Abstract
Polystichum kuankuoshuiense (Dryopteridaceae), a new species from Kuankuoshui Nature Reserve, Suiyang County, Guizhou, China, is described and illustrated. It is a member of P. sect. Sphaenopolystichum and similar to P. auriculum, P. bifidum, and P. suiyangense but differs from them in its rhizome, scale color, and pinna morphology. Quite surprisingly, the new species is not closely related to these morphologically similar species. The new species is currently known only from the type locality and is classified as CR (critically endangered) following the IUCN criteria.
References
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ching, R.-C. (1958) Icones Filicum Sinicarum, vol. 5. Science Press, Beijing, 250 plates.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ching, R.-C. (1949) The studies of Chinese ferns – XXXV. Bulletin of Fan Memorial Institute of Biology New Series 1: 267–317.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Duan, Y.-F., Lu, N.T., Zhang, L. & Zhang, L.-B. (2017a) Polystichum luteoviride (subg. Haplopolystichum; Dryopteridaceae), a new cave species from Guizhou, China—One of the most endangered species. Phytotaxa 313: 296–300. https://doi.org/10.11646/phytotaxa.313.3.10</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Duan, Y.-F., Kropf, M. & Zhang, L.-B. (2017b) Polystichum zhijinense (subg. Haplopolystichum; Dryopteridaceae), a new cave species of Polystichum from Guizhou, China. Phytotaxa 331: 124–130. https://doi.org/10.11646/phytotaxa.331.1.11</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Han, M.-Q., Liu, Y. & Zhang, L.-B. (2016) Seven new species of Polystichum (subg. Haplopolystichum; Dryopteridaceae) from southern China. Phytotaxa 280: 201–221. https://doi.org/10.11646/phytotaxa.280.3.1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Han, M.-Q., Liu, Y. & Zhang, L.-B. (2018) Eight new species of Polystichum (subg. Haplopolystichum; Dryopteridaceae) from limestone caves in Guangdong and Yunnan, China, with reference to species diversity in the karst terrains at high elevations in subtropical areas. Phytotaxa 365: 145–168. https://doi.org/10.11646/phytotaxa.365.2.3</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">He, H. & Zhang, L.-B. (2017) Polystichum tiandengense (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangxi, China. Novon 25: 153–157. https://doi.org/10.3417/2016025</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">He, H. & Zhang, L.-B. (2022) Polystichum gonggashanense (Dryopteridaceae): a new fern from Sichuan, China. Phytotaxa 559: 207–212.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Kung, H.-S., Chu, W.-M., He, Z.-R. & Zhang, L.-B. (2001) Polystichum. In: Kung, H.-S. (Ed.) Flora Reipublicae Popularis Sinicae, vol. 5 (2). Science Press, Beijing, pp. 1–246.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Le Péchon, T., Zhang, L., Zhou, X.-M., He, H., Gao, X.-F. & Zhang, L.-B. (2016a) A well-sampled phylogenetic analysis of the polystichoid ferns (Dryopteridaceae) suggests a complex biogeographical history involving both boreotropical migrations and recent transoceanic dispersals. Molecular Phylogenetics and Evolution 98: 324–336. https://doi.org/10.1016/j.ympev.2016.02.018</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Le Péchon, T., He, H., Zhang, L., Zhou, X.-M., Gao, X.-F. & Zhang, L.-B. (2016b) Using a multilocus phylogeny to test morphology-based classifications of one of the largest fern genera Polystichum (Dryopteridaceae). BMC Evolutionary Biology 16: 55. https://doi.org/10.1186/s12862-016-0626-z</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Li, C.-X., Patel, N.R. & Zhang, L.-B. (2016) Polystichum clarinervium (subg. Haplopolystichum; Dryopteridaceae), a new fern from Emei Shan, China. Phytotaxa 280: 271–277. https://doi.org/10.11646/phytotaxa.280.3.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Liang, Z.-L., He, Z.-R. & Zhang, L.-B. (2021) Polystichum dongchuanense and P. menglaense (subg. Haplopolystichum; Dryopteridaceae), two new ferns from Yunnan, China. Phytotaxa 479: 114–120. https://doi.org/10.11646/phytotaxa.539.1.8</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Liang, Z.-L., Yu, Z.-Y., Shui, Y.-M., Chen, W.-H., Tunckol, B., Zhang, L. & Zhang, L.-B. (2022) Polystichum jinpingense (subg. Haplopolystichum; Dryopteridaceae), a new fern from southeastern Yunnan, China. Phytotaxa 539: 69–73. https://doi.org/10.11646/phytotaxa.479.1.9</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Liang, Y.-Y. & Zhang, L.-B. (2022) Polystichum neocavernicola (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangxi, China. Phytotaxa 559: 95–98.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lu, N.T., Zhang, L., Nguyen, D.T. & Zhang, L.-B. (2016) Polystichum quangbinhense sp. nov. (subg. Haplopolystichum; Dryopteridaceae), the southernmost cave species of Polystichum from central Vietnam. Phytotaxa 283: 295–299. https://doi.org/10.11646/phytotaxa.283.3.6</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lu, N.T., Duan, Y.-F. & Zhang, L.-B. (2017) Polystichum alluvium (subg. Haplopolystichum; Dryopteridaceae), a new cave species from Guizhou, China, with reference to new distribution records of P. leveillei. Phytotaxa 323: 88–92. https://doi.org/10.11646/phytotaxa.323.1.8</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lu, N.T., Zhang, L. & Zhang, L.-B. (2014) Ten new records of Polystichum (Dryopteridaceae) for Vietnam and lectotypification of Polystichum atroviridissimum and P. fimbriatum. Phytotaxa 283: 295–299. https://doi.org/10.11646/phytotaxa.164.2.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Miao, X.-Y., Li, C.-X. & Zhang, L.-B. (2017) Polystichum duyunense (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guizhou, China. Phytotaxa 292: 296–300. https://doi.org/10.11646/phytotaxa.292.3.10</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Roth, A.W. (1800) Tentamen Florae Germanicae, vol. 3. Gleditsch, Leipzig, 578 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Shen, X.-L, Liu, J. & Jiang, R.-H. (2015) Polystichum mulunense (Dryopteridaceae), a New Species from Karst Caves in Guangxi, China. Plant Diversity and Resources 37: 737–740. https://doi.org/10.7677/ynzwyj201515068</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tagawa, M. (1940) Polystichum of Japan, Korea, and Formosa I. Acta Phytotaxononica Geobotanica 9: 119–138.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tang, G.-D., Huang, L., Li, J.-Y., He, Z.-L. & Zhang, L.-B. (2017) Polystichum hastipinnum (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangdong, China. Phytotaxa 309: 66–72. https://doi.org/10.11646/phytotaxa.309.1.6</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, P.-S. & Wang, X.-Y. (2001) Pteridophyte flora of Guizhou. Guizhou Science & Technology Press, Guiyang, 727 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, P.-S. & Pan, L.-T. (2018) Lycophytes and Ferns of Guizhou, (vol. II). Guizhou Science & Technology Press, Guiyang, 976 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, B., Sun, Q.-W., Wang, Z.-H., Chen, C.-L., Xu, W.-F. & Zhang, L.-B. (2022) Polystichum suiyangense (sect. Sphaenopolystichum, Dryopteridaceae), a new fern from Guizhou, China. Phytotaxa 567: 93–99.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wei, H.-J. & Zhang, L.-B. (2018) Polystichum recavum (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangxi, China. Phytotaxa 374: 167–171. https://doi.org/10.11646/phytotaxa.374.2.9</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhang, L., Zhu, Z.-M., Gao, X.-F. & Zhang, L.-B. (2012) Polystichum hubeiense (Dryopteridaceae), a new fern species from Hubei, China. Annales Botanici Fennici 50: 107–110. https://doi.org/10.5735/085.050.0121</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhang, L.-B. & Barrington, D.S. (2013) Polystichum. In: Wu, Z.-Y., Raven, P.H. & Hong, D.-Y. (Eds.) Flora of China, vols. 2–3. Science Press, Beijing & Missouri Botanical Garden Press, St. Louis, pp. 629–713.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhang, L.-B., Duan, Y.-F. & Kropf, M. (2019) Eight new species of the fern genus Polystichum (subg. Haplopolystichum; Dryopteridaceae) from limestone caves in Guizhou, China. Phytotaxa 404: 209–230. https://doi.org/10.11646/phytotaxa.404.6.1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhang, L.-B. & He, H. (2009) Polystichum peishanii, a new species known only from three populations on limestones in southern Guizhou, China. Botanical Studies 50: 101–106.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ching, R.-C. (1949) The studies of Chinese ferns – XXXV. Bulletin of Fan Memorial Institute of Biology New Series 1: 267–317.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Duan, Y.-F., Lu, N.T., Zhang, L. & Zhang, L.-B. (2017a) Polystichum luteoviride (subg. Haplopolystichum; Dryopteridaceae), a new cave species from Guizhou, China—One of the most endangered species. Phytotaxa 313: 296–300. https://doi.org/10.11646/phytotaxa.313.3.10</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Duan, Y.-F., Kropf, M. & Zhang, L.-B. (2017b) Polystichum zhijinense (subg. Haplopolystichum; Dryopteridaceae), a new cave species of Polystichum from Guizhou, China. Phytotaxa 331: 124–130. https://doi.org/10.11646/phytotaxa.331.1.11</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Han, M.-Q., Liu, Y. & Zhang, L.-B. (2016) Seven new species of Polystichum (subg. Haplopolystichum; Dryopteridaceae) from southern China. Phytotaxa 280: 201–221. https://doi.org/10.11646/phytotaxa.280.3.1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Han, M.-Q., Liu, Y. & Zhang, L.-B. (2018) Eight new species of Polystichum (subg. Haplopolystichum; Dryopteridaceae) from limestone caves in Guangdong and Yunnan, China, with reference to species diversity in the karst terrains at high elevations in subtropical areas. Phytotaxa 365: 145–168. https://doi.org/10.11646/phytotaxa.365.2.3</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">He, H. & Zhang, L.-B. (2017) Polystichum tiandengense (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangxi, China. Novon 25: 153–157. https://doi.org/10.3417/2016025</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">He, H. & Zhang, L.-B. (2022) Polystichum gonggashanense (Dryopteridaceae): a new fern from Sichuan, China. Phytotaxa 559: 207–212.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Kung, H.-S., Chu, W.-M., He, Z.-R. & Zhang, L.-B. (2001) Polystichum. In: Kung, H.-S. (Ed.) Flora Reipublicae Popularis Sinicae, vol. 5 (2). Science Press, Beijing, pp. 1–246.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Le Péchon, T., Zhang, L., Zhou, X.-M., He, H., Gao, X.-F. & Zhang, L.-B. (2016a) A well-sampled phylogenetic analysis of the polystichoid ferns (Dryopteridaceae) suggests a complex biogeographical history involving both boreotropical migrations and recent transoceanic dispersals. Molecular Phylogenetics and Evolution 98: 324–336. https://doi.org/10.1016/j.ympev.2016.02.018</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Le Péchon, T., He, H., Zhang, L., Zhou, X.-M., Gao, X.-F. & Zhang, L.-B. (2016b) Using a multilocus phylogeny to test morphology-based classifications of one of the largest fern genera Polystichum (Dryopteridaceae). BMC Evolutionary Biology 16: 55. https://doi.org/10.1186/s12862-016-0626-z</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Li, C.-X., Patel, N.R. & Zhang, L.-B. (2016) Polystichum clarinervium (subg. Haplopolystichum; Dryopteridaceae), a new fern from Emei Shan, China. Phytotaxa 280: 271–277. https://doi.org/10.11646/phytotaxa.280.3.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Liang, Z.-L., He, Z.-R. & Zhang, L.-B. (2021) Polystichum dongchuanense and P. menglaense (subg. Haplopolystichum; Dryopteridaceae), two new ferns from Yunnan, China. Phytotaxa 479: 114–120. https://doi.org/10.11646/phytotaxa.539.1.8</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Liang, Z.-L., Yu, Z.-Y., Shui, Y.-M., Chen, W.-H., Tunckol, B., Zhang, L. & Zhang, L.-B. (2022) Polystichum jinpingense (subg. Haplopolystichum; Dryopteridaceae), a new fern from southeastern Yunnan, China. Phytotaxa 539: 69–73. https://doi.org/10.11646/phytotaxa.479.1.9</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Liang, Y.-Y. & Zhang, L.-B. (2022) Polystichum neocavernicola (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangxi, China. Phytotaxa 559: 95–98.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lu, N.T., Zhang, L., Nguyen, D.T. & Zhang, L.-B. (2016) Polystichum quangbinhense sp. nov. (subg. Haplopolystichum; Dryopteridaceae), the southernmost cave species of Polystichum from central Vietnam. Phytotaxa 283: 295–299. https://doi.org/10.11646/phytotaxa.283.3.6</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lu, N.T., Duan, Y.-F. & Zhang, L.-B. (2017) Polystichum alluvium (subg. Haplopolystichum; Dryopteridaceae), a new cave species from Guizhou, China, with reference to new distribution records of P. leveillei. Phytotaxa 323: 88–92. https://doi.org/10.11646/phytotaxa.323.1.8</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lu, N.T., Zhang, L. & Zhang, L.-B. (2014) Ten new records of Polystichum (Dryopteridaceae) for Vietnam and lectotypification of Polystichum atroviridissimum and P. fimbriatum. Phytotaxa 283: 295–299. https://doi.org/10.11646/phytotaxa.164.2.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Miao, X.-Y., Li, C.-X. & Zhang, L.-B. (2017) Polystichum duyunense (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guizhou, China. Phytotaxa 292: 296–300. https://doi.org/10.11646/phytotaxa.292.3.10</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Roth, A.W. (1800) Tentamen Florae Germanicae, vol. 3. Gleditsch, Leipzig, 578 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Shen, X.-L, Liu, J. & Jiang, R.-H. (2015) Polystichum mulunense (Dryopteridaceae), a New Species from Karst Caves in Guangxi, China. Plant Diversity and Resources 37: 737–740. https://doi.org/10.7677/ynzwyj201515068</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tagawa, M. (1940) Polystichum of Japan, Korea, and Formosa I. Acta Phytotaxononica Geobotanica 9: 119–138.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tang, G.-D., Huang, L., Li, J.-Y., He, Z.-L. & Zhang, L.-B. (2017) Polystichum hastipinnum (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangdong, China. Phytotaxa 309: 66–72. https://doi.org/10.11646/phytotaxa.309.1.6</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, P.-S. & Wang, X.-Y. (2001) Pteridophyte flora of Guizhou. Guizhou Science & Technology Press, Guiyang, 727 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, P.-S. & Pan, L.-T. (2018) Lycophytes and Ferns of Guizhou, (vol. II). Guizhou Science & Technology Press, Guiyang, 976 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, B., Sun, Q.-W., Wang, Z.-H., Chen, C.-L., Xu, W.-F. & Zhang, L.-B. (2022) Polystichum suiyangense (sect. Sphaenopolystichum, Dryopteridaceae), a new fern from Guizhou, China. Phytotaxa 567: 93–99.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wei, H.-J. & Zhang, L.-B. (2018) Polystichum recavum (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangxi, China. Phytotaxa 374: 167–171. https://doi.org/10.11646/phytotaxa.374.2.9</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhang, L., Zhu, Z.-M., Gao, X.-F. & Zhang, L.-B. (2012) Polystichum hubeiense (Dryopteridaceae), a new fern species from Hubei, China. Annales Botanici Fennici 50: 107–110. https://doi.org/10.5735/085.050.0121</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhang, L.-B. & Barrington, D.S. (2013) Polystichum. In: Wu, Z.-Y., Raven, P.H. & Hong, D.-Y. (Eds.) Flora of China, vols. 2–3. Science Press, Beijing & Missouri Botanical Garden Press, St. Louis, pp. 629–713.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhang, L.-B., Duan, Y.-F. & Kropf, M. (2019) Eight new species of the fern genus Polystichum (subg. Haplopolystichum; Dryopteridaceae) from limestone caves in Guizhou, China. Phytotaxa 404: 209–230. https://doi.org/10.11646/phytotaxa.404.6.1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhang, L.-B. & He, H. (2009) Polystichum peishanii, a new species known only from three populations on limestones in southern Guizhou, China. Botanical Studies 50: 101–106.</span></span></span>