Abstract
Tibet is a very important biodiversity hot spot in China and the diatom flora of this region has drawn a lot of attention. Diatom samples were collected from several places across Tibet. During research on the biodiversity of diatoms from Tibet, one Achnanthidium species showed unique features compared with other known taxa. It has linear-lanceolate valves with rounded apices, tear-drop shaped proximal raphe ends and ?-shaped distal raphe ends. The most distinctive character of this species, however, is the shape of areolae; since this species has nearly round external areolar openings on the valve face but more typical slit-like areolae present on the mantle. Here we describe it as new to science as Achnanthidium tibeticum Y. Liu & Kociolek. We compare this new species with other, similar taxa within the genus.
References
Bukhtiyarova, L. & Round, F.E. (1996) Revision of the genus Achnanthes senu lato. Psammothidium, a new genus based on A. marginulatum. Diatom Research 11: 1–30. https://doi.org/10.1080/0269249X.1996.9705361
Chudaev, D.A. & Georgiev, A.A. (2016) A New taxa of Navicula sensu stricto (Bacillariophyta, Naviculaceae) from high-altitude lake in Tibet, China. Phytotaxa 243: 180–184. https://doi.org/10.11646/phytotaxa.243.2.9
Czarnecki, D.B. & Edlund, M.B. (1995) New combinations for some taxa of Achnanthes. Diatom Research 10: 207–209. https://doi.org/10.1080/0269249X.1995.9705338
Desrosiers, C., Witkowski, A., Riaux-Gobin, C., Zglobicka, I., Kurzydlowski, K.J., Eulin, A., Leflaive, J. & Ten-Hage, L. (2014) Madinithidium gen. nov. (Bacillariophyceae), a new monoraphid diatom genus from the tropical marine coastal zone. Phycologia 53: 583–592. https://doi.org/10.2216/14-21R2
Hlúbiková, D., Ector, L. & Hoffmann, L. (2011) Examination of the type material of some diatom species related to Achnanthidium minutissimum (Kütz.) Czarn. (Bacillariophyceae). Algological Studies 136/137: 19–43. https://doi.org/10.1127/1864-1318/2011/0136-0019
Hustedt, F. (1910) Beitrag zur Algenflora von Afrika. Bacillariales aus Dahomé. Archiv für Hydrobiologies and Planktonkunde 5: 365–382.
Hustedt, F. (1922) Bacillariales aus Innerasien. Gesammelt von Dr. Sven Hedin. In: Hedin, S. (Ed.) Southern Tibet, discoveries in former times compared with my own researches in 1906-1908. Lithographic Institute of the General Staff of the Swedish Army, Stockholm, 6: 107–152. https://doi.org/10.5962/bhl.title.64226
Jao, C., Zhu, H. & Lee, Y. (1974) The fresh-water algae from Mount Qomolangma district (in Tibet). Report of the Scientific Survey of Mount Qomolangma district 1966–1968: 92–126.
Kobayasi, H. (1997) Comparative studies among four linear-lanceolate Achnanthidium species (Bacillariophyceae) with curved terminal raphe endings. Nova Hedwigia 65 (1/4): 147–163. https://doi.org/10.1127/nova.hedwigia/65/1997/147
Kobayasi, H., Idei, M., Mayama, S., Nagumo, T. & Osada, K. (2006) Atlas of Japanese Diatoms based on electron microscopy. Volume 1. Uchida Rokakuko Publishing Co., Tokyo. 59 + 533 pp.
Kobayasi, H., Nagumo, T. & Mayama, S. (1986) Observations on the two rheophilic species of the genus Achnanthes (Bacillariophyceae), A. convergens H. Kob. and A. japonica H. Kob. Diatom 2: 83–93.
Kociolek, J.P., You, Q.M., Yu, P., Li, Y.L., Wang, Y.L., Lowe, R. & Wang, Q.X. (2021) Description of Gomphothidium gen. nov., with light and scanning electron microscopy: a new freshwater monoraphid diatom genus from Asia. Fottea 21: 1–7. https://doi.org/10.5507/fot.2020.011
Krammer, K. & Lange-Bertalot, H. (2004) Bacillariophyceae 4. Teil: Achnanthaceae, Kritische Erganzungen zu Navicula (Lineolatae), Gomphonema Gesamtliteraturverzeichnis [second revised edition] In: Suesswasserflora von Mitteleuropa. Spektrum Akademischer Verlad Heidelberg, 2, 4, 468 pp.
Kulikovskiy, M., Glushchenko, A., Genkal, S.I., Kuznetsova, I.V. & Kociolek, J.P. (2020) Platebaikalia - a new monoraphid diatom genus from ancient Lake Baikal with comments on the genus Platessa. Fottea, Olomouc 20: 58–67. https://doi.org/10.5507/fot.2019.014
Kulikovskiy, M., Lange-Bertalot , H. & Witkowski, A. (2013) Gliwiczia gen. nov. a new monoraphid diatom genus from Lake Baikal with a description of four species new for science. Phytotaxa 109: 1–16. https://doi.org/10.11646/phytotaxa.109.1.1
Kulikovskiy, M., Lange-Bertalot, H. & Kuznetsova, I.V. (2015) Lake Baikal: hotspot of endemic diatoms II. Iconographia Diatomologica 26: 1–656.
Kulikovskiy, M., Maltsev,Y., Glushchenko, A., Kuznetsova, I., Kapustin, D., Gusev, E., Lange-Bertalot, H., Genkal, S. & Kociolek, J.P. (2020) Gogorevia, a new monoraphid diatom genus for Achnanthes exigua and allied taxa (Achnanthidiaceae) described on the basis of an integrated molecular and morphological approach. Journal of Phycology 56: 1601–1613. https://doi.org/10.1111/jpy.13064
Kulikovskiy, M.S., Lange-Bertalot, H., Metzeltin, D. & Witkowski, A. (2012) Lake Baikal: hotspot of endemic diatoms I. Iconographia Diatomologica 23: 7–607.
Kützing, F.T. (1844) Die Kieselschaligen. Bacillarien oder Diatomeen. Nordhausen., 152 pp. https://doi.org/10.5962/bhl.title.64360
Li, S. & Zhang, Q. (2008) Geochemistry of the upper Han River basin, China, 1: Spatial distribution of major ion compositions and their controlling factors. Applied Geochemistry 23: 3535–3544. https://doi.org/10.1016/j.apgeochem.2008.08.012
Lange-Bertalot, H. (1999) Neue Kombinationen von Taxa aus Achnanthes Bory (sensu lato). Iconographia Diatomologica 6: 270–283
Li, Y., Williams, D.M. , Metzeltin, D. , Kociolek, J.P. & Gong, Z. (2010) Tibetiella pulchra gen. nov. et sp. nov. a new freshwater epilithic diatom (Bacillariophyta) from river nujiang in Tibet, China. Journal of Phycology 46: 325–330. https://doi.org/10.1111/j.1529-8817.2009.00776.x
Li, Y., You, Q.M., Kociolek, J.P., Wang, L.Q. & Zhang, W. (2019) Gomphosinica selincuoensis sp. nov. a new diatom (Bacillariophyceae) from North Tibet, China. Phytotaxa 423: 195–205. https://doi.org/10.11646/phytotaxa.423.3.8
Liu, Q., Li, J.J., Nan, F.R., Feng, J., Lv, J.P., Xie, S.L. & Kociolek, J.P. (2019) New and interesting diatoms from Tibet: I. Description of a new species of Clipeoparvus Woodbridge et al. Diatom Research 34: 33–38. https://doi.org/10.1080/0269249X.2019.1578697
Liu, Q., Xiang, Y.D., Yu, P., Xie, S.L. & Kociolek, J.P. (2020) New and interesting diatoms from Tibet. II. Description of two new species of monoraphid diatoms. Diatom Research 35: 353–361. https://doi.org/10.1080/0269249X.2020.1810781
Liu, Q., Yang, L., Nan, F.R., Feng, J., Lv, J.P., Kociolek, J.P. & Xie, S.L. (2018) A new diatom species of Aneumastus D. G. Mann & Stickle ((Bacillariophyta, Bacillariophyceae, Mastogloiales, Mastogloiaceae) from Tibet, China. Phytotaxa 373: 231–235. https://doi.org/10.11646/phytotaxa.373.3.6
Liu, Y., Kociolek, J.P., Liu, Q., Tan, X. & Fan, Y.W. (2020) A new aerophilic Neidium Pfister (Neidiaceae, Bacillariophyta) species from Guangxi Zhuang Autonomous Region, China. Phytotaxa 432 (2): 171–180. https://doi.org/10.11646/phytotaxa.432.2.6
Liu, Y., Tan, X., Kociolek, J.P., Kulikovskiy, M., Lu, X.X. & Fan, Y.W. (2021) One new species of Achnanthidium Kützing (Bacillariophyta, Achnanthidiaceae) from the upper Han River, China. Phytotaxa 516 (2): 187–194. https://doi.org/10.11646/phytotaxa.516.2.6
Monnier, O., Lange-Bertalot, H., Rimet, F., Hoffmann, L. & Ector, L. (2004) Achnanthidium atomoides sp. nov., A new diatom from the Grand-Duchy of Luxembourg. Vie Milieu 54 (2–3): 127–136.
Monnier, O., Lange-Bertalot, H., Hoffmann, L. & Ector, L. (2007) The genera Achnanthidium Ku?tzing and Psammothidium Bukhtiyarova & Round in the family Achnanthidiaceae (Bacillariophyceae): a reappraisal of the differential criteria. Cryptogamie, Algologie 28: 141–158.
Moser, G., Lange-Bertalot, H. & Metzeltin, D. (1998) Insel der Endemiten Geobotanisches Phänomen Neukaledonien (Island of endemics New Caledonia - a geobotanical phenomenon). Bibliotheca Diatomologica. 38: 1–464.
Novais, M.H., Jüttner, I., Hoffmann, L. & Ector, L. (2015) Morphological variability within the Achnanthidium minutissimum species complex (Bacillariophyta): comparison between the type material of Achnanthes minutissima and related taxa, and new freshwater Achnanthidium species from Portugal. Phytotaxa 224: 101–139. https://doi.org/10.11646/phytotaxa.224.2.1
Pérès, F., Barthès, A., Ponton, E., Coste, M., Ten-Hage, L. & Le-Cohu, R. (2012) Achnanthidium delmontii sp. nov., a new species from French rivers. Fottea 12 (2): 189–198. https://doi.org/10.5507/fot.2012.014
Potapova, M. (2006) Achnanthidiumn zhakovschikovii sp. nov. (Bacillariophyta) and related species from rivers of Northwestern Russia. Nova Hedwigia 82 (3–4): 399–408. https://doi.org/10.1127/0029-5035/2006/0082-0399
Round, F.E. (1998) Validation of some previously published “achnanthoid” genera. Diatom Research 13: 181. https://doi.org/10.1080/0269249X.1998.9705442
Round, F.E. & Basson, P.W. (1997) A new monoraphid diatom genus (Pogoneis) from Bahrain and the transfer of previously described species A. hungarica and A. taeniata to new genera. Diatom Research 12: 71–81. https://doi.org/10.1080/0269249X.1997.9705403
Round, F.E. & Bukhtiyarova, L. (1996) Four new genera based on Achnanthes (Achnanthidium) together with a re-definition of Achnanthidium. Diatom Research 11 (2): 345–361. https://doi.org/10.1080/0269249X.1996.9705389
Round, F.E., Crawford, R.M. & Mann, D.G. (1990) The Diatoms. Biology and Morphology of the Genera, Cambridge University Press, Cambridge. 747 pp.
Silva, P.C. (1962) Classification of Algae. Appendix A. In: Ralph, A.L. (Ed.) Physiology and Biochemstry of Algae. Academic Press. N. R., London. pp. 827–837.
Simonsen, R. (1987) Atlas and Catalogue of the Diatom Types of Friedrich Hustedt. 3 Volumes. J. Cramer, Berlin.
Smith, W. (1855) Notes of an excursion to the south of France and the Auvergne in search of Diatomaceae. Annals and Magazine of Natural History 15: 1–9. https://doi.org/10.1080/037454809495381
Van de Vijver, B., Ector, L., Beltrami, M.E., De Haan, M., Falasco, E., Hlúbiková, D., Jarlman, A., Kelly, M., Novais, M.H. & Wojtal, A.Z. (2011) A critical analysis of the type material of Achnanthidium lineare W. Smith (Bacillariophyceae). Algological Studies 136/137: 167–191. https://doi.org/10.1127/1864-1318/2011/0136-0167
Wojtal, A.Z. (2013) Species composition and distribution of diatom assemblages in spring waters from various geological formations in Southern Poland. Bibliotheca Diatomologica 59: 1–436.
You, Q.M., Cao, Y., Yu, P., Kociolek, J.P., Zhang, L.X., Wu, B., Lowe, R. & Wang, Q.X. (2019) Three new subaerial Achnanthidium (Bacillariophyta) species from a karst landform in the Guizhou Province, China. Fottea 19: 138–150. https://doi.org/10.5507/fot.2019.005
You, Q., Wang, Y.L., Yu, P., Kociolek, J.P., Pang, W.T. & Wang, Q.X. (2021) Four new species of monoraphid diatoms from Western Sichuan Plateau in China. Phytotaxa 479: 257–274. https://doi.org/10.11646/phytotaxa.479.3.3
Yu, P., You, Q.M., Kociolek, J.P. & Wang, Q.X. (2019) Three new freshwater species of the genus Achnanthidium (Bacillariophyta, Achnanthidiaceae) from Taiping Lake, China. Fottea, Olomouc 19: 33–49. https://doi.org/10.5507/fot.2018.015
Zhu, H.Z. & Chen, J.Y. (2000) Bacillariophyta of the Xizang Plateau. Science Press, Beijing, China. 353 pp.
Chudaev, D.A. & Georgiev, A.A. (2016) A New taxa of Navicula sensu stricto (Bacillariophyta, Naviculaceae) from high-altitude lake in Tibet, China. Phytotaxa 243: 180–184. https://doi.org/10.11646/phytotaxa.243.2.9
Czarnecki, D.B. & Edlund, M.B. (1995) New combinations for some taxa of Achnanthes. Diatom Research 10: 207–209. https://doi.org/10.1080/0269249X.1995.9705338
Desrosiers, C., Witkowski, A., Riaux-Gobin, C., Zglobicka, I., Kurzydlowski, K.J., Eulin, A., Leflaive, J. & Ten-Hage, L. (2014) Madinithidium gen. nov. (Bacillariophyceae), a new monoraphid diatom genus from the tropical marine coastal zone. Phycologia 53: 583–592. https://doi.org/10.2216/14-21R2
Hlúbiková, D., Ector, L. & Hoffmann, L. (2011) Examination of the type material of some diatom species related to Achnanthidium minutissimum (Kütz.) Czarn. (Bacillariophyceae). Algological Studies 136/137: 19–43. https://doi.org/10.1127/1864-1318/2011/0136-0019
Hustedt, F. (1910) Beitrag zur Algenflora von Afrika. Bacillariales aus Dahomé. Archiv für Hydrobiologies and Planktonkunde 5: 365–382.
Hustedt, F. (1922) Bacillariales aus Innerasien. Gesammelt von Dr. Sven Hedin. In: Hedin, S. (Ed.) Southern Tibet, discoveries in former times compared with my own researches in 1906-1908. Lithographic Institute of the General Staff of the Swedish Army, Stockholm, 6: 107–152. https://doi.org/10.5962/bhl.title.64226
Jao, C., Zhu, H. & Lee, Y. (1974) The fresh-water algae from Mount Qomolangma district (in Tibet). Report of the Scientific Survey of Mount Qomolangma district 1966–1968: 92–126.
Kobayasi, H. (1997) Comparative studies among four linear-lanceolate Achnanthidium species (Bacillariophyceae) with curved terminal raphe endings. Nova Hedwigia 65 (1/4): 147–163. https://doi.org/10.1127/nova.hedwigia/65/1997/147
Kobayasi, H., Idei, M., Mayama, S., Nagumo, T. & Osada, K. (2006) Atlas of Japanese Diatoms based on electron microscopy. Volume 1. Uchida Rokakuko Publishing Co., Tokyo. 59 + 533 pp.
Kobayasi, H., Nagumo, T. & Mayama, S. (1986) Observations on the two rheophilic species of the genus Achnanthes (Bacillariophyceae), A. convergens H. Kob. and A. japonica H. Kob. Diatom 2: 83–93.
Kociolek, J.P., You, Q.M., Yu, P., Li, Y.L., Wang, Y.L., Lowe, R. & Wang, Q.X. (2021) Description of Gomphothidium gen. nov., with light and scanning electron microscopy: a new freshwater monoraphid diatom genus from Asia. Fottea 21: 1–7. https://doi.org/10.5507/fot.2020.011
Krammer, K. & Lange-Bertalot, H. (2004) Bacillariophyceae 4. Teil: Achnanthaceae, Kritische Erganzungen zu Navicula (Lineolatae), Gomphonema Gesamtliteraturverzeichnis [second revised edition] In: Suesswasserflora von Mitteleuropa. Spektrum Akademischer Verlad Heidelberg, 2, 4, 468 pp.
Kulikovskiy, M., Glushchenko, A., Genkal, S.I., Kuznetsova, I.V. & Kociolek, J.P. (2020) Platebaikalia - a new monoraphid diatom genus from ancient Lake Baikal with comments on the genus Platessa. Fottea, Olomouc 20: 58–67. https://doi.org/10.5507/fot.2019.014
Kulikovskiy, M., Lange-Bertalot , H. & Witkowski, A. (2013) Gliwiczia gen. nov. a new monoraphid diatom genus from Lake Baikal with a description of four species new for science. Phytotaxa 109: 1–16. https://doi.org/10.11646/phytotaxa.109.1.1
Kulikovskiy, M., Lange-Bertalot, H. & Kuznetsova, I.V. (2015) Lake Baikal: hotspot of endemic diatoms II. Iconographia Diatomologica 26: 1–656.
Kulikovskiy, M., Maltsev,Y., Glushchenko, A., Kuznetsova, I., Kapustin, D., Gusev, E., Lange-Bertalot, H., Genkal, S. & Kociolek, J.P. (2020) Gogorevia, a new monoraphid diatom genus for Achnanthes exigua and allied taxa (Achnanthidiaceae) described on the basis of an integrated molecular and morphological approach. Journal of Phycology 56: 1601–1613. https://doi.org/10.1111/jpy.13064
Kulikovskiy, M.S., Lange-Bertalot, H., Metzeltin, D. & Witkowski, A. (2012) Lake Baikal: hotspot of endemic diatoms I. Iconographia Diatomologica 23: 7–607.
Kützing, F.T. (1844) Die Kieselschaligen. Bacillarien oder Diatomeen. Nordhausen., 152 pp. https://doi.org/10.5962/bhl.title.64360
Li, S. & Zhang, Q. (2008) Geochemistry of the upper Han River basin, China, 1: Spatial distribution of major ion compositions and their controlling factors. Applied Geochemistry 23: 3535–3544. https://doi.org/10.1016/j.apgeochem.2008.08.012
Lange-Bertalot, H. (1999) Neue Kombinationen von Taxa aus Achnanthes Bory (sensu lato). Iconographia Diatomologica 6: 270–283
Li, Y., Williams, D.M. , Metzeltin, D. , Kociolek, J.P. & Gong, Z. (2010) Tibetiella pulchra gen. nov. et sp. nov. a new freshwater epilithic diatom (Bacillariophyta) from river nujiang in Tibet, China. Journal of Phycology 46: 325–330. https://doi.org/10.1111/j.1529-8817.2009.00776.x
Li, Y., You, Q.M., Kociolek, J.P., Wang, L.Q. & Zhang, W. (2019) Gomphosinica selincuoensis sp. nov. a new diatom (Bacillariophyceae) from North Tibet, China. Phytotaxa 423: 195–205. https://doi.org/10.11646/phytotaxa.423.3.8
Liu, Q., Li, J.J., Nan, F.R., Feng, J., Lv, J.P., Xie, S.L. & Kociolek, J.P. (2019) New and interesting diatoms from Tibet: I. Description of a new species of Clipeoparvus Woodbridge et al. Diatom Research 34: 33–38. https://doi.org/10.1080/0269249X.2019.1578697
Liu, Q., Xiang, Y.D., Yu, P., Xie, S.L. & Kociolek, J.P. (2020) New and interesting diatoms from Tibet. II. Description of two new species of monoraphid diatoms. Diatom Research 35: 353–361. https://doi.org/10.1080/0269249X.2020.1810781
Liu, Q., Yang, L., Nan, F.R., Feng, J., Lv, J.P., Kociolek, J.P. & Xie, S.L. (2018) A new diatom species of Aneumastus D. G. Mann & Stickle ((Bacillariophyta, Bacillariophyceae, Mastogloiales, Mastogloiaceae) from Tibet, China. Phytotaxa 373: 231–235. https://doi.org/10.11646/phytotaxa.373.3.6
Liu, Y., Kociolek, J.P., Liu, Q., Tan, X. & Fan, Y.W. (2020) A new aerophilic Neidium Pfister (Neidiaceae, Bacillariophyta) species from Guangxi Zhuang Autonomous Region, China. Phytotaxa 432 (2): 171–180. https://doi.org/10.11646/phytotaxa.432.2.6
Liu, Y., Tan, X., Kociolek, J.P., Kulikovskiy, M., Lu, X.X. & Fan, Y.W. (2021) One new species of Achnanthidium Kützing (Bacillariophyta, Achnanthidiaceae) from the upper Han River, China. Phytotaxa 516 (2): 187–194. https://doi.org/10.11646/phytotaxa.516.2.6
Monnier, O., Lange-Bertalot, H., Rimet, F., Hoffmann, L. & Ector, L. (2004) Achnanthidium atomoides sp. nov., A new diatom from the Grand-Duchy of Luxembourg. Vie Milieu 54 (2–3): 127–136.
Monnier, O., Lange-Bertalot, H., Hoffmann, L. & Ector, L. (2007) The genera Achnanthidium Ku?tzing and Psammothidium Bukhtiyarova & Round in the family Achnanthidiaceae (Bacillariophyceae): a reappraisal of the differential criteria. Cryptogamie, Algologie 28: 141–158.
Moser, G., Lange-Bertalot, H. & Metzeltin, D. (1998) Insel der Endemiten Geobotanisches Phänomen Neukaledonien (Island of endemics New Caledonia - a geobotanical phenomenon). Bibliotheca Diatomologica. 38: 1–464.
Novais, M.H., Jüttner, I., Hoffmann, L. & Ector, L. (2015) Morphological variability within the Achnanthidium minutissimum species complex (Bacillariophyta): comparison between the type material of Achnanthes minutissima and related taxa, and new freshwater Achnanthidium species from Portugal. Phytotaxa 224: 101–139. https://doi.org/10.11646/phytotaxa.224.2.1
Pérès, F., Barthès, A., Ponton, E., Coste, M., Ten-Hage, L. & Le-Cohu, R. (2012) Achnanthidium delmontii sp. nov., a new species from French rivers. Fottea 12 (2): 189–198. https://doi.org/10.5507/fot.2012.014
Potapova, M. (2006) Achnanthidiumn zhakovschikovii sp. nov. (Bacillariophyta) and related species from rivers of Northwestern Russia. Nova Hedwigia 82 (3–4): 399–408. https://doi.org/10.1127/0029-5035/2006/0082-0399
Round, F.E. (1998) Validation of some previously published “achnanthoid” genera. Diatom Research 13: 181. https://doi.org/10.1080/0269249X.1998.9705442
Round, F.E. & Basson, P.W. (1997) A new monoraphid diatom genus (Pogoneis) from Bahrain and the transfer of previously described species A. hungarica and A. taeniata to new genera. Diatom Research 12: 71–81. https://doi.org/10.1080/0269249X.1997.9705403
Round, F.E. & Bukhtiyarova, L. (1996) Four new genera based on Achnanthes (Achnanthidium) together with a re-definition of Achnanthidium. Diatom Research 11 (2): 345–361. https://doi.org/10.1080/0269249X.1996.9705389
Round, F.E., Crawford, R.M. & Mann, D.G. (1990) The Diatoms. Biology and Morphology of the Genera, Cambridge University Press, Cambridge. 747 pp.
Silva, P.C. (1962) Classification of Algae. Appendix A. In: Ralph, A.L. (Ed.) Physiology and Biochemstry of Algae. Academic Press. N. R., London. pp. 827–837.
Simonsen, R. (1987) Atlas and Catalogue of the Diatom Types of Friedrich Hustedt. 3 Volumes. J. Cramer, Berlin.
Smith, W. (1855) Notes of an excursion to the south of France and the Auvergne in search of Diatomaceae. Annals and Magazine of Natural History 15: 1–9. https://doi.org/10.1080/037454809495381
Van de Vijver, B., Ector, L., Beltrami, M.E., De Haan, M., Falasco, E., Hlúbiková, D., Jarlman, A., Kelly, M., Novais, M.H. & Wojtal, A.Z. (2011) A critical analysis of the type material of Achnanthidium lineare W. Smith (Bacillariophyceae). Algological Studies 136/137: 167–191. https://doi.org/10.1127/1864-1318/2011/0136-0167
Wojtal, A.Z. (2013) Species composition and distribution of diatom assemblages in spring waters from various geological formations in Southern Poland. Bibliotheca Diatomologica 59: 1–436.
You, Q.M., Cao, Y., Yu, P., Kociolek, J.P., Zhang, L.X., Wu, B., Lowe, R. & Wang, Q.X. (2019) Three new subaerial Achnanthidium (Bacillariophyta) species from a karst landform in the Guizhou Province, China. Fottea 19: 138–150. https://doi.org/10.5507/fot.2019.005
You, Q., Wang, Y.L., Yu, P., Kociolek, J.P., Pang, W.T. & Wang, Q.X. (2021) Four new species of monoraphid diatoms from Western Sichuan Plateau in China. Phytotaxa 479: 257–274. https://doi.org/10.11646/phytotaxa.479.3.3
Yu, P., You, Q.M., Kociolek, J.P. & Wang, Q.X. (2019) Three new freshwater species of the genus Achnanthidium (Bacillariophyta, Achnanthidiaceae) from Taiping Lake, China. Fottea, Olomouc 19: 33–49. https://doi.org/10.5507/fot.2018.015
Zhu, H.Z. & Chen, J.Y. (2000) Bacillariophyta of the Xizang Plateau. Science Press, Beijing, China. 353 pp.