Abstract
The French botanist Louis-René ‘Edmond’ Tulasne validly published three new names in Kalanchoe (Crassulaceae subfam. Kalanchooideae) in 1857. These are K. eriophylla, K. miniata, and K. floribunda, with the first two of these names being based on material that was collected more than 30 years earlier in Madagascar by Carl Hilsenberg and Wenceslas Bojer, who are jointly credited as having made the first significant plant collections in the interior of the island. The third name, K. floribunda, was published for a species from the Comoros, but is an illegitimate later homonym, for which a lectotype is designated here. The fourth species that Tulasne dealt with in his treatment of Malagasy kalanchoes was included under the name Bryophyllum calycinum, which is now regarded as a synonym of K. pinnata. In the past, Bryophyllum was sometimes accepted as a genus distinct from Kalanchoe, but at present it is treated as K. subg. Bryophyllum. The nomenclature of the three names published in Kalanchoe and the one treated in Bryophyllum by Tulasne is reviewed and discussed in a historical context. The type of the name K. tubiflora, which was collected during a British expedition that lasted from 1822 to 1826 to South and East Africa, is clarified.
References
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Figueiredo, E. & Smith, G.F. (2022) Notes on the itinerary, fleet, and collectors of botanical specimens during the 1822–1826 expedition of William Fitz William Owen to South and East Africa, and beyond. Phytotaxa 568 (2): 170–190. https://doi.org/10.11646/phytotaxa.568.2.3
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Hamet, R. (1912) Observations sur le Kalanchoe tubiflora nom. nov. Beihefte zum Botanischen Centralblatt 29: 41–44. [https://www.biodiversitylibrary.org/page/4986270#page/523/mode/1up]
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Mabberley, D.J. (2020) Botanical revelation. European encounters with Australian plants before Darwin. [The Peter Crossing Collection.] Corrected reprint. NewSouth Publishing, University of New South Wales Press Ltd, Sydney, Australia, 372 pp.
Natural History Museum (2022) Natural History Museum Archives Catalogue. Available from: https://www.nhm.ac.uk/CalmView/Record.aspx?src=CalmView.Catalog&id=DF+BOT (accessed 31 December 2022)
Oken, L. (1841) Allgemeine Naturgeschichte für alle Stande, von Professor Oken. Vol. 3(3). Hoffmann’sche Verlags-Buchhandlung, Stuttgart, pp. 1459–2142; Index (“Register”): pp. 1–44. https://doi.org/10.5962/bhl.title.100031
Owen, W.F. (1833) Narrative of voyages to explore the shores of Africa, Arabia, and Madagascar; performed in H.M. ships Leven and Barracouta, under the direction of Capt. W. F. W. Owen, R.N. Edited by H. B. Robinson. Published in two vols. J. & J. Harper, New York, 259 pp. and 240 pp., respectively.
Perrier de la Bâthie, [J.M.]H.[A.] (1924 [1922–1923]) Les Crassulacées Malgaches. [Observations sur quelques espèces du genre Kalanchoe, pp. 21–24; Crassulacées Malgaches nouvelles, pp. 24–26; Stations et distribution géographique, pp. 26–28; Synopsis des Crassulacées Malgaches pp. 29–36.] Bulletin de l’Académie Malgache [Nouvelle série Tome VI] 6: 21–36. [https://www.biodiversitylibrary.org/item/50784#page/80/mode/1up]
Perrier de la Bâthie, [J.M.]H.[A.] (1928 [Février]) Observations nouvelles sur le genre Kalanchoe. Archives de Botanique II, Bulletin Mensuel. Caen 2 (2): 17–31.
Persoon, C.H. (1805 [1 April–15 June]) Synopsis plantarum, seu enchiridium botanicum, complectens enumerationem systematicam specierum hucusque cognitorum. Pars prima. Apud Carol. Frid. Cramerum, Parisiis lutetiorum [Paris] et apud J.G. Cottam, Tubingæ [London], 546 pp. https://doi.org/10.5962/bhl.title.638
Raymond-Hamet (1916) Sur un groupe de transition reliant le genre Kalanchoe au genre Cotyledon. Revue Générale de Botanique. Paris XXVIII: 80–84. [https://www.biodiversitylibrary.org/item/45180#page/83/mode/1up]
Raymond-Hamet (1933) Observations sur le Kalanchoe uniflora (Stapf) Raymond-Hamet. Bulletin de la Société Botanique de France 80 (4): 542–548. https://doi.org/10.1080/00378941.1933.10833878
Salisbury, R.A. (1805 [1 June 1805]) The Paradisus Londinensis: or coloured figures of plants cultivated in the vicinity of the metropolis 1. Published by William Hooker, No. 6, Frith Street, city not stated, presumably London, 157 pp. & 117 plates. https://doi.org/10.5962/bhl.title.53520
Shtein, R., Smith, G.F. & Ikeda, J. (2021) Aspects of the taxonomy of the Kalanchoe daigremontiana species complex (Crassulaceae subfam. Kalanchooideae) and associated interspecific hybrids in southern Madagascar, with the description of a new nothospecies, K. ×descoingsii (=K. laetivirens × K. tubiflora). Phytotaxa 524 (4): 235–260. https://doi.org/10.11646/phytotaxa.524.4.2
Smith, G.F. (2020a) Kalanchoe Adans. subg. Fernandesiae Gideon F.Sm. (Crassulaceae subfam. Kalanchooideae), a new subgenus from southern and south-tropical Africa. Haseltonia 27: 2–7. https://doi.org/10.2985/026.027.0101
Smith, G.F. (2020b) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). Bradleya 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15
Smith, G.F. (2021) Reinstatement of Kalanchoe [subg. Bryophyllum] sect. Alatae (Crassulaceae subfam. Kalanchooideae) for K. porphyrocalyx and K. uniflora, two Madagascan endemics. Bradleya 39: 177–181. https://doi.org/10.25223/brad.n39.2021.a17
Smith, G.F. (2022a) Notes on Kalanchoe subg. Bryophyllum (Crassulaceae subfam. Kalanchooideae) and the kalanchoes of Sijfert H. Koorders (1919). Phytotaxa 549 (1): 87–96. https://doi.org/10.11646/phytotaxa.549.1.7
Smith, G.F. (2022b) Kalanchoe sect. Raveta (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1916): a long-overlooked name in K. subg. Kalanchoe. Phytotaxa 550 (2): 208–212. https://doi.org/10.11646/phytotaxa.550.2.10
Smith, G.F. (2022c) Date and place of publication and author attribution of the combination Kalanchoe sect. Raveta (Crassulaceae subfam. Kalanchooideae). Phytotaxa 560 (1): 131–134. https://doi.org/10.11646/phytotaxa.560.1.12
Smith, G.F. (2023) A reassessment of combinations previously proposed or published for Kalanchoe subg. Alatae (Crassulaceae subfam. Kalanchooideae) and its valid publication at the rank of subgenus. Phytotaxa. [in press]
Smith, G.F. & Figueiredo, E. (2018a) Nomenclatural notes on Kalanchoe pinnata (Lam.) Pers. (Crassulaceae). Bradleya 36: 220–223. https://doi.org/10.25223/brad.n36.2018.a18
Smith, G.F. & Figueiredo, E. (2018b) The infrageneric classification and nomenclature of Kalanchoe Adans. (Crassulaceae), with special reference to the southern African species. Bradleya 36: 162–172. https://doi.org/10.25223/brad.n36.2018.a10
Smith, G.F. & Figueiredo, E. (2021) The taxonomy and nomenclature of Kalanchoe streptantha Baker (Crassulaceae subfam. Kalanchooideae). Bradleya 39: 195–201. https://doi.org/10.25223/brad.n39.2021.a20
Smith, G.F. & Figueiredo, E. (2022a) Kalanchoe beharensis ‘Pixel’ (Crassulaceae subfam. Kalanchooideae), eine neue Sorte einer populären Zierpflanze aus Madagaskar. Kakteen und andere Sukkulenten 73 (11): 329–333.
Smith, G.F. & Figueiredo, E. (2022b) Clarifying the typification of the name Kalanchoe orgyalis (Crassulaceae subfam. Kalanchooideae). Phytotaxa 543 (1): 99–102. https://doi.org/10.11646/phytotaxa.543.1.11
Smith, G.F. & Figueiredo, E. (2023) Kalanchoe orgyalis. Crassulaceae subfam. Kalanchooideae. Madagascar. Flowering Plants of Africa 68. [in press]
Smith, G.F., Figueiredo, E. & Van Wyk, A.E. (2019) Kalanchoe (Crassulaceae) in southern Africa. Classification, biology, and cultivation. Academic Press, an imprint of Elsevier, London, San Diego, Cambridge (U.S.A.), and Oxford, 328 pp. https://doi.org/10.1016/C2017-0-00602-X
Smith, G.F., Klein, D.-P. & Shtein, R. (2021c) A revision of the Malagasy Kalanchoe subg. Kitchingia (Crassulaceae subfam. Kalanchooideae): history, taxonomy, and nomenclature. Phytotaxa 507 (1): 67–80. https://doi.org/10.11646/phytotaxa.507.1.3
Smith, G.F. & Monro, A.M. (2022a) The nomenclature and taxonomy of Crassulaceae subfam. Kalanchooideae resolved. Phytotaxa 555 (3): 241–251. https://doi.org/10.11646/phytotaxa.555.3.3
Smith, G.F. & Monro, A.M. (2022b) (2908) Proposal to reject the name Crassulaceae subfam. Cotyledonoideae (Crassulaceae). Taxon 71(4): 910–911. https://doi.org/10.1002/tax.12778
Smith, G.F. & Shtein, R. (2021) Nomenclature and taxonomic placement of the infraspecific taxa recognised in the Madagascan Kalanchoe miniata (Crassulaceae subfam. Kalanchooideae) by Perrier de la Bâthie and Boiteau and Mannoni in the 20<sup>th</sup> century. Phytotaxa 514 (2): 118–128. https://doi.org/10.11646/phytotaxa.514.2.3
Smith, G.F., Shtein, R. & Klein, D.-P. (2021b) Kalanchoe subg. Alatae (Crassulaceae subfam. Kalanchooideae), a new rank for K. sect. Alatae. Phytotaxa 509(2): 255–258. https://doi.org/10.11646/phytotaxa.509.2.9
Smith, G.F., Shtein, R., Klein, D.-P., Parihar, B., Almeida, A., Rodewald, S. & Kadereit, G. (2021a) Special Review. Sexual and asexual reproduction in Kalanchoe (Crassulaceae): a review of known and newly recorded strategies. Haseltonia 28: 2–22. https://doi.org/10.2985/026.028.0101
Sprengel, C. (1825) Systema vegetabilium. Editio decima sexta. Volumen II. Sumtibus Librariae Dieterichianae, Gottingae, 939 pp. https://doi.org/10.5962/bhl.title.822
Tulasne, L.-R. [‘Edmond’]. (1856) Floræ madagascariensis. Fragmenta scripsit collectave digessit. Fragmentum primum. Combreteas, Myrobalaneis Junctas, Alangieas et Rhizophoreas includes. Annales des Sciences Naturelles; Botanique sér. 4, 6: 75–138. [https://www.biodiversitylibrary.org/item/129814#page/79/mode/1up]
Tulasne, L.-R. [‘Edmond’]. (1857) Floræ madagascariensis. Fragmenta scripsit collectave digessit. Fragmentum alterum (1). Passifloreas, Homalineas, Chailletieas, Hippocrateaceas, Celastrineas, Ilicineas, Rhamneas, Pittosporeas, Hamamelideas, Crassulaceas et Saxifragaceas exhibens. [Crassulaceæ. I. Bryophyllum. II. Calanchoe (sic, Kalanchoe) on pp. 147–150.] Annales des Sciences Naturelles; Botanique sér. 4, 8: 44–163. [https://www.biodiversitylibrary.org/item/129817#page/151/mode/1up]
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