Abstract
Graptopetalum trujilloi (subg. Glassia, Crassulaceae), a new haplostemonous species from San Gabriel municipality, Jalisco, Mexico is described and illustrated. Its morphologically closest species is G. rosanevadoense. It differs from the latter in having a smaller habit, more numerous primary branches per inflorescence, larger mean flower density per 10 cm of the scape, fewer flowers per basal branch, smaller flower diameter, petals greenish with transversal inconspicuous red stripes, narrower petals, filaments greenish to distally reddish and carpels abaxially obtuse. A key to the haplostemonous species of Graptopetalum is provided and, according to IUCN criteria, the species was assessed as critically endangered (CR).
References
<p>Acevedo-Rosas, R., Cameron, K., Sosa, V. & Pell, S. (2004a) A molecular phylogenetic study of <em>Graptopetalum</em> (Crassulaceae) based on ETS, ITS, <em>rpl</em>16, and <em>trn</em>L-F nucleotide sequences. <em>American Journal of Botany</em> 91: 1099–1104. https://doi.org/10.3732/ajb.91.7.1099</p>
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<p>Acevedo-Rosas, R., Cházaro-Basáñez, M.J., Jimeno-Sevilla, H.D. & Zuno-Delgadillo, O. (2018) Crassulaceae from Western-Central Mexico. <em>Cactus and Succulent Journal</em> 90: 146–151. https://doi.org/10.2985/015.090.0201</p>
<p>Acevedo-Rosas, R., Sosa, V. & Lorea, F.G. (2004b) Phylogenetic relationships and morphological patterns in <em>Graptopetalum</em> (Crassulaceae). <em>Brittonia</em> 56: 185–194. https://doi.org/10.1663/0007-196X(2004)056[0185:PRAMPI]2.0.CO;2</p>
<p>Bachman, S., Moat, J., Hill, A.W., De La Torre, J. & Scott, B. (2011) Supporting red list threat assessments with GeoCAT: geospatial conservation assessment tool. <em>ZooKeys</em> 150: 117. https://doi.org/10.3897/zookeys.150.2109</p>
<p>Benavides-Solorio, J. de D., González-Guillén, M. de J., López-Paniagua, C. & Valdez-Lazalde, J.R. (2008) Oferta hídrica de la cuenca forestal Tapalpa, Jalisco, orientada hacia los servicios ambientales. <em>Madera y Bosques</em> 14: 5–28. https://doi.org/10.21829/myb.2008.1421209</p>
<p>Berger, A. (1930) Crassulaceae.<em> In: </em>Engler, A., Prantl, K. <em>Die natürlichen Pflanzenfamilien 2</em>, W. Engelmann, Leipzig, pp. 352–483.</p>
<p>Byalt, V.V. [Бялт, В.В.] (2012) Новая система рода <em>Graptopetalum</em> Rose (Crassulaceae) [A new system of the genus <em>Graptopetalum </em>Rose (Crassulaceae)]. <em>Новости систематики высших растений</em> [<em>Novitates Systematicae Plantarum Vascularium</em>] 43: 70–83.</p>
<p>Cházaro-Basáñez, M.J. & Acevedo-Rosas, R. (2008) <em>Graptopetalum</em> <em>glassii</em>, recently described from Colima, Mexico. <em>Cactus and Succulents Journal </em>(<em>US</em>) 80: 202–203. https://doi.org/10.2985/0007-9367(2008)80[202:GG]2.0.CO;2</p>
<p>Cházaro-Basáñez, M.J., Acevedo-Rosas, R. & Machuca-Núñez, J.A. (2012) On <em>Graptopetalum marginatum</em> (Crassulaceae). <em>Cactus and Succulents Journal</em> <em>(US</em>) 84: 155–158. https://doi.org/10.2985/0007-9367-84.3.155</p>
<p>Cházaro-Basáñez, M.J. & Flores, A. (1992) <em>Graptopetalum pentandrum </em>subsp. <em>superbum</em> Kimnach from the wild. <em>Cactus and Succulents Journal </em>(<em>US</em>) 64: 187–189.</p>
<p>Cházaro-Basáñez, M.J. & Flores, A. (1999) <em>Graptopetalum amethystinum </em>(Rose) Walther, rediscovered! <em>International Cactus Adventures</em> 43: 9–12.</p>
<p>Cházaro-Basáñez, M.J., García-Ruíz, I., Flores-Macías, A. & Machuca-Núñez, J.A. (2004) In Situ: <em>Graptopetalum pentandrum</em> Moran. <em>International Cactus Adventures</em> 61: 12–17.</p>
<p>Cházaro-Basáñez, M.J. & Thiede, J. (1995) Floristic and phytogeographic studies on the Crassulaceae of Jalisco (Mexico).<em> In: </em>T´Hart, H. & Eggli, U. (Eds.) <em>Evolution and systematics on the Crassulaceae</em>. Backhuys Publishers, Leiden, The Netherlands, pp. 89–123.</p>
<p>Christenhusz, M. (2020) On species concepts, phylogenetics and the science of natural history—three current issues facing taxonomy. <em>Megataxa</em> 1: 67–72. https://doi.org/10.11646/megataxa.1.1.14</p>
<p>De Queiroz, K. (2007) Species Concepts and Species Delimitation. <em>Systematic Biology</em> 56: 879–886. https://doi.org/10.1080/10635150701701083</p>
<p>Greenland, D. (2005) Mountain and valleywinds.<em> In: </em>Oliver, J.E. (Ed.) <em>Encyclopedia of World Climatology</em>. Springer, Dordrecht, The Netherlands, pp. 516–523.</p>
<p>IUCN (2019) <em>Guidelines for using the IUCN Red List Categories and Criteria</em>. Version 14. Prepared by the Standards and Petitions Committee<em>. </em>Available from: http://www.iucnredlist.org/documents/RedListGuidelines.pdf (accessed 1 February 2022)</p>
<p>Kimnach, M. (1987) A new succulent from Mexico: <em>Graptopetalum pentandrum</em> subsp. <em>superbum</em>. <em>Cactus & Succulent Journal (US)</em> 59: 140–143.</p>
<p>Kimnach, M. & Moran, R. (2002) G<em>raptopetalum marginatum</em>, a new species from Nayarit, Mexico. <em>Cactus & Succulent Journal (US)</em> 74: 196–198.</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum</em>, Laurentius Salvius, Stockholm, 560 pp.</p>
<p>Lomelí-Sención, J.A. (1988) <em>Graptopetalum fruticosum</em> (Crassulaceae), en el sur de Jalisco. <em>Cactáceas y Suculentas Mexicanas</em> 33: 89–91.</p>
<p>Mayuzumi, S. & Ohba, H. (2004) The phylogenetic position of eastern Asia <em>Sedoideae</em> (Crassulaceae) inferred from chloroplast and nuclear DNA sequences. <em>Systematic Botany</em> 29: 587–598. https://doi.org/10.1600/0363644041744329</p>
<p>Moran, R. (1971) <em>Graptopetalum pentandrum</em> a new species of Mexico. <em>Cactus and Succulent Journal</em> (<em>US</em>) 43: 255–258.</p>
<p>Moran, R. & Uhl, Ch. (1968) <em>Graptopetalum fruticosum</em>, a new species from Jalisco, Mexico. <em>Cactus and Succulents Journal</em> (<em>US</em>) 40: 152–156.</p>
<p>Morrone, J.J., Escalante, T. & Rodríguez-Tapia, G. (2017) Mexican biogeographic provinces: Map and shapefiles. <em>Zootaxa</em> 4277: 277–279. https://doi.org/10.11646/zootaxa.4277.2.8</p>
<p>Mort, M.E., Soltis, D.E., Soltis, P.S., Francisco-Ortega, J. & Santos-Guerra, A. (2001) Phylogenetic relationships and evolution of Crassulaceae inferred from matK sequence data. <em>American Journal of Botany</em> 88: 76–91. https://doi.org/10.2307/2657129</p>
<p>NaturaLista (2021) <em>Graptopetalum glassii</em>. Consultado el 21 de julio de 2021, Comisión Nacional para el Conocimiento y Uso de la Biodiversidad. [https://www.naturalista.mx/taxa/1145953-Graptopetalum-glassii]</p>
<p>Rose, J.N. (1905) <em>Pachyphytum amethystinum</em>. <em>North American Flora</em> 22: 11.</p>
<p>Rose, J.N. (1911) <em>Graptopetalum</em>, a new genus, Crassulaceae. Studies of Mexican and Central American Plants-No. 7, <em>Contributions from the United States National Herbarium</em> 13: 296.</p>
<p>Rose, J.N. (1922) <em>Graptopetalum pachyphyllum</em>: thick leaved Graptopetalum. <em>Addisonia</em> 7: 45–46.</p>
<p>Secretaría de Programación y Presupuesto (1981a) <em>Síntesis geográfica del Estado de Jalisco (anexo cartográfico escala 1:1’000,000).</em> Secretaría de Programación y Presupuesto (SPP) Coordinación General de los Servicios Nacional de Estadística, Geografía e Informática, Mexico, DF. 22 pp.</p>
<p>Secretaría de Programación y Presupuesto (1981b) <em>Síntesis geográfica de Colima.</em> SPP, Coordinación Nacional de Estadística, Geografía e Informática, Mexico, D. F. 131 pp.</p>
<p>SEMARNAT (2005) <em>Manifestación de Impacto Ambiental, Salada Zona Norte, Ampliación de cantera de yeso.</em> Exploración Yeso S. A. de C. V., SINAT, México. 106 pp. [http://sinat.semarnat.gob.mx/dgiraDocs/documentos/col/estudios/2005/06CL2005MD028.pdf]</p>
<p>SEMARNAT (2010) Norma Oficial Mexicana NOM-059-SEMARNAT-2010, Protección ambiental-Especies nativas de México de flora y fauna silvestres-Categorías de riesgo y especificaciones para su inclusión, exclusión o cambio-Lista de especies en riesgo. Secretaría del Medio Ambiente y Recursos Naturales. Diario Oficial de la Federación. Cd. Mx., México. Available from: http://dof.gob.mx/ nota_detalle.php?codigo=5173091&fecha=30/12/2010 (accessed 9 September 2021)</p>
<p>Thiede, J. (2003) <em>Graptopetalum</em>.<em> In: </em>Eggli, U. (Ed.) <em>Illustrated handbook of succulent plants</em>, <em>Vol. VI, Crassulaceae</em>. Springer, Berlin, Heidelberg, pp. 128–134. https://doi.org/10.2307/2657129</p>
<p>Thiede, J. & Eggli, U. (2007) Crassulaceae.<em> In: </em>Kubitzki, K. (Ed.) <em>Flowering Plants· Eudicots, The families and genera of vascular plants</em>, vol. 9, 9. Springer, Berlin, Heidelberg, pp. 83–118. https://doi.org/10.1007/978-3-540-32219-1_12</p>
<p>Vázquez-García, J.A., Acevedo-Rosas, R., Cházaro-Basáñez, M., Nieves-Hernández, G. & Sahagun-Godínez, E. (2021a) <em>Graptopetalum rosanevadoense </em>(Crassulaceae): A new haplostemonous species from the Nevado de Colima, Jalisco, Mexico. <em>Phytotaxa</em> 496: 159–169. https://doi.org/10.11646/phytotaxa.496.2.5</p>
<p>Vázquez-García, J.A., Muñiz-Castro, M.A., Dahua-Machoa, A., Osorio-Muñoz, E.A., Hernández-Vera, G., Ortega-Peña, A.S., Romo-Campos, R.L., Jacobo-Pereira, C., Álvarez de Román, N. & Shalisko, V. (2021b) How to Save Endangered Magnolias? From Population Biology to Conservation Action: The Case of Allopatric Radiation in Western Mexico. <em>In</em>: Kumar, S. (Ed.) <em>Endangered Plants</em>. IntechOpen. https://doi.org/10.5772/intechopen.94346</p>
<p>Vázquez-García, J.A., Muñiz-Castro, M.A, Sahagún-Godínez, E., Cházaro-Basáñez, M., De Castro-Arce, E., Nieves-Hernández, G. & Padilla-Lepe, J. (2013) Four New Species of <em>Agave</em> (Agavaceae) of the Marmoratae Group. <em>Systematic Botany</em> 38: 320–331. https://doi.org/10.1600/036364413X666642</p>
<p>Vega Aviña, R., Delgado Vargas, F. & Pío León, J.F. (2020) <em>Graptopetalum sinaloensis </em>(Crassulaceae), una nueva especie de Sinaloa, México. <em>Acta Botánica Mexicana</em> 127: e1550. https://doi.org/10.21829/abm127.2020.1550</p>
<p>Vogelmann, H.W. (1973) Fog precipitation in the cloud forests of eastern Mexico. <em>Bioscience</em> 23: 96–100. https://doi.org/10.2307/1296569</p>
<p>Walther, E. (1931) <em>Graptopetalum amethystinum</em> (Rose) Walther, New combination. <em>Cactus & Succulent Journal of America</em> 3: 73–75.</p>
<p>Acevedo-Rosas, R. & Cházaro-Basáñez, M.J. (2003) A new species and a nomenclature change in <em>Graptopetalum</em> (Crassulaceae). <em>Novon</em> 13: 377–380. https://doi.org/10.2307/3393365</p>
<p>Acevedo-Rosas, R., Cházaro-Basáñez, M.J., Jimeno-Sevilla, H.D. & Zuno-Delgadillo, O. (2018) Crassulaceae from Western-Central Mexico. <em>Cactus and Succulent Journal</em> 90: 146–151. https://doi.org/10.2985/015.090.0201</p>
<p>Acevedo-Rosas, R., Sosa, V. & Lorea, F.G. (2004b) Phylogenetic relationships and morphological patterns in <em>Graptopetalum</em> (Crassulaceae). <em>Brittonia</em> 56: 185–194. https://doi.org/10.1663/0007-196X(2004)056[0185:PRAMPI]2.0.CO;2</p>
<p>Bachman, S., Moat, J., Hill, A.W., De La Torre, J. & Scott, B. (2011) Supporting red list threat assessments with GeoCAT: geospatial conservation assessment tool. <em>ZooKeys</em> 150: 117. https://doi.org/10.3897/zookeys.150.2109</p>
<p>Benavides-Solorio, J. de D., González-Guillén, M. de J., López-Paniagua, C. & Valdez-Lazalde, J.R. (2008) Oferta hídrica de la cuenca forestal Tapalpa, Jalisco, orientada hacia los servicios ambientales. <em>Madera y Bosques</em> 14: 5–28. https://doi.org/10.21829/myb.2008.1421209</p>
<p>Berger, A. (1930) Crassulaceae.<em> In: </em>Engler, A., Prantl, K. <em>Die natürlichen Pflanzenfamilien 2</em>, W. Engelmann, Leipzig, pp. 352–483.</p>
<p>Byalt, V.V. [Бялт, В.В.] (2012) Новая система рода <em>Graptopetalum</em> Rose (Crassulaceae) [A new system of the genus <em>Graptopetalum </em>Rose (Crassulaceae)]. <em>Новости систематики высших растений</em> [<em>Novitates Systematicae Plantarum Vascularium</em>] 43: 70–83.</p>
<p>Cházaro-Basáñez, M.J. & Acevedo-Rosas, R. (2008) <em>Graptopetalum</em> <em>glassii</em>, recently described from Colima, Mexico. <em>Cactus and Succulents Journal </em>(<em>US</em>) 80: 202–203. https://doi.org/10.2985/0007-9367(2008)80[202:GG]2.0.CO;2</p>
<p>Cházaro-Basáñez, M.J., Acevedo-Rosas, R. & Machuca-Núñez, J.A. (2012) On <em>Graptopetalum marginatum</em> (Crassulaceae). <em>Cactus and Succulents Journal</em> <em>(US</em>) 84: 155–158. https://doi.org/10.2985/0007-9367-84.3.155</p>
<p>Cházaro-Basáñez, M.J. & Flores, A. (1992) <em>Graptopetalum pentandrum </em>subsp. <em>superbum</em> Kimnach from the wild. <em>Cactus and Succulents Journal </em>(<em>US</em>) 64: 187–189.</p>
<p>Cházaro-Basáñez, M.J. & Flores, A. (1999) <em>Graptopetalum amethystinum </em>(Rose) Walther, rediscovered! <em>International Cactus Adventures</em> 43: 9–12.</p>
<p>Cházaro-Basáñez, M.J., García-Ruíz, I., Flores-Macías, A. & Machuca-Núñez, J.A. (2004) In Situ: <em>Graptopetalum pentandrum</em> Moran. <em>International Cactus Adventures</em> 61: 12–17.</p>
<p>Cházaro-Basáñez, M.J. & Thiede, J. (1995) Floristic and phytogeographic studies on the Crassulaceae of Jalisco (Mexico).<em> In: </em>T´Hart, H. & Eggli, U. (Eds.) <em>Evolution and systematics on the Crassulaceae</em>. Backhuys Publishers, Leiden, The Netherlands, pp. 89–123.</p>
<p>Christenhusz, M. (2020) On species concepts, phylogenetics and the science of natural history—three current issues facing taxonomy. <em>Megataxa</em> 1: 67–72. https://doi.org/10.11646/megataxa.1.1.14</p>
<p>De Queiroz, K. (2007) Species Concepts and Species Delimitation. <em>Systematic Biology</em> 56: 879–886. https://doi.org/10.1080/10635150701701083</p>
<p>Greenland, D. (2005) Mountain and valleywinds.<em> In: </em>Oliver, J.E. (Ed.) <em>Encyclopedia of World Climatology</em>. Springer, Dordrecht, The Netherlands, pp. 516–523.</p>
<p>IUCN (2019) <em>Guidelines for using the IUCN Red List Categories and Criteria</em>. Version 14. Prepared by the Standards and Petitions Committee<em>. </em>Available from: http://www.iucnredlist.org/documents/RedListGuidelines.pdf (accessed 1 February 2022)</p>
<p>Kimnach, M. (1987) A new succulent from Mexico: <em>Graptopetalum pentandrum</em> subsp. <em>superbum</em>. <em>Cactus & Succulent Journal (US)</em> 59: 140–143.</p>
<p>Kimnach, M. & Moran, R. (2002) G<em>raptopetalum marginatum</em>, a new species from Nayarit, Mexico. <em>Cactus & Succulent Journal (US)</em> 74: 196–198.</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum</em>, Laurentius Salvius, Stockholm, 560 pp.</p>
<p>Lomelí-Sención, J.A. (1988) <em>Graptopetalum fruticosum</em> (Crassulaceae), en el sur de Jalisco. <em>Cactáceas y Suculentas Mexicanas</em> 33: 89–91.</p>
<p>Mayuzumi, S. & Ohba, H. (2004) The phylogenetic position of eastern Asia <em>Sedoideae</em> (Crassulaceae) inferred from chloroplast and nuclear DNA sequences. <em>Systematic Botany</em> 29: 587–598. https://doi.org/10.1600/0363644041744329</p>
<p>Moran, R. (1971) <em>Graptopetalum pentandrum</em> a new species of Mexico. <em>Cactus and Succulent Journal</em> (<em>US</em>) 43: 255–258.</p>
<p>Moran, R. & Uhl, Ch. (1968) <em>Graptopetalum fruticosum</em>, a new species from Jalisco, Mexico. <em>Cactus and Succulents Journal</em> (<em>US</em>) 40: 152–156.</p>
<p>Morrone, J.J., Escalante, T. & Rodríguez-Tapia, G. (2017) Mexican biogeographic provinces: Map and shapefiles. <em>Zootaxa</em> 4277: 277–279. https://doi.org/10.11646/zootaxa.4277.2.8</p>
<p>Mort, M.E., Soltis, D.E., Soltis, P.S., Francisco-Ortega, J. & Santos-Guerra, A. (2001) Phylogenetic relationships and evolution of Crassulaceae inferred from matK sequence data. <em>American Journal of Botany</em> 88: 76–91. https://doi.org/10.2307/2657129</p>
<p>NaturaLista (2021) <em>Graptopetalum glassii</em>. Consultado el 21 de julio de 2021, Comisión Nacional para el Conocimiento y Uso de la Biodiversidad. [https://www.naturalista.mx/taxa/1145953-Graptopetalum-glassii]</p>
<p>Rose, J.N. (1905) <em>Pachyphytum amethystinum</em>. <em>North American Flora</em> 22: 11.</p>
<p>Rose, J.N. (1911) <em>Graptopetalum</em>, a new genus, Crassulaceae. Studies of Mexican and Central American Plants-No. 7, <em>Contributions from the United States National Herbarium</em> 13: 296.</p>
<p>Rose, J.N. (1922) <em>Graptopetalum pachyphyllum</em>: thick leaved Graptopetalum. <em>Addisonia</em> 7: 45–46.</p>
<p>Secretaría de Programación y Presupuesto (1981a) <em>Síntesis geográfica del Estado de Jalisco (anexo cartográfico escala 1:1’000,000).</em> Secretaría de Programación y Presupuesto (SPP) Coordinación General de los Servicios Nacional de Estadística, Geografía e Informática, Mexico, DF. 22 pp.</p>
<p>Secretaría de Programación y Presupuesto (1981b) <em>Síntesis geográfica de Colima.</em> SPP, Coordinación Nacional de Estadística, Geografía e Informática, Mexico, D. F. 131 pp.</p>
<p>SEMARNAT (2005) <em>Manifestación de Impacto Ambiental, Salada Zona Norte, Ampliación de cantera de yeso.</em> Exploración Yeso S. A. de C. V., SINAT, México. 106 pp. [http://sinat.semarnat.gob.mx/dgiraDocs/documentos/col/estudios/2005/06CL2005MD028.pdf]</p>
<p>SEMARNAT (2010) Norma Oficial Mexicana NOM-059-SEMARNAT-2010, Protección ambiental-Especies nativas de México de flora y fauna silvestres-Categorías de riesgo y especificaciones para su inclusión, exclusión o cambio-Lista de especies en riesgo. Secretaría del Medio Ambiente y Recursos Naturales. Diario Oficial de la Federación. Cd. Mx., México. Available from: http://dof.gob.mx/ nota_detalle.php?codigo=5173091&fecha=30/12/2010 (accessed 9 September 2021)</p>
<p>Thiede, J. (2003) <em>Graptopetalum</em>.<em> In: </em>Eggli, U. (Ed.) <em>Illustrated handbook of succulent plants</em>, <em>Vol. VI, Crassulaceae</em>. Springer, Berlin, Heidelberg, pp. 128–134. https://doi.org/10.2307/2657129</p>
<p>Thiede, J. & Eggli, U. (2007) Crassulaceae.<em> In: </em>Kubitzki, K. (Ed.) <em>Flowering Plants· Eudicots, The families and genera of vascular plants</em>, vol. 9, 9. Springer, Berlin, Heidelberg, pp. 83–118. https://doi.org/10.1007/978-3-540-32219-1_12</p>
<p>Vázquez-García, J.A., Acevedo-Rosas, R., Cházaro-Basáñez, M., Nieves-Hernández, G. & Sahagun-Godínez, E. (2021a) <em>Graptopetalum rosanevadoense </em>(Crassulaceae): A new haplostemonous species from the Nevado de Colima, Jalisco, Mexico. <em>Phytotaxa</em> 496: 159–169. https://doi.org/10.11646/phytotaxa.496.2.5</p>
<p>Vázquez-García, J.A., Muñiz-Castro, M.A., Dahua-Machoa, A., Osorio-Muñoz, E.A., Hernández-Vera, G., Ortega-Peña, A.S., Romo-Campos, R.L., Jacobo-Pereira, C., Álvarez de Román, N. & Shalisko, V. (2021b) How to Save Endangered Magnolias? From Population Biology to Conservation Action: The Case of Allopatric Radiation in Western Mexico. <em>In</em>: Kumar, S. (Ed.) <em>Endangered Plants</em>. IntechOpen. https://doi.org/10.5772/intechopen.94346</p>
<p>Vázquez-García, J.A., Muñiz-Castro, M.A, Sahagún-Godínez, E., Cházaro-Basáñez, M., De Castro-Arce, E., Nieves-Hernández, G. & Padilla-Lepe, J. (2013) Four New Species of <em>Agave</em> (Agavaceae) of the Marmoratae Group. <em>Systematic Botany</em> 38: 320–331. https://doi.org/10.1600/036364413X666642</p>
<p>Vega Aviña, R., Delgado Vargas, F. & Pío León, J.F. (2020) <em>Graptopetalum sinaloensis </em>(Crassulaceae), una nueva especie de Sinaloa, México. <em>Acta Botánica Mexicana</em> 127: e1550. https://doi.org/10.21829/abm127.2020.1550</p>
<p>Vogelmann, H.W. (1973) Fog precipitation in the cloud forests of eastern Mexico. <em>Bioscience</em> 23: 96–100. https://doi.org/10.2307/1296569</p>
<p>Walther, E. (1931) <em>Graptopetalum amethystinum</em> (Rose) Walther, New combination. <em>Cactus & Succulent Journal of America</em> 3: 73–75.</p>