Abstract
Begonia fritschiana from Mindanao Island is described as a new species endemic to the Philippines. This is the latest addition to the species-rich Begonia section Baryandra. It resembles Begonia caramoanensis but is distinguished by the consistent obliquely ovate leaves with shorter petioles, lamina glaucous abaxially and with marginal hairs, mostly solitary to singly-branched inflorescences with shorter white-glandular hairs on the pedicels and peduncle, one of the outer tepals pink, and glabrous ovary and ovary wings. Begonia fritschiana is known only from the type locality with one population and less than 100 individuals. The area is not currently protected under the country’s National Integrated Protected Areas System by the Department of Environment and Natural Resources. Because of ongoing threats such as illegal logging, poaching, and over collection of biological resources, the single area of occurrence with estimated extent < 10 km2, and the small number of < 100 individuals, we recommend a classification of B. fritschiana as Critically Endangered.
References
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Amoroso, V.B., Coritico, F.P. & Fritsch, P.W. (2020) <em>Actinostachys minuta</em>, a new species of grass fern from Mindanao, Philippines. <em>PhytoKeys</em> 151: 59–66. https://doi.org/10.3897/phytokeys.151.53100</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ang, Y.P., Aumentado, J.A. & Magtoto L.M. (2022) <em>Begonia naemma</em> (sect. <em>Petermannia</em>), a new species from Nueva Ecija, Luzon Island, the Philippines. <em>Edinburgh Journal of Botany</em> 79, <em>Begonia</em> special issue, Article 390: 1–10. https://doi.org/10.24823/EJB.2022.390 </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Blasco, F.A., Alejandro, G.J., Tandang, D.N., Vales, M.B. & Rubite, R.R. (2022) <em>Begonia amparoi</em> (Begoniaceae, section <em>Baryandra</em>) a new species from Linungaw Twin Islands, Surigao del Sur, Mindanao, Philippines. <em>Phytotaxa</em> 549 (2): 136–140. https://doi.org/10.11646/phytotaxa.549.2.2</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Coritico, F.P. & Fleischmann, A. (2016) The first record of the boreal bog species <em>Drosera rotundifolia</em> (Droseraceae) from the Philippines, and a key to the Philippine sundews. <em>Blumea</em> 61: 24–28. https://doi.org/10.3767/000651916X691330</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Coritico, F.P., Amoroso, V.B. & Lehnert, M. (2017) New records, names, and combinations of scaly tree ferns <em>(Cyatheaceae)</em> in eastern Malesia. <em>Blumea</em> 62 (5): 92–96. https://doi.org/10.3767/blumea.2017.62.02.03</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">de Candolle, A. (1859) Meìmoire sur la famille des Beìgoniaceìes. [ser. 4.] <em>Annales des Sciences Naturelles; Botanique </em>11: 128–129. </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Dela Cruz, C.J.P., Concepcion S.R. & Ang Y.P. (2022) <em>Begonia francisabuidii,</em> (section <em>Baryandra</em>, Begoniaceae) a new species endemic to Albay, Luzon Island, Philippines. <em>Taiwania </em>67 (2): 223‒228. https://doi.org/10.6165/tai.2022.67.223</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Delos Angeles M.D., Rubite R.R., Chung K.-F., Buot, Jr. I.E. & Tandang D.N. (2022) <em>Begonia normaaguilariae </em>(section <em>Baryandra</em>, Begoniaceae), a new species from the limestone forests of Samar Island, Philippines. <em>Phytotaxa </em>541 (1): 49–56. https://doi.org/10.11646/phytotaxa.541.1.4</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Elmer, A.D.E. (1915) Two Hundred Twenty Six New Species I. <em>Leaflets of Philippine Botany </em>7: 2543–2700.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hughes, M., Moonlight, P.W., Jara-MunÞoz, A., Tebbitt, M.C., Wilson, H.P. & Pullan, M. (2015–) Begonia Resource Centre. Online database available from [http://padme.rbge.org.uk/begonia/]</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">IUCN Standards and Petitions Committee (2022) <em>Guidelines for Using the IUCN Red List Categories and Criteria. Version 15. Prepared by the Standards and Petitions Committee.</em> Available from: https://www.iucnredlist.org/documents/RedListGuidelines.pdf (accessed 5 February 2023)</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Klotzsch, J.F. (1854) <em>Begoniaceen-Gattungen und Arten</em>. <em>Monatsberichte der Königlich preussischen Akademie der Wissenschaften zu Berlin </em>1854: 119–128.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Klotzsch, J.F. (1855) <em>Begoniaceen-Gattungen und Arten</em>. <em>Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin </em>1855: 121–25.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lagunday, N.E., Acma, F.M., Cabana, V.G., Sabas, N.M. & Amoroso, V.B. (2017) Two new <em>Nephenthes </em>species from the unexplored mountains of Central Mindanao, Philippines. <em>Philippine Journal of Science</em> 146 (2): 159–165.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lagunday N.E., Cabana, V.G., Armecin, H.C., Sabas, N.M., Escarlos, J.A. & Mohagan, A.B. (2019) Rodents and some non-rodent mammals of Mt. Tangkulan Range, Central Mindanao, Philippines. <em>Journal of Biodiversity and Environmental Sciences</em> 14 (1): 814–2019.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lagunday N.E., Dela Rosa S., Cleofei, C.M.B, Patanao, R. Jr., Coritico, F.P. & Amoroso, V.B. (2022) <em>Nepenthes candalaga</em> (Nepenthaceae), a new species from eastern Mindanao, Philippines. <em>Phytotaxa </em>554 (3): 285–292. https://doi.org/10.11646/phytotaxa.554.3.7</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Linnaeus, C. (1753) <em>Species plantarum</em>, vol .1. Salvius, Stockholm, 1200 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Mazo, K.R.M. & Rubite, R.R. (2022) Two new species of <em>Begonia</em> (section <em>Petermannia</em>, Begoniaceae) from the Zamboanga Peninsula, Philippines and a redescription of <em>Begonia</em> <em>parvilimba</em>. <em>Phytotaxa </em>538 (2): 163–171. https://doi.org/10.11646/phytotaxa.538.2.9</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Mazo K.R.F., Salatan N.L., Santos I.E.A. & Rubite, R.R. (2022) Two new species of <em>Begonia</em> (section <em>Petermannia</em>, Begoniaceae) from Zamboanga Peninsula, Philippines with notes on an amended description of <em>B. elatostematoides</em>. <em>Taiwania</em> 67(3): 441–449. https://doi.org/10.6165/tai.2022.67.441</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Merrill, E.D. (1912 [1911]) The Philippine species of <em>Begonia</em>. <em>Philippine Journal of Science, section C, Botany</em> 6: 369–406.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Merrill, E.D. (1918) New or noteworthy Philippine plants. <em>The Philippine Journal of Science section C, Botany</em> 13: 1–66.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Moonlight, P.W., Ardi, W.H., Padilla, L.A., Chung, K.-F., Fuller, D., Girmansyah, D., Hollands, R., Jara-MunÞoz, A., Kiew, R., Leong, W.-C., Liu, Y., Mahardika, A., Marasinghe, L.D.K., O’Connor, M., Peng, C.-I, Peìrez, A.J., Phutthai, T., Pullan, M., Rajbhandary, S., Reynel, C., Rubite, R.R., Sang, J., Scherberich, D., Shui, Y.-M., Tebbitt, M.C., Thomas, D.C., Wilson, H.P., Zaini, N.H. & Hughes, M. (2018) Dividing and conquering the fastest-growing genus: Towards a natural sectional classification of the mega-diverse genus <em>Begonia </em>(Begoniaceae). <em>Taxon </em>67: 267–323. https://doi.org/10.12705/672.3</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite R.R., Ubaldo D.B.H., Salcedo J.C., Chung K.F., Evangelista L.T., Tandang D.N. & Hughes M. (2022) <em>Begonia hemicardia</em> (sect. <em>Petermannia</em>, Begoniaceae), a resurrected heterotypic synonym and <em>nomen nudum</em>. <em>Edinburgh Journal of Botany</em> 79, <em>Begonia</em> special issue, Article 403: 1–7. https://doi.org/10.24823/EJB.2022.403</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R., Irabagon, M.L., Palacio, D.J.E., Ang, Y.P. & Bustamante, R.A.A. (2020) <em>Begonia caramoanensis</em> (sect. <em>Baryandra</em>, Begoniaceae) a new species from Camarines Sur, Philippines. <em>Phytotaxa</em> 439 (3): 287–294. https://doi.org/10.11646/phytotaxa.439.3.10</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rule, M.G.Q., Ang, Y.P., Rubite, R.R., Docot, R.V.A., Bustamante, R.A.A. & Robinson, A.S. (2020) <em>Begonia makuruyot </em>(Begoniaceae section <em>Baryandra</em>), a new species from Surigao del Norte Province, Philippines. <em>Phytotaxa </em>470: 226–234. https://doi.org/10.11646/phytotaxa.470.3.2 </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Steudel, E.G. von. (1821) <em>Nomenclator botanicus: enumerans ordine alphabetico nomina atque synonyma, tum generica tum specifica, et a linnaeo et recentioribus de re botanica scriptoribus plantis phanerogamis imposita</em>. Cottae, Stuttgartiae & Tubingae, 900 pp. </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Warburg, O. (1904) Begoniaceae. <em>In</em>: Perkins, J.R. (Ed.) <em>Fragmenta florae Philippinae: contributions to the flora of the Philippine Islands. </em>Leipzig, Germany, pp. 51–56. </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ang, Y.P., Aumentado, J.A. & Magtoto L.M. (2022) <em>Begonia naemma</em> (sect. <em>Petermannia</em>), a new species from Nueva Ecija, Luzon Island, the Philippines. <em>Edinburgh Journal of Botany</em> 79, <em>Begonia</em> special issue, Article 390: 1–10. https://doi.org/10.24823/EJB.2022.390 </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Blasco, F.A., Alejandro, G.J., Tandang, D.N., Vales, M.B. & Rubite, R.R. (2022) <em>Begonia amparoi</em> (Begoniaceae, section <em>Baryandra</em>) a new species from Linungaw Twin Islands, Surigao del Sur, Mindanao, Philippines. <em>Phytotaxa</em> 549 (2): 136–140. https://doi.org/10.11646/phytotaxa.549.2.2</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Coritico, F.P. & Fleischmann, A. (2016) The first record of the boreal bog species <em>Drosera rotundifolia</em> (Droseraceae) from the Philippines, and a key to the Philippine sundews. <em>Blumea</em> 61: 24–28. https://doi.org/10.3767/000651916X691330</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Coritico, F.P., Amoroso, V.B. & Lehnert, M. (2017) New records, names, and combinations of scaly tree ferns <em>(Cyatheaceae)</em> in eastern Malesia. <em>Blumea</em> 62 (5): 92–96. https://doi.org/10.3767/blumea.2017.62.02.03</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">de Candolle, A. (1859) Meìmoire sur la famille des Beìgoniaceìes. [ser. 4.] <em>Annales des Sciences Naturelles; Botanique </em>11: 128–129. </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Dela Cruz, C.J.P., Concepcion S.R. & Ang Y.P. (2022) <em>Begonia francisabuidii,</em> (section <em>Baryandra</em>, Begoniaceae) a new species endemic to Albay, Luzon Island, Philippines. <em>Taiwania </em>67 (2): 223‒228. https://doi.org/10.6165/tai.2022.67.223</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Delos Angeles M.D., Rubite R.R., Chung K.-F., Buot, Jr. I.E. & Tandang D.N. (2022) <em>Begonia normaaguilariae </em>(section <em>Baryandra</em>, Begoniaceae), a new species from the limestone forests of Samar Island, Philippines. <em>Phytotaxa </em>541 (1): 49–56. https://doi.org/10.11646/phytotaxa.541.1.4</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Elmer, A.D.E. (1915) Two Hundred Twenty Six New Species I. <em>Leaflets of Philippine Botany </em>7: 2543–2700.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hughes, M., Moonlight, P.W., Jara-MunÞoz, A., Tebbitt, M.C., Wilson, H.P. & Pullan, M. (2015–) Begonia Resource Centre. Online database available from [http://padme.rbge.org.uk/begonia/]</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">IUCN Standards and Petitions Committee (2022) <em>Guidelines for Using the IUCN Red List Categories and Criteria. Version 15. Prepared by the Standards and Petitions Committee.</em> Available from: https://www.iucnredlist.org/documents/RedListGuidelines.pdf (accessed 5 February 2023)</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Klotzsch, J.F. (1854) <em>Begoniaceen-Gattungen und Arten</em>. <em>Monatsberichte der Königlich preussischen Akademie der Wissenschaften zu Berlin </em>1854: 119–128.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Klotzsch, J.F. (1855) <em>Begoniaceen-Gattungen und Arten</em>. <em>Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin </em>1855: 121–25.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lagunday, N.E., Acma, F.M., Cabana, V.G., Sabas, N.M. & Amoroso, V.B. (2017) Two new <em>Nephenthes </em>species from the unexplored mountains of Central Mindanao, Philippines. <em>Philippine Journal of Science</em> 146 (2): 159–165.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lagunday N.E., Cabana, V.G., Armecin, H.C., Sabas, N.M., Escarlos, J.A. & Mohagan, A.B. (2019) Rodents and some non-rodent mammals of Mt. Tangkulan Range, Central Mindanao, Philippines. <em>Journal of Biodiversity and Environmental Sciences</em> 14 (1): 814–2019.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lagunday N.E., Dela Rosa S., Cleofei, C.M.B, Patanao, R. Jr., Coritico, F.P. & Amoroso, V.B. (2022) <em>Nepenthes candalaga</em> (Nepenthaceae), a new species from eastern Mindanao, Philippines. <em>Phytotaxa </em>554 (3): 285–292. https://doi.org/10.11646/phytotaxa.554.3.7</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Linnaeus, C. (1753) <em>Species plantarum</em>, vol .1. Salvius, Stockholm, 1200 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Mazo, K.R.M. & Rubite, R.R. (2022) Two new species of <em>Begonia</em> (section <em>Petermannia</em>, Begoniaceae) from the Zamboanga Peninsula, Philippines and a redescription of <em>Begonia</em> <em>parvilimba</em>. <em>Phytotaxa </em>538 (2): 163–171. https://doi.org/10.11646/phytotaxa.538.2.9</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Mazo K.R.F., Salatan N.L., Santos I.E.A. & Rubite, R.R. (2022) Two new species of <em>Begonia</em> (section <em>Petermannia</em>, Begoniaceae) from Zamboanga Peninsula, Philippines with notes on an amended description of <em>B. elatostematoides</em>. <em>Taiwania</em> 67(3): 441–449. https://doi.org/10.6165/tai.2022.67.441</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Merrill, E.D. (1912 [1911]) The Philippine species of <em>Begonia</em>. <em>Philippine Journal of Science, section C, Botany</em> 6: 369–406.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Merrill, E.D. (1918) New or noteworthy Philippine plants. <em>The Philippine Journal of Science section C, Botany</em> 13: 1–66.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Moonlight, P.W., Ardi, W.H., Padilla, L.A., Chung, K.-F., Fuller, D., Girmansyah, D., Hollands, R., Jara-MunÞoz, A., Kiew, R., Leong, W.-C., Liu, Y., Mahardika, A., Marasinghe, L.D.K., O’Connor, M., Peng, C.-I, Peìrez, A.J., Phutthai, T., Pullan, M., Rajbhandary, S., Reynel, C., Rubite, R.R., Sang, J., Scherberich, D., Shui, Y.-M., Tebbitt, M.C., Thomas, D.C., Wilson, H.P., Zaini, N.H. & Hughes, M. (2018) Dividing and conquering the fastest-growing genus: Towards a natural sectional classification of the mega-diverse genus <em>Begonia </em>(Begoniaceae). <em>Taxon </em>67: 267–323. https://doi.org/10.12705/672.3</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite R.R., Ubaldo D.B.H., Salcedo J.C., Chung K.F., Evangelista L.T., Tandang D.N. & Hughes M. (2022) <em>Begonia hemicardia</em> (sect. <em>Petermannia</em>, Begoniaceae), a resurrected heterotypic synonym and <em>nomen nudum</em>. <em>Edinburgh Journal of Botany</em> 79, <em>Begonia</em> special issue, Article 403: 1–7. https://doi.org/10.24823/EJB.2022.403</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R., Irabagon, M.L., Palacio, D.J.E., Ang, Y.P. & Bustamante, R.A.A. (2020) <em>Begonia caramoanensis</em> (sect. <em>Baryandra</em>, Begoniaceae) a new species from Camarines Sur, Philippines. <em>Phytotaxa</em> 439 (3): 287–294. https://doi.org/10.11646/phytotaxa.439.3.10</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rule, M.G.Q., Ang, Y.P., Rubite, R.R., Docot, R.V.A., Bustamante, R.A.A. & Robinson, A.S. (2020) <em>Begonia makuruyot </em>(Begoniaceae section <em>Baryandra</em>), a new species from Surigao del Norte Province, Philippines. <em>Phytotaxa </em>470: 226–234. https://doi.org/10.11646/phytotaxa.470.3.2 </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Steudel, E.G. von. (1821) <em>Nomenclator botanicus: enumerans ordine alphabetico nomina atque synonyma, tum generica tum specifica, et a linnaeo et recentioribus de re botanica scriptoribus plantis phanerogamis imposita</em>. Cottae, Stuttgartiae & Tubingae, 900 pp. </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Warburg, O. (1904) Begoniaceae. <em>In</em>: Perkins, J.R. (Ed.) <em>Fragmenta florae Philippinae: contributions to the flora of the Philippine Islands. </em>Leipzig, Germany, pp. 51–56. </span></span></span></p>