Abstract
A new subspecies of Stachys cretica (Lamiaceae) from Turkey, Stachys cretica subsp. gulendamiana, is described and its relationships with allied subspecies is discussed. The new subspecies is characterized by having decumbent flowering stems. Stachys cretica subsp. gulendamiana is similar to S. cretica subsp. anatolica, but it differs mainly by its marine sandy habitat, indumentum adpressed white lanate—tomentose, flowering stems decumbent, basal leaves undulate, nutlets dull, greyish—brown. IUCN threat category, geographical distribution map and ecology of the new subspecies are also given.
References
Akçiçek, E. (2010) A new subspecies of Stachys cretica (section Eriostomum, Lamiaceae) from Turkey. Turkish Journal of Botany 32: 113–121.
Akçiçek, E. (2020) Taxonomic revision of Stachys sect. Olisia (Lamiaceae: Lamioideae) in Turkey. Phytotaxa 449 (2): 109–148. https://doi.org/10.11646/phytotaxa.449.2.2
Akçiçek, E., Dirmenci, T. & Dündar, E. (2012) Taxonomical notes on Stachys L. Sect. Eriostomum (Hoffmanns. & Link) Dumort. (Lamiaceae) in Turkey. Turkish Journal of Botany 36: 217–234.
Akçiçek, E., F?rat, M. & Güner, Ö. (2016) Stachys hakkariensis (Lamiaceae), a new species from eastern Anatolia (Turkey) belonging to Stachys sect. Olisia. Phytotaxa 257 (2): 167–173. https://doi.org/10.11646/phytotaxa.257.2.6
Ball, P.W. (1972) Stachys L. In: Tutin, T.G., Heywood, V.H., Burgess, N.A., Moore, D.M., Valentine, D.H., Walter, S.M. & Webb, D.A. (Eds.) Flora Europaea, vol. 3. Cambridge University Press, Cambridge, pp. 151–157.
Bendiksby, M., Lisbeth, T., Scheen, A.C., Lindqvist, C. & Ryding, O. (2011) An update phylogeny and classification of Lamiaceae subfamily Lamioideae. Taxon 60: 471–484. https://doi.org/10.1002/tax.602015
Bhattacharjee, R. (1974) Taxonomic studies in Stachys I: New species and infra-specific taxa from Turkey. Notes from the Royal Botanic Garden, Edinburgh 33: 275–292.
Bhattacharjee, R. (1980) Taxonomic studies in Stachys II: A new infrageneric classification of Stachys L. Notes from the Roy. Bot. Gard. Edinburgh 38 (1): 65–96.
Bhattacharjee, R. (1982) Stachys L. In: Davis, P.H. (Ed.) Flora of Turkey and the East Aegean Islands, vol. 7. Edinburgh University Press, Edinburgh, pp. 199–262.
Boissier, E. (1879) Flora Orientalis, vol. 4. H. Georg., Basel, Genève, pp. 714–752.
Brown, R. (1812) Prodromus florae Novae-Hollandiae et insulae van-Diemen. Vol. 4. Hortus Kew. pp. 120.
Crantz, H.J.N. (1763) Henrici Joa. Nepom. Crantz. Stirpium austriarum. Vol. 2. Krauss. pp. 131.
Davidson-Arnott, R. (2010) An Introduction to Coastal Processes and Geomorphology. Cambridge University Press, UK. https://doi.org/10.1017/CBO9780511841507
Davis, P.H., Mill, R.R. & Tan, K. (1988) Flora of Turkey and East Aegean Islands, vol. 10. Edinburgh University Press, Edinburgh. pp. 204–206.
Dimopoulos, P., Raus, Th., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A. & Tzanoudakis, D. (2013) Vascular plants of Greece: An annotated checklist. Berlin: Botanic Garden and Botanical Museum Berlin-Dahlem; Athens: Hellenic Botanical Society. [Englera 31].
Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, Th., Iatrou, G., Kokkini, S., Strid, A. & Tzanoudakis, D. (2016) Vascular plants of Greece: An annotated checklist. Suppl. Willdenowia 46: 301–347. https://doi.org/10.3372/wi.46.46303
Dirmenci, T., Y?ld?z, B., Akçiçek, E., Martin, E. & Dündar, E. (2011) Stachys vuralii (Lamiaceae), a new species from north Anatolia, Turkey. Annales Botanici Fennici 48: 401–408. https://doi.org/10.5735/085.048.0503
Duman, H. (2000) Stachys L. In: Güner, A., Özhatay, N., Ekim, T. & Ba?er, K.H.C. (Eds.) Flora of Turkey and the East Aegean Islands, vol. 11. Edinburgh University Press, Edinburgh. pp. 204–206.
Dumortier, B.C. (1827) Florula Belgica. Staminacia, Tornaci Nerviorum: Casterman, 172 pp.
Falciani, L. (1997) Systematic Revision of Stachys Sect. Eriostomum (Hoffmans. & Link) Dumort. in Italy. Lagascalia 19: 187–238.
http://apps.kew.org/wcsp/ (accessed 11 November 2016).
Feinbrun-Dothan, N. (1978) Flora Palaestina, vol. 3. The Israel Academy of Sciences and Humanities, Jerusalem, pp. 128–133.
Gören, A.C., Piozzi, F., Akcicek, E. K?l?ç, T., Çar?kçI, S., Mozio?lu, E. & Setzer, W.N. (2011) Essential oil composition of twenty-two Stachys species (mountain tea) and their biological activities. Phytochemistry Letters 4 (4): 448–453. https://doi.org/10.1016/j.phytol.2011.04.013
Govaerts, R., Paton, A., Harve, Y., Navarro, T. & Pena, M.R.G. (2022) [continuously updated] World Checklist of Lamiaceae. Facilitated by the Royal Botanic Gardens, Kew (accessed 25 January 2022).
Güner, Ö., Akçiçek, E. & Dirmenci, T. (2021) A new Stachys species from Turkey: Stachys siirtensis (Lamiaceae). Phytotaxa 516 (3): 252–262. https://doi.org/10.11646/phytotaxa.516.3.4
Güner, Ö. & Ferrer-Galego, P.P. (2021) Nomenclatural and taxonomic notes on some Stachys taxa (Lamiaceae). Turkish Journal of Botany 45: 69–82. https://doi.org/10.3906/bot-2010-12
Hacio?ullari, I., Gücel, S., Fuller, W.J. & Özden, Ö. (2019) Abundance and spatial distribution analysis of Salvia veneris: a critically endangered plant species endemic to Cyprus. Biodiversity 20 (2–3): 98–105. https://doi.org/10.1080/14888386.2019.1662327
Harley, R.M., Atkins, S., Budantsev, A.L., Cantino, P.D., Conn, B.J., Grayer, R., Harley, MM., de Kok, R., Krestovskaya, T., Morales, R., Paton, A.J., Ryding, O. & Upson, T. (2004) Labiatae. In: Kubitzki, K. & Kadereit, J.W. (Eds.) The families and genera of vascular plants 7. Springer, Berlin, Heidelberg, pp. 167–275.
Hoffmannsegg, J.C. & Link, H.F. (1809) Flora Portugaise, vol. 1. Berlino 458 pp. https://doi.org/10.1007/978-3-642-18617-2_11
Hudson, W. (1762) Flora Anglica, Exhibens Plantas per Regnum Angliae Sponte Crescentes, Distributas Secundum Systema Sexuale. J. Nourse & C. Moran, London. https://doi.org/10.5962/bhl.title.124863
IUCN. (2021) The IUCN Red List of Threatened Species. Version 2021-3. Available from: https://www.iucnredlist.org (accessed 16 February 2022)
Krestovskaya, T.V. (2011) The critical review of species of the genus Stachys L. section Eriostomum Hoffmanns. et Link (Labiatae). Novosti Sistematiki Vysshikh Rastenii 42: 198–221.
Linnaeus, C. (1753) Species Plantarum, vols. 1–2. Holmiæ, Impensis. Laurentii Salvii, [Stockholm], 1200 pp.
Linnaeus, C. (1756) “Centuria II Plantarum.” Centuria II Plantarum.
Martinov (1820) Tekhno-Bot. Slovar: 355 (nom. cons.)
Miller, P. (1768) The gardeners dictionary. ed. VIII. London.
Orsenigo, S., Bacchetta, G., Calevo, J., Castello, M., Cogoni, D., Gennai, M., Licht, W., Montagnani, C., Perrino, E.V., Pinna, S.M., Silletti, G.N., Vela, E., Viciani, D., Vidali, M., Wagensommer, R.P., Zappa, E. & Fenu, G. (2016) Global and regional IUCN Red List Assessments: 1. Italian Botanist 1: 61–85. https://doi.org/10.3897/italianbotanist.1.8647
Özdöl, T., Güner, Ö., Sefal?, A., Akçiçek, E., Dirmenci, T. &Y?ld?r?m, H. (2022) Three new records for the flora of Turkey: Campanula lyrata subsp. icarica (Campanulaceae), Erysimum aureum (Brassicaceae) and Stachys benthamiana (Lamiaceae). Phytotaxa 531 (2): 147–150. https://doi.org/10.11646/phytotaxa.531.2.8
Öztürk, M., Duru, M.E., Aydo?mus-Öztürk, F., Harmandar, M., Mahlicli, M., Kolak, U. & Ulubelen, A. (2009) GC–MS analysis and antimicrobial activity of essential oil of Stachys cretica subsp. smyrnaea. Natural Product Communications 4 (1): 109–114. https://doi.org/10.1177/1934578X0900400124
Perrino, E.V., Valerio, F., Gannouchi, A., Trani, A. & Mezzapesa, G. (2021) Ecological and Plant Community Implication on Essential Oils Composition in Useful Wild Officinal Species: A Pilot Case Study in Apulia (Italy). Plants 10 (3): 574. https://doi.org/10.3390/plants10091952
Perrino, E.V. & Wagensommer, R.P. (2022) Crop Wild Relatives (CWRs) Threatened and Endemic to Italy: Urgent Actions for Protection and Use. Biology 11 (2): 193. https://doi.org/10.3390/biology11020193
Reed, J.D., Davidson-Arnott, R. & Perillo, G.M.E. (2009) Estuaries, coastal marshes, tidal flats and coastal dunes.” Geomorphology and global environmental change. Cambridge University Press, UK. pp. 130–157. https://doi.org/10.1017/CBO9780511627057.006
Rechinger, K.H. (1937) Revision des Formenkreises der Stachys cretica. In: Keissler, K. (ed.) Ann. Naturhist. Mus. in Wien. Vol. 48. Vienna: Natural History Museum. pp. 167–178.
Salmaki, Y., Zarre, S., Govaerts, R. & Bräuchler, C. (2012) A taxonomic revision of the genus Stachys (Lamiaceae: Lamioideae) in Iran. Botanical Journal of the Linnean Society 170: 573–617. https://doi.org/10.1111/j.1095-8339.2012.01317.x
Salmaki, Y., Zarre, S., Ryding, O., Lindqvist, C., Bräuchler, C., Heubl, G., Barber, J. & Bendiksby, M. (2013) Molecular phylogeny of the tribe Stachydeae (Lamiaceae subfamily Lamioideae). Molecular Phylogenetics and Evolution 69: 535–551. https://doi.org/10.1016/j.ympev.2013.07.024
Sat?l, F., Tümen, G. & Selvi, S. (2019) Ayval?k (Bal?kesir/Turkey) Dune Plant Diversity, Threatening Factors and Solution Proposals. Journal of the Institute of Science and Technology 9 (3): 1289–129. https://doi.org/10.21597/jist.550131
Scheen, A.C., Bendiksby, M., Ryding, O., Mathiesen, C., Albert, V.A. & Lindqvist, C. (2010) Molecular phylogenetics, character evolution, and suprageneric classification of Lamioideae (Lamiaceae). Annals of the Missouri Botanical Garden 97: 191–219. https://doi.org/10.3417/2007174
Schultes, J.A. & Schultes, J.H. (1827) “Mantissa [...] Volumen tertium.”
Smith, J.E. (1798) Remarks on some Foreign Species of Orobanche. Transactions of the Linnean Society 4: 169. https://doi.org/10.1111/j.1096-3642.1798.tb00524.x
Soares, A.D.S., Negrão, R., Harley, R.M., Pastore, J.F.B. & Jardim, J.G. (2021) Rediscovery of Oocephalus foliosus (Hyptidinae-Lamiaceae): notes on taxonomy and conservation of a species endemic to the campo rupestre of Central Brazil. Phytotaxa 525 (2): 156–162. https://doi.org/10.11646/phytotaxa.525.2.6
Strid, A. & Tan, K. (Eds.) (1991) Mountain Flora of Greece, vol. 2. Edinburgh University Press, Edinburgh, pp. 97–107.
Thiers, B. (2022) Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 30 January 2022)
Turo?lu, H. (2009) 3621 Say?l? K?y? Kanunu ve Onun Uygulama Problemleri. Türk Co?rafya Dergisi 53: 31–40. [in Turkish]
Valerio, F., Mezzapesa, G.N., Ghannouchi, A., Mondelli, D., Logrieco, A.F. & Perrino, E.V. (2021) Characterization and antimicrobial properties of essential oils from four wild taxa of Lamiaceae family growing in Apulia. Agronomy 11 (7): 1431. https://doi.org/10.3390/agronomy11071431
Wagensommer, R.P., Bartolucci, F., Fiorentino, M., Licht, W., Peccenini, S., Perrino, E.V. & Venanzoni, R. (2017) First record for the flora of Italy and lectotypification of the name Linum elegans (Linaceae). Phytotaxa 296 (2): 161–170. https://doi.org/10.11646/phytotaxa.296.2.5
Wagensommer, R.P., Perrino, E.V. & Silletti, G.N. (2014) Carex phyllostachys C.A. Mey. (Cyperaceae) new for Italy and phytogeographical considerations. Phyton (Horn, Austria) 54 (2): 215–222.
Akçiçek, E. (2020) Taxonomic revision of Stachys sect. Olisia (Lamiaceae: Lamioideae) in Turkey. Phytotaxa 449 (2): 109–148. https://doi.org/10.11646/phytotaxa.449.2.2
Akçiçek, E., Dirmenci, T. & Dündar, E. (2012) Taxonomical notes on Stachys L. Sect. Eriostomum (Hoffmanns. & Link) Dumort. (Lamiaceae) in Turkey. Turkish Journal of Botany 36: 217–234.
Akçiçek, E., F?rat, M. & Güner, Ö. (2016) Stachys hakkariensis (Lamiaceae), a new species from eastern Anatolia (Turkey) belonging to Stachys sect. Olisia. Phytotaxa 257 (2): 167–173. https://doi.org/10.11646/phytotaxa.257.2.6
Ball, P.W. (1972) Stachys L. In: Tutin, T.G., Heywood, V.H., Burgess, N.A., Moore, D.M., Valentine, D.H., Walter, S.M. & Webb, D.A. (Eds.) Flora Europaea, vol. 3. Cambridge University Press, Cambridge, pp. 151–157.
Bendiksby, M., Lisbeth, T., Scheen, A.C., Lindqvist, C. & Ryding, O. (2011) An update phylogeny and classification of Lamiaceae subfamily Lamioideae. Taxon 60: 471–484. https://doi.org/10.1002/tax.602015
Bhattacharjee, R. (1974) Taxonomic studies in Stachys I: New species and infra-specific taxa from Turkey. Notes from the Royal Botanic Garden, Edinburgh 33: 275–292.
Bhattacharjee, R. (1980) Taxonomic studies in Stachys II: A new infrageneric classification of Stachys L. Notes from the Roy. Bot. Gard. Edinburgh 38 (1): 65–96.
Bhattacharjee, R. (1982) Stachys L. In: Davis, P.H. (Ed.) Flora of Turkey and the East Aegean Islands, vol. 7. Edinburgh University Press, Edinburgh, pp. 199–262.
Boissier, E. (1879) Flora Orientalis, vol. 4. H. Georg., Basel, Genève, pp. 714–752.
Brown, R. (1812) Prodromus florae Novae-Hollandiae et insulae van-Diemen. Vol. 4. Hortus Kew. pp. 120.
Crantz, H.J.N. (1763) Henrici Joa. Nepom. Crantz. Stirpium austriarum. Vol. 2. Krauss. pp. 131.
Davidson-Arnott, R. (2010) An Introduction to Coastal Processes and Geomorphology. Cambridge University Press, UK. https://doi.org/10.1017/CBO9780511841507
Davis, P.H., Mill, R.R. & Tan, K. (1988) Flora of Turkey and East Aegean Islands, vol. 10. Edinburgh University Press, Edinburgh. pp. 204–206.
Dimopoulos, P., Raus, Th., Bergmeier, E., Constantinidis, T., Iatrou, G., Kokkini, S., Strid, A. & Tzanoudakis, D. (2013) Vascular plants of Greece: An annotated checklist. Berlin: Botanic Garden and Botanical Museum Berlin-Dahlem; Athens: Hellenic Botanical Society. [Englera 31].
Dimopoulos, P., Raus, T., Bergmeier, E., Constantinidis, Th., Iatrou, G., Kokkini, S., Strid, A. & Tzanoudakis, D. (2016) Vascular plants of Greece: An annotated checklist. Suppl. Willdenowia 46: 301–347. https://doi.org/10.3372/wi.46.46303
Dirmenci, T., Y?ld?z, B., Akçiçek, E., Martin, E. & Dündar, E. (2011) Stachys vuralii (Lamiaceae), a new species from north Anatolia, Turkey. Annales Botanici Fennici 48: 401–408. https://doi.org/10.5735/085.048.0503
Duman, H. (2000) Stachys L. In: Güner, A., Özhatay, N., Ekim, T. & Ba?er, K.H.C. (Eds.) Flora of Turkey and the East Aegean Islands, vol. 11. Edinburgh University Press, Edinburgh. pp. 204–206.
Dumortier, B.C. (1827) Florula Belgica. Staminacia, Tornaci Nerviorum: Casterman, 172 pp.
Falciani, L. (1997) Systematic Revision of Stachys Sect. Eriostomum (Hoffmans. & Link) Dumort. in Italy. Lagascalia 19: 187–238.
http://apps.kew.org/wcsp/ (accessed 11 November 2016).
Feinbrun-Dothan, N. (1978) Flora Palaestina, vol. 3. The Israel Academy of Sciences and Humanities, Jerusalem, pp. 128–133.
Gören, A.C., Piozzi, F., Akcicek, E. K?l?ç, T., Çar?kçI, S., Mozio?lu, E. & Setzer, W.N. (2011) Essential oil composition of twenty-two Stachys species (mountain tea) and their biological activities. Phytochemistry Letters 4 (4): 448–453. https://doi.org/10.1016/j.phytol.2011.04.013
Govaerts, R., Paton, A., Harve, Y., Navarro, T. & Pena, M.R.G. (2022) [continuously updated] World Checklist of Lamiaceae. Facilitated by the Royal Botanic Gardens, Kew (accessed 25 January 2022).
Güner, Ö., Akçiçek, E. & Dirmenci, T. (2021) A new Stachys species from Turkey: Stachys siirtensis (Lamiaceae). Phytotaxa 516 (3): 252–262. https://doi.org/10.11646/phytotaxa.516.3.4
Güner, Ö. & Ferrer-Galego, P.P. (2021) Nomenclatural and taxonomic notes on some Stachys taxa (Lamiaceae). Turkish Journal of Botany 45: 69–82. https://doi.org/10.3906/bot-2010-12
Hacio?ullari, I., Gücel, S., Fuller, W.J. & Özden, Ö. (2019) Abundance and spatial distribution analysis of Salvia veneris: a critically endangered plant species endemic to Cyprus. Biodiversity 20 (2–3): 98–105. https://doi.org/10.1080/14888386.2019.1662327
Harley, R.M., Atkins, S., Budantsev, A.L., Cantino, P.D., Conn, B.J., Grayer, R., Harley, MM., de Kok, R., Krestovskaya, T., Morales, R., Paton, A.J., Ryding, O. & Upson, T. (2004) Labiatae. In: Kubitzki, K. & Kadereit, J.W. (Eds.) The families and genera of vascular plants 7. Springer, Berlin, Heidelberg, pp. 167–275.
Hoffmannsegg, J.C. & Link, H.F. (1809) Flora Portugaise, vol. 1. Berlino 458 pp. https://doi.org/10.1007/978-3-642-18617-2_11
Hudson, W. (1762) Flora Anglica, Exhibens Plantas per Regnum Angliae Sponte Crescentes, Distributas Secundum Systema Sexuale. J. Nourse & C. Moran, London. https://doi.org/10.5962/bhl.title.124863
IUCN. (2021) The IUCN Red List of Threatened Species. Version 2021-3. Available from: https://www.iucnredlist.org (accessed 16 February 2022)
Krestovskaya, T.V. (2011) The critical review of species of the genus Stachys L. section Eriostomum Hoffmanns. et Link (Labiatae). Novosti Sistematiki Vysshikh Rastenii 42: 198–221.
Linnaeus, C. (1753) Species Plantarum, vols. 1–2. Holmiæ, Impensis. Laurentii Salvii, [Stockholm], 1200 pp.
Linnaeus, C. (1756) “Centuria II Plantarum.” Centuria II Plantarum.
Martinov (1820) Tekhno-Bot. Slovar: 355 (nom. cons.)
Miller, P. (1768) The gardeners dictionary. ed. VIII. London.
Orsenigo, S., Bacchetta, G., Calevo, J., Castello, M., Cogoni, D., Gennai, M., Licht, W., Montagnani, C., Perrino, E.V., Pinna, S.M., Silletti, G.N., Vela, E., Viciani, D., Vidali, M., Wagensommer, R.P., Zappa, E. & Fenu, G. (2016) Global and regional IUCN Red List Assessments: 1. Italian Botanist 1: 61–85. https://doi.org/10.3897/italianbotanist.1.8647
Özdöl, T., Güner, Ö., Sefal?, A., Akçiçek, E., Dirmenci, T. &Y?ld?r?m, H. (2022) Three new records for the flora of Turkey: Campanula lyrata subsp. icarica (Campanulaceae), Erysimum aureum (Brassicaceae) and Stachys benthamiana (Lamiaceae). Phytotaxa 531 (2): 147–150. https://doi.org/10.11646/phytotaxa.531.2.8
Öztürk, M., Duru, M.E., Aydo?mus-Öztürk, F., Harmandar, M., Mahlicli, M., Kolak, U. & Ulubelen, A. (2009) GC–MS analysis and antimicrobial activity of essential oil of Stachys cretica subsp. smyrnaea. Natural Product Communications 4 (1): 109–114. https://doi.org/10.1177/1934578X0900400124
Perrino, E.V., Valerio, F., Gannouchi, A., Trani, A. & Mezzapesa, G. (2021) Ecological and Plant Community Implication on Essential Oils Composition in Useful Wild Officinal Species: A Pilot Case Study in Apulia (Italy). Plants 10 (3): 574. https://doi.org/10.3390/plants10091952
Perrino, E.V. & Wagensommer, R.P. (2022) Crop Wild Relatives (CWRs) Threatened and Endemic to Italy: Urgent Actions for Protection and Use. Biology 11 (2): 193. https://doi.org/10.3390/biology11020193
Reed, J.D., Davidson-Arnott, R. & Perillo, G.M.E. (2009) Estuaries, coastal marshes, tidal flats and coastal dunes.” Geomorphology and global environmental change. Cambridge University Press, UK. pp. 130–157. https://doi.org/10.1017/CBO9780511627057.006
Rechinger, K.H. (1937) Revision des Formenkreises der Stachys cretica. In: Keissler, K. (ed.) Ann. Naturhist. Mus. in Wien. Vol. 48. Vienna: Natural History Museum. pp. 167–178.
Salmaki, Y., Zarre, S., Govaerts, R. & Bräuchler, C. (2012) A taxonomic revision of the genus Stachys (Lamiaceae: Lamioideae) in Iran. Botanical Journal of the Linnean Society 170: 573–617. https://doi.org/10.1111/j.1095-8339.2012.01317.x
Salmaki, Y., Zarre, S., Ryding, O., Lindqvist, C., Bräuchler, C., Heubl, G., Barber, J. & Bendiksby, M. (2013) Molecular phylogeny of the tribe Stachydeae (Lamiaceae subfamily Lamioideae). Molecular Phylogenetics and Evolution 69: 535–551. https://doi.org/10.1016/j.ympev.2013.07.024
Sat?l, F., Tümen, G. & Selvi, S. (2019) Ayval?k (Bal?kesir/Turkey) Dune Plant Diversity, Threatening Factors and Solution Proposals. Journal of the Institute of Science and Technology 9 (3): 1289–129. https://doi.org/10.21597/jist.550131
Scheen, A.C., Bendiksby, M., Ryding, O., Mathiesen, C., Albert, V.A. & Lindqvist, C. (2010) Molecular phylogenetics, character evolution, and suprageneric classification of Lamioideae (Lamiaceae). Annals of the Missouri Botanical Garden 97: 191–219. https://doi.org/10.3417/2007174
Schultes, J.A. & Schultes, J.H. (1827) “Mantissa [...] Volumen tertium.”
Smith, J.E. (1798) Remarks on some Foreign Species of Orobanche. Transactions of the Linnean Society 4: 169. https://doi.org/10.1111/j.1096-3642.1798.tb00524.x
Soares, A.D.S., Negrão, R., Harley, R.M., Pastore, J.F.B. & Jardim, J.G. (2021) Rediscovery of Oocephalus foliosus (Hyptidinae-Lamiaceae): notes on taxonomy and conservation of a species endemic to the campo rupestre of Central Brazil. Phytotaxa 525 (2): 156–162. https://doi.org/10.11646/phytotaxa.525.2.6
Strid, A. & Tan, K. (Eds.) (1991) Mountain Flora of Greece, vol. 2. Edinburgh University Press, Edinburgh, pp. 97–107.
Thiers, B. (2022) Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 30 January 2022)
Turo?lu, H. (2009) 3621 Say?l? K?y? Kanunu ve Onun Uygulama Problemleri. Türk Co?rafya Dergisi 53: 31–40. [in Turkish]
Valerio, F., Mezzapesa, G.N., Ghannouchi, A., Mondelli, D., Logrieco, A.F. & Perrino, E.V. (2021) Characterization and antimicrobial properties of essential oils from four wild taxa of Lamiaceae family growing in Apulia. Agronomy 11 (7): 1431. https://doi.org/10.3390/agronomy11071431
Wagensommer, R.P., Bartolucci, F., Fiorentino, M., Licht, W., Peccenini, S., Perrino, E.V. & Venanzoni, R. (2017) First record for the flora of Italy and lectotypification of the name Linum elegans (Linaceae). Phytotaxa 296 (2): 161–170. https://doi.org/10.11646/phytotaxa.296.2.5
Wagensommer, R.P., Perrino, E.V. & Silletti, G.N. (2014) Carex phyllostachys C.A. Mey. (Cyperaceae) new for Italy and phytogeographical considerations. Phyton (Horn, Austria) 54 (2): 215–222.