468
  157. Williams, B.S. (1852) The orchid-grower’s manual, containing a brief description of upwards of two hundred and sixty orchidaceous plant. Chapman and Hall, London, 108 pp. https://doi.org/10.5962/bhl.title.84830
  158. Wood, J.J. (2014) Dendrobium of Borneo. Natural History Publications (Borneo), Kota Kinabalu, and Royal Botanic Gardens, Kew, London, 946 pp.
  159. Wood, J.J. & Cribb, P.J. (1994) A checklist of the orchids of Borneo. Royal Botanic Gardens, Kew, 409 pp.
  160. Xu, Z., Jiang, H., Ye, D. & Liu, E. (2010) The wild orchids in Yunnan. Yunnan Science and Technology Press, Kunming, 500 pp. [in Chinese]
  161. Ya, J.-D., Zhang, T., Pandey, T.R., Liu, C., Han, Z.-D., Ye, D.-P., He, D.-M., Liu, Q., Yang, L., Huang, L., Zhang, R.-Z., Jiang, H. & Cai, J. (2021) New contributions to Goodyerinae and Dendrobiinae (Orchidaceae) in the flora of China. Plant Diversity 443 (5): 362–378. https://doi.org/10.1016/j.pld.2021.05.006
  162. Ya, J.-D., Jiang, H., He, Z.-J., Liao, Q.-C., Zhao, Y.-H., Cai, J., Wang, H. & Xiong, Z. (2023) Cylindrolobus gaoligongensis sp. nov. (Orchidaceae, Podochileae) from Yunnan, China, and improved description of C. arunachalensis. Nordic Journal of Botany 2023 (12): e04075. https://doi.org/10.1111/njb.04075
  163. Zhou, X., Cheng, Z., Liu, Q., Zhang, J., Hu, A., Huang, M., Hu, C. & Tian, H. (2016) An updated checklist of Orchidaceae for China, with two new national records. Phytotaxa 276 (1): 1–148. https://doi.org/10.11646/phytotaxa.276.1.1