Abstract
Article 35.2 of the Shenzhen Code requires that, for a combination to be validly published, authors of plant names must: “[…] definitely associate the final epithet with the name of the genus or species, or with its abbreviation […]”. Correct application of this Article indicates that neither “Kalanchoe sect. Alatae” nor “Kalanchoe subg. Alatae”, was validly published by, respectively, Raymond-Hamet in 1933 and Smith et al. in 2021. However, Kalanchoe sect. Alatae was validly published by Smith in 2021. The new status, at the rank of subgenus, Kalanchoe subg. Alatae, is validly published here.
References
<p>Adanson, M. (1763 [July–August]) <em>Familles des plantes. Contenant une Préface Historike fur l’état ancien & actuel de la Botanike, & une Téorie de cette Science</em>. II. Partie. Chez Vincent, Imprimeur-Libraire de M<sup>gr</sup> le Comte de Provence, rue S, Severin, Paris, 640 pp. [https://www.biodiversitylibrary.org/item/6958#page/794/mode/1up]</p>
<p>Baillon, M.H. (1885 [séance du 18 février]) Liste des plantes de Madagascar (suite de la page 464). [<em>Kalanchoe</em> § <em>Kitchingia</em>]. <em>Bulletin Mensuel de la Société Linnéenne de Paris (Paris)</em> 1 (59): 465–472. [https://www.biodiversitylibrary.org/item/41447#page/60/mode/1up]</p>
<p>Baker, J.G. (1881 [21 February]) Notes on a collection of flowering plants made by L. Kitching, Esq., in Madagascar in 1879 (Plates VII. & VIII.). <em>Journal of the Linnean Society, Botany</em> 18: 264–281. https://doi.org/10.1111/j.1095-8339.1881.tb01257.x</p>
<p>Baker, J.G. (1883 [1884 published 24 March 1883]) Contributions to the flora of Madagascar.—Part I. Polypetalæ [Plates XXII & XXIII]. <em>Journal of the Linnean Society, Botany</em> (<em>London</em>) 20 (126): 87–158. https://doi.org/10.1111/j.1095-8339.1883.tb00195.x</p>
<p>Berger, A. (1930 [Mai]) Crassulaceae. Unterfam. II. Kalanchoideae [sic, Kalanchooideae]. 6. <em>Kalanchoe</em> Adanson. 7. <em>Kitchingia</em> Baker. 8. <em>Bryophyllum</em> Salisb. <em>In</em>: Engler, A. & Prantl, K. (Eds.) <em>Die natürlichen Pflanzenfamilien</em>. ed. 2, 18a. Verlag von Wilhelm Engelmann, Leipzig, pp. 402–412.</p>
<p>Boiteau, P. (1947 [novembre-décembre]). Les plantes grasses de Madagascar. [Overview of sections in <em>Kalanchoe</em>.] <em>Cactus (Paris)</em> 1947 (12): 5–10, figures 1–4.</p>
<p>Boiteau, P. & Mannoni, O. (1948 [octobre-décembre]) Les <em>Kalanchoe</em> (suite). [§ 4. <em>Epidendrae</em>; <em>Kalanchoe porphyrocalyx</em> (Baker) Baill. and <em>Kalanchoe uniflora</em> Raym.-Hamet (in the key).] (à suivre). <em>Cactus</em> <em>(Paris)</em> 17–18: 57–58 + two unnumbered figures.</p>
<p>Boiteau, P. & Mannoni, O. (1949 [janvier-mars]) Les <em>Kalanchoe</em> (suite). [<em>Kalanchoe uniflora</em> Raym.-Hamet; § 5. <em>Scandentes</em>; § (unnumbered) <em>Bulbilliferae</em>.] (à suivre). <em>Cactus</em> <em>(Paris)</em> 1949 (19): 9–14, + seven unnumbered figures.</p>
<p>Dauphiné, A. & Hamet, R. (1911) Contribution a l’étude anatomique du genre <em>Kalanchoe</em>. <em>Annales des Sciences Naturelles, Botanique</em>, neuvième série XIV (N<sup>os</sup> 4 à 6): 195–219. [https://www.biodiversitylibrary.org/item/155873#page/7/mode/1up]</p>
<p>Hamet, R. (1910 [Janvier]) Sur quelques <em>Kalanchoe</em> peu connus (Suite). <em>Bulletin de la Société Botanique de France</em> [Tome Cinquante-Septième], Quatrième serie—Tome X) Quatrième series 57: 49–54. https://doi.org/10.1080/00378941.1910.10832165</p>
<p>Koorders, S.H. (1919 [Mai] [24 October 1918]) Beitrag zur Kenntniss der Flora von Java No. 18. Beschreibung von <em>Kalanchoë Schumacheri</em> vom Idjen-Plateau und Revision der Crassulaceae von Java. (Mit 2 lithographischen Tafeln.). <em>Bulletin du Jardin Botanique de Buitenzorg</em> sér. 3, 1 (3): 169–180, Figures 14 and 15. [https://www.biodiversitylibrary.org/item/27173#page/205/mode/1up] (text) [https://www.biodiversitylibrary.org/item/27173#page/243/mode/1up] (figures)</p>
<p>Raymond-Hamet (1933) Observations sur le <em>Kalanchoe uniflora</em> (Stapf) Raymond-Hamet. <em>Bulletin de la Société Botanique de France</em> 80 (4): 542–548. https://doi.org/10.1080/00378941.1933.10833878</p>
<p>Raymond-Hamet (1954a [juillet-août-septembre]) Sur un <em>Kalanchoe</em> Malgache dont les fleurs étaient encore inconnues. [<em>Kalanchoe rhombopilosa</em> Mannoni & Boiteau.] <em>Journal d’Agriculture Traditionnelle et de Botanique Appliquée</em> 1 (7–9): 281–292. https://doi.org/10.3406/jatba.1954.2160</p>
<p>Raymond-Hamet (1954b [décembre]) Sur le <em>Kalanchoe rhombopilosa</em> Mannoni et Boiteau. <em>Bulletin du Jardin Botanique de l’État a Bruxelles</em> 24 (4): 329–333. https://doi.org/10.2307/3667043</p>
<p>Salisbury, R.A. (1805 [1 June 1805]) <em>The Paradisus Londinensis: or coloured figures of plants cultivated in the vicinity of the metropolis</em> 1. Published by William Hooker, No. 6, Frith Street, city not stated, presumably London, 157 pp. & 117 plates. https://doi.org/10.5962/bhl.title.53520</p>
<p>Smith, G.F. (2020) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). <em>Bradleya</em> 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15</p>
<p>Smith, G.F. (2021) Reinstatement of <em>Kalanchoe</em> [subg. <em>Bryophyllum</em>] sect. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae) for <em>K. porphyrocalyx</em> and <em>K. uniflora</em>, two Madagascan endemics. <em>Bradleya</em> 39: 177–181. https://doi.org/10.25223/brad.n39.2021.a17</p>
<p>Smith, G.F. (2022) Notes on <em>Kalanchoe</em> subg. <em>Bryophyllum</em> (Crassulaceae subfam. Kalanchooideae) and the kalanchoes of Sijfert H. Koorders (1919). <em>Phytotaxa</em> 549 (1): 87–96. https://doi.org/10.11646/phytotaxa.549.1.7</p>
<p>Smith, G.F. & Figueiredo, E. (2018) The infrageneric classification and nomenclature of <em>Kalanchoe </em>Adans. (Crassulaceae), with special reference to the southern African species. <em>Bradleya </em>36: 162–172. https://doi.org/10.25223/brad.n36.2018.a10</p>
<p>Smith, G.F. & Figueiredo, E. (2019) <em>Kalanchoe porphyrocalyx</em> var. <em>sambiranensis</em>, a new variety in a Madagascan <em>Kalanchoe</em> (Crassulaceae subfam. Kalanchooideae). <em>Phytotaxa</em> 422 (2): 206–208. https://doi.org/10.11646/phytotaxa.422.2.8</p>
<p>Smith, G.F., Figueiredo, E. & Van Wyk, A.E. (2019) <em>Kalanchoe (Crassulaceae) in southern Africa. Classification, biology, and cultivation</em>. Academic Press, an imprint of Elsevier, London, San Diego, Cambridge (USA), and Oxford, 328 pp. https://doi.org/10.1016/C2017-0-00602-X</p>
<p>Smith, G.F., Klein, D.-P. & Shtein, R. (2021a) A revision of the Malagasy <em>Kalanchoe</em> subg. <em>Kitchingia</em> (Crassulaceae subfam. Kalanchooideae): history, taxonomy, and nomenclature. <em>Phytotaxa</em> 507 (1): 67–80. https://doi.org/10.11646/phytotaxa.507.1.3</p>
<p>Smith, G.F. & Monro, A.M. (2022a) The nomenclature and taxonomy of Crassulaceae subfam. Kalanchooideae resolved. <em>Phytotaxa</em> 555 (3): 241–251. https://doi.org/10.11646/phytotaxa.555.3.3</p>
<p>Smith, G.F. & Monro, A.M. (2022b) (2908) Proposal to reject the name Crassulaceae subfam. Cotyledonoideae (Crassulaceae). <em>Taxon</em> 71 (4): 910–911. https://doi.org/10.1002/tax.12778</p>
<p>Smith, G.F., Shtein, R. & Klein, D.-P. (2021b) <em>Kalanchoe</em> subg. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae), a new rank for <em>K</em>. sect. <em>Alatae</em>. <em>Phytotaxa</em> 509 (2): 255–258. https://doi.org/10.11646/phytotaxa.509.2.9</p>
<p>Stapf, O. (1908) 943. <em>Kitchingia uniflora Stapf [Crassulaceae]. In: XXXIV. Diagnoses africanae: XXIII, pp. 257–262</em>. Bulletin of Miscellaneous Information, Royal Gardens, Kew, pp. 258–259. https://doi.org/10.2307/4111576</p>
<p>Tillson, A.H. (1940) The floral anatomy of the Kalanchoideae [sic, Kalanchooideae]. <em>American Journal of Botany</em> 27 (8): 595–600. https://doi.org/10.2307/2436977</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) <em>International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. Koeltz Botanical Books, Glashütten. [Regnum Vegetabile 159], 254 pp. https://doi.org/10.12705/Code.2018</p>
<p>Baillon, M.H. (1885 [séance du 18 février]) Liste des plantes de Madagascar (suite de la page 464). [<em>Kalanchoe</em> § <em>Kitchingia</em>]. <em>Bulletin Mensuel de la Société Linnéenne de Paris (Paris)</em> 1 (59): 465–472. [https://www.biodiversitylibrary.org/item/41447#page/60/mode/1up]</p>
<p>Baker, J.G. (1881 [21 February]) Notes on a collection of flowering plants made by L. Kitching, Esq., in Madagascar in 1879 (Plates VII. & VIII.). <em>Journal of the Linnean Society, Botany</em> 18: 264–281. https://doi.org/10.1111/j.1095-8339.1881.tb01257.x</p>
<p>Baker, J.G. (1883 [1884 published 24 March 1883]) Contributions to the flora of Madagascar.—Part I. Polypetalæ [Plates XXII & XXIII]. <em>Journal of the Linnean Society, Botany</em> (<em>London</em>) 20 (126): 87–158. https://doi.org/10.1111/j.1095-8339.1883.tb00195.x</p>
<p>Berger, A. (1930 [Mai]) Crassulaceae. Unterfam. II. Kalanchoideae [sic, Kalanchooideae]. 6. <em>Kalanchoe</em> Adanson. 7. <em>Kitchingia</em> Baker. 8. <em>Bryophyllum</em> Salisb. <em>In</em>: Engler, A. & Prantl, K. (Eds.) <em>Die natürlichen Pflanzenfamilien</em>. ed. 2, 18a. Verlag von Wilhelm Engelmann, Leipzig, pp. 402–412.</p>
<p>Boiteau, P. (1947 [novembre-décembre]). Les plantes grasses de Madagascar. [Overview of sections in <em>Kalanchoe</em>.] <em>Cactus (Paris)</em> 1947 (12): 5–10, figures 1–4.</p>
<p>Boiteau, P. & Mannoni, O. (1948 [octobre-décembre]) Les <em>Kalanchoe</em> (suite). [§ 4. <em>Epidendrae</em>; <em>Kalanchoe porphyrocalyx</em> (Baker) Baill. and <em>Kalanchoe uniflora</em> Raym.-Hamet (in the key).] (à suivre). <em>Cactus</em> <em>(Paris)</em> 17–18: 57–58 + two unnumbered figures.</p>
<p>Boiteau, P. & Mannoni, O. (1949 [janvier-mars]) Les <em>Kalanchoe</em> (suite). [<em>Kalanchoe uniflora</em> Raym.-Hamet; § 5. <em>Scandentes</em>; § (unnumbered) <em>Bulbilliferae</em>.] (à suivre). <em>Cactus</em> <em>(Paris)</em> 1949 (19): 9–14, + seven unnumbered figures.</p>
<p>Dauphiné, A. & Hamet, R. (1911) Contribution a l’étude anatomique du genre <em>Kalanchoe</em>. <em>Annales des Sciences Naturelles, Botanique</em>, neuvième série XIV (N<sup>os</sup> 4 à 6): 195–219. [https://www.biodiversitylibrary.org/item/155873#page/7/mode/1up]</p>
<p>Hamet, R. (1910 [Janvier]) Sur quelques <em>Kalanchoe</em> peu connus (Suite). <em>Bulletin de la Société Botanique de France</em> [Tome Cinquante-Septième], Quatrième serie—Tome X) Quatrième series 57: 49–54. https://doi.org/10.1080/00378941.1910.10832165</p>
<p>Koorders, S.H. (1919 [Mai] [24 October 1918]) Beitrag zur Kenntniss der Flora von Java No. 18. Beschreibung von <em>Kalanchoë Schumacheri</em> vom Idjen-Plateau und Revision der Crassulaceae von Java. (Mit 2 lithographischen Tafeln.). <em>Bulletin du Jardin Botanique de Buitenzorg</em> sér. 3, 1 (3): 169–180, Figures 14 and 15. [https://www.biodiversitylibrary.org/item/27173#page/205/mode/1up] (text) [https://www.biodiversitylibrary.org/item/27173#page/243/mode/1up] (figures)</p>
<p>Raymond-Hamet (1933) Observations sur le <em>Kalanchoe uniflora</em> (Stapf) Raymond-Hamet. <em>Bulletin de la Société Botanique de France</em> 80 (4): 542–548. https://doi.org/10.1080/00378941.1933.10833878</p>
<p>Raymond-Hamet (1954a [juillet-août-septembre]) Sur un <em>Kalanchoe</em> Malgache dont les fleurs étaient encore inconnues. [<em>Kalanchoe rhombopilosa</em> Mannoni & Boiteau.] <em>Journal d’Agriculture Traditionnelle et de Botanique Appliquée</em> 1 (7–9): 281–292. https://doi.org/10.3406/jatba.1954.2160</p>
<p>Raymond-Hamet (1954b [décembre]) Sur le <em>Kalanchoe rhombopilosa</em> Mannoni et Boiteau. <em>Bulletin du Jardin Botanique de l’État a Bruxelles</em> 24 (4): 329–333. https://doi.org/10.2307/3667043</p>
<p>Salisbury, R.A. (1805 [1 June 1805]) <em>The Paradisus Londinensis: or coloured figures of plants cultivated in the vicinity of the metropolis</em> 1. Published by William Hooker, No. 6, Frith Street, city not stated, presumably London, 157 pp. & 117 plates. https://doi.org/10.5962/bhl.title.53520</p>
<p>Smith, G.F. (2020) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). <em>Bradleya</em> 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15</p>
<p>Smith, G.F. (2021) Reinstatement of <em>Kalanchoe</em> [subg. <em>Bryophyllum</em>] sect. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae) for <em>K. porphyrocalyx</em> and <em>K. uniflora</em>, two Madagascan endemics. <em>Bradleya</em> 39: 177–181. https://doi.org/10.25223/brad.n39.2021.a17</p>
<p>Smith, G.F. (2022) Notes on <em>Kalanchoe</em> subg. <em>Bryophyllum</em> (Crassulaceae subfam. Kalanchooideae) and the kalanchoes of Sijfert H. Koorders (1919). <em>Phytotaxa</em> 549 (1): 87–96. https://doi.org/10.11646/phytotaxa.549.1.7</p>
<p>Smith, G.F. & Figueiredo, E. (2018) The infrageneric classification and nomenclature of <em>Kalanchoe </em>Adans. (Crassulaceae), with special reference to the southern African species. <em>Bradleya </em>36: 162–172. https://doi.org/10.25223/brad.n36.2018.a10</p>
<p>Smith, G.F. & Figueiredo, E. (2019) <em>Kalanchoe porphyrocalyx</em> var. <em>sambiranensis</em>, a new variety in a Madagascan <em>Kalanchoe</em> (Crassulaceae subfam. Kalanchooideae). <em>Phytotaxa</em> 422 (2): 206–208. https://doi.org/10.11646/phytotaxa.422.2.8</p>
<p>Smith, G.F., Figueiredo, E. & Van Wyk, A.E. (2019) <em>Kalanchoe (Crassulaceae) in southern Africa. Classification, biology, and cultivation</em>. Academic Press, an imprint of Elsevier, London, San Diego, Cambridge (USA), and Oxford, 328 pp. https://doi.org/10.1016/C2017-0-00602-X</p>
<p>Smith, G.F., Klein, D.-P. & Shtein, R. (2021a) A revision of the Malagasy <em>Kalanchoe</em> subg. <em>Kitchingia</em> (Crassulaceae subfam. Kalanchooideae): history, taxonomy, and nomenclature. <em>Phytotaxa</em> 507 (1): 67–80. https://doi.org/10.11646/phytotaxa.507.1.3</p>
<p>Smith, G.F. & Monro, A.M. (2022a) The nomenclature and taxonomy of Crassulaceae subfam. Kalanchooideae resolved. <em>Phytotaxa</em> 555 (3): 241–251. https://doi.org/10.11646/phytotaxa.555.3.3</p>
<p>Smith, G.F. & Monro, A.M. (2022b) (2908) Proposal to reject the name Crassulaceae subfam. Cotyledonoideae (Crassulaceae). <em>Taxon</em> 71 (4): 910–911. https://doi.org/10.1002/tax.12778</p>
<p>Smith, G.F., Shtein, R. & Klein, D.-P. (2021b) <em>Kalanchoe</em> subg. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae), a new rank for <em>K</em>. sect. <em>Alatae</em>. <em>Phytotaxa</em> 509 (2): 255–258. https://doi.org/10.11646/phytotaxa.509.2.9</p>
<p>Stapf, O. (1908) 943. <em>Kitchingia uniflora Stapf [Crassulaceae]. In: XXXIV. Diagnoses africanae: XXIII, pp. 257–262</em>. Bulletin of Miscellaneous Information, Royal Gardens, Kew, pp. 258–259. https://doi.org/10.2307/4111576</p>
<p>Tillson, A.H. (1940) The floral anatomy of the Kalanchoideae [sic, Kalanchooideae]. <em>American Journal of Botany</em> 27 (8): 595–600. https://doi.org/10.2307/2436977</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) <em>International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. Koeltz Botanical Books, Glashütten. [Regnum Vegetabile 159], 254 pp. https://doi.org/10.12705/Code.2018</p>