Abstract
The French botanist Louis-René ‘Edmond’ Tulasne validly published three new names in Kalanchoe (Crassulaceae subfam. Kalanchooideae) in 1857. These are K. eriophylla, K. miniata, and K. floribunda, with the first two of these names being based on material that was collected more than 30 years earlier in Madagascar by Carl Hilsenberg and Wenceslas Bojer, who are jointly credited as having made the first significant plant collections in the interior of the island. The third name, K. floribunda, was published for a species from the Comoros, but is an illegitimate later homonym, for which a lectotype is designated here. The fourth species that Tulasne dealt with in his treatment of Malagasy kalanchoes was included under the name Bryophyllum calycinum, which is now regarded as a synonym of K. pinnata. In the past, Bryophyllum was sometimes accepted as a genus distinct from Kalanchoe, but at present it is treated as K. subg. Bryophyllum. The nomenclature of the three names published in Kalanchoe and the one treated in Bryophyllum by Tulasne is reviewed and discussed in a historical context. The type of the name K. tubiflora, which was collected during a British expedition that lasted from 1822 to 1826 to South and East Africa, is clarified.
References
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<p>Ecklon, C.F. & Zeyher, C. (1837) <em>Enumeratio plantarum Africae australis extratropicae</em>. Publication info: Sumtibus auctorum. Prostat apud Perthes & Besser, Hamburgi, 400 pp. https://doi.org/10.5962/bhl.title.48</p>
<p>Elliott, B. (2004) <em>The Royal Horticultural Society. A history 1804–2004</em>. Phillimore & Co., Ltd, Chicester, West Sussex, England, 432 pp.</p>
<p>Figueiredo, E. & Smith, G.F. (2017) (56) Request for a binding decision on the descriptive statement associated with <em>Kalanchoe delagoensis </em>(Crassulaceae). <em>Taxon </em>66 (3): 771. https://doi.org/10.12705/663.37</p>
<p>Figueiredo, E. & Smith, G.F. (2022) Notes on the itinerary, fleet, and collectors of botanical specimens during the 1822–1826 expedition of William Fitz William Owen to South and East Africa, and beyond. <em>Phytotaxa</em> 568 (2): 170–190. https://doi.org/10.11646/phytotaxa.568.2.3</p>
<p>Forsskål, P. (1775) <em>Flora Ægyptiaco-Arabica. Sive descriptiones plantarum, Quas per Ægyptum inferiorem et Arabiam felicem detexit</em> […]. Ex officina Mölleri, aulæ Typographi. Prostat apud Heineck et Faber, Hauniæ, 220 pp. https://doi.org/10.5962/bhl.title.6625</p>
<p>Hamet, R. (1908) Monographie du genre <em>Kalanchoe</em>. Suite et fin. <em>Bulletin de l’Herbier Boissier</em>, sér. 2, 8: 17–48. [https://www.biodiversitylibrary.org/item/104945#page/31/mode/1up]</p>
<p>Hamet, R. (1912) Observations sur le <em>Kalanchoe tubiflora</em> nom. nov. <em>Beihefte zum Botanischen Centralblatt </em>29: 41–44. [https://www.biodiversitylibrary.org/page/4986270#page/523/mode/1up]</p>
<p>Harvey, W.H. (1862) Order LIII. Crassulaceae, D.C. VIII. <em>Kalanchoe</em>, Adans. <em>In</em>: Harvey, W.H. & Sonder, O.W. (Eds.) <em>Flora capensis</em> [<em>being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal</em>] 2. L. Reeve & Co., Ltd, Kent, pp. 378–380. [https://www.biodiversitylibrary.org/item/15233#page/386/mode/1up]</p>
<p>IPNI (2022+) <em>The International Plant Names Index</em>. Available from: https://www.ipni.org/ (continuously updated; accessed 31 December 2022)</p>
<p>Koorders, S.H. (1919 [Mai] [24 October 1918]) Beitrag zur Kenntniss der Flora von Java No. 18. Beschreibung von <em>Kalanchoë Schumacheri</em> vom Idjen-Plateau und Revision der Crassulaceae von Java. (Mit 2 lithographischen Tafeln.). <em>Bulletin du Jardin Botanique de Buitenzorg</em> sér. 3, 1(3): 169–180, Figures 14 and 15. [https://www.biodiversitylibrary.org/item/27173#page/205/mode/1up] (text) [https://www.biodiversitylibrary.org/item/27173#page/243/mode/1up] (figures)</p>
<p>Kurz, W.S. (1871) On some new or imperfectly known Indian Plants, continuation from Journal, Vol. XXXIX, Part II, pp. 61–91. <em>Journal of the Asiatic Society of Bengal. [Part 2. (Natural History, &c.)]. Calcutta </em>40 (2): 45–78. [https://www.biodiversitylibrary.org/item/110241#page/63/mode/1up]</p>
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<p>Mabberley, D.J. (2020) <em>Botanical revelation. European encounters with Australian plants before Darwin</em>. [The Peter Crossing Collection.] Corrected reprint. NewSouth Publishing, University of New South Wales Press Ltd, Sydney, Australia, 372 pp.</p>
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<p>Shtein, R., Smith, G.F. & Ikeda, J. (2021) Aspects of the taxonomy of the <em>Kalanchoe daigremontiana</em> species complex (Crassulaceae subfam. Kalanchooideae) and associated interspecific hybrids in southern Madagascar, with the description of a new nothospecies, <em>K</em>. ×<em>descoingsii</em> (=<em>K</em>. <em>laetivirens</em> × <em>K</em>. <em>tubiflora</em>). <em>Phytotaxa</em> 524 (4): 235–260. https://doi.org/10.11646/phytotaxa.524.4.2</p>
<p>Smith, G.F. (2020a) <em>Kalanchoe</em> Adans. subg. <em>Fernandesiae</em> Gideon F.Sm. (Crassulaceae subfam. Kalanchooideae), a new subgenus from southern and south-tropical Africa. <em>Haseltonia</em> 27: 2–7. https://doi.org/10.2985/026.027.0101</p>
<p>Smith, G.F. (2020b) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). <em>Bradleya</em> 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15</p>
<p>Smith, G.F. (2021) Reinstatement of <em>Kalanchoe</em> [subg. <em>Bryophyllum</em>] sect. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae) for <em>K. porphyrocalyx</em> and <em>K. uniflora</em>, two Madagascan endemics. <em>Bradleya</em> 39: 177–181. https://doi.org/10.25223/brad.n39.2021.a17</p>
<p>Smith, G.F. (2022a) Notes on <em>Kalanchoe</em> subg. <em>Bryophyllum</em> (Crassulaceae subfam. Kalanchooideae) and the kalanchoes of Sijfert H. Koorders (1919). <em>Phytotaxa</em> 549 (1): 87–96. https://doi.org/10.11646/phytotaxa.549.1.7</p>
<p>Smith, G.F. (2022b) <em>Kalanchoe</em> sect. <em>Raveta</em> (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1916): a long-overlooked name in <em>K</em>. subg. <em>Kalanchoe</em>. <em>Phytotaxa</em> 550 (2): 208–212. https://doi.org/10.11646/phytotaxa.550.2.10</p>
<p>Smith, G.F. (2022c) Date and place of publication and author attribution of the combination <em>Kalanchoe</em> sect. <em>Raveta</em> (Crassulaceae subfam. Kalanchooideae). <em>Phytotaxa</em> 560 (1): 131–134. https://doi.org/10.11646/phytotaxa.560.1.12</p>
<p>Smith, G.F. (2023) A reassessment of combinations previously proposed or published for <em>Kalanchoe</em> subg. <em>Alatae</em> (Crassulaceae subfam. Kalanchooideae) and its valid publication at the rank of subgenus. <em>Phytotaxa. </em>[in press]</p>
<p>Smith, G.F. & Figueiredo, E. (2018a) Nomenclatural notes on <em>Kalanchoe pinnata</em> (Lam.) Pers. (Crassulaceae). <em>Bradleya</em> 36: 220–223. https://doi.org/10.25223/brad.n36.2018.a18</p>
<p>Smith, G.F. & Figueiredo, E. (2018b) The infrageneric classification and nomenclature of <em>Kalanchoe </em>Adans. (Crassulaceae), with special reference to the southern African species. <em>Bradleya </em>36: 162–172. https://doi.org/10.25223/brad.n36.2018.a10</p>
<p>Smith, G.F. & Figueiredo, E. (2021) The taxonomy and nomenclature of <em>Kalanchoe streptantha</em> Baker (Crassulaceae subfam. Kalanchooideae). <em>Bradleya</em> 39: 195–201. https://doi.org/10.25223/brad.n39.2021.a20</p>
<p>Smith, G.F. & Figueiredo, E. (2022a) <em>Kalanchoe beharensis </em>‘Pixel’ (Crassulaceae subfam. Kalanchooideae), eine neue Sorte einer populären Zierpflanze aus Madagaskar. <em>Kakteen und andere Sukkulenten</em> 73 (11): 329–333.</p>
<p>Smith, G.F. & Figueiredo, E. (2022b) Clarifying the typification of the name <em>Kalanchoe orgyalis</em> (Crassulaceae subfam. Kalanchooideae). <em>Phytotaxa</em> 543 (1): 99–102. https://doi.org/10.11646/phytotaxa.543.1.11</p>
<p>Smith, G.F. & Figueiredo, E. (2023) <em>Kalanchoe orgyalis</em>. Crassulaceae subfam. Kalanchooideae. Madagascar. <em>Flowering Plants of Africa</em> 68. [in press]</p>
<p>Smith, G.F., Figueiredo, E. & Van Wyk, A.E. (2019) Kalanchoe <em>(Crassulaceae) in southern Africa. Classification, biology, and cultivation</em>. Academic Press, an imprint of Elsevier, London, San Diego, Cambridge (U.S.A.), and Oxford, 328 pp. https://doi.org/10.1016/C2017-0-00602-X</p>
<p>Smith, G.F., Klein, D.-P. & Shtein, R. (2021c) A revision of the Malagasy <em>Kalanchoe</em> subg. <em>Kitchingia</em> (Crassulaceae subfam. Kalanchooideae): history, taxonomy, and nomenclature. <em>Phytotaxa</em> 507 (1): 67–80. https://doi.org/10.11646/phytotaxa.507.1.3</p>
<p>Smith, G.F. & Monro, A.M. (2022a) The nomenclature and taxonomy of Crassulaceae subfam. Kalanchooideae resolved. <em>Phytotaxa</em> 555 (3): 241–251. https://doi.org/10.11646/phytotaxa.555.3.3</p>
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<p>Smith, G.F. & Shtein, R. (2021) Nomenclature and taxonomic placement of the infraspecific taxa recognised in the Madagascan <em>Kalanchoe miniata</em> (Crassulaceae subfam. Kalanchooideae) by Perrier de la Bâthie and Boiteau and Mannoni in the 20<sup>th</sup> century. <em>Phytotaxa</em> 514 (2): 118–128. https://doi.org/10.11646/phytotaxa.514.2.3</p>
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<p>Smith, G.F., Shtein, R., Klein, D.-P., Parihar, B., Almeida, A., Rodewald, S. & Kadereit, G. (2021a) Special Review. Sexual and asexual reproduction in <em>Kalanchoe</em> (Crassulaceae): a review of known and newly recorded strategies. <em>Haseltonia</em> 28: 2–22. https://doi.org/10.2985/026.028.0101</p>
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