Abstract
An account of the Drepanolejeunea subgenus Rhaphidolejeunea in Thailand has been updated based on herbarium specimens and recent collections from field surveys. Five species of Drepanolejeunea, i.e. Drepanolejeunea commutata, D. fleischeri, D. siamensis, D. spicata, and D. yunnanensis are recognized. Of these, Drepanolejeunea commutata, D. fleischeri, and D. yunnanensis are newly reported for Thailand. A key to species, descriptions, distribution ranges and illustrations of each species are provided. The ecology and geographical distribution of the species are noted. Additionally, Drepanolejeunea longicruris is excluded from the country.
References
Asthana, G. & Shukla, A. (2009) Two epiphyllous species of Drepanolejeunea (Spruce) Schiffn. new to the Indian bryoflora. Journal of Bryology 31: 139–142. https://doi.org/10.1179/037366809X400419
Bai, Y., Chen, S.W., Qian, H.Y., Yu, S.H., Xu, Y.M., Zhang, Z.X., Shen, C., Chen, Y.Q., Zhang, M.Q., Yu, J.P. & Zhu, R.L. (2020) Species diversity of epiphyllous liverworts in Qianjiangyuan National Park, Zhejiang. Biodiversity Science 28: 231–237. https://doi.org/10.17520/biods.2019273
Bakalin, V.A. & Nguyen, V.S. (2016) The checklist of liverworts (Hepaticae) and hornworts (Anthocerotae) of Vietnam updated based on literature survey. Tap Chi Sinh Hoc 38: 480–491. https://doi.org/10.15625/0866-7160/v38n4.8771
Bischler, H. (1968) Monographie du genre Rhaphidolejeunea Herzog. Revue Bryologique et Lichenologique 36: 56–104.
But, P.P.-H. & Gao, C.-H. (1991) Epiphyllous Hepaticae of Kowloon Peninsula (Hong Kong). Tropical Bryology 4: 17–22.
Chandini, V.K., Mufeed, B., Manju, C.N. & Rajesh, K.P. (2020) The genus Drepanolejeunea (Spruce) Schiffn. (Lejeuneaceae; Marchantiophyta) in the Western Ghats with special reference to Kerala. Plant Science Today 7: 476–493. https://doi.org/10.14719/pst.2020.7.3.789
Chantanaorrapint, S., Boonkerd, T. & Thaithong, O. (2004) Checklist of bryophytes at the summit of Khao Luang, Huai Yang waterfall National Park, Prachuap Khiri Khan Province, Thailand. National History Bulletin of the Siam Society 52: 163–179.
Chen, P.C. (1955) Bryophyta nova Sinica. Feddes Repertorium Specierum Novarum Regni Vegetabilis 58: 23–52.
Dandotiya, D., Govindapyari, H., Sumam, S. & Uniyal, P.L. (2011) Checklist of the bryophytes of India. Archive for Bryology 88: 1–126.
Dey, M. & Singh, D.K. (2010) Drepanolejeunea fleischeri (Steph.) Grolle & R.L. Zhu (Hepaticae: Lejeuneaceae): an addition to Indian Himalaya. Geophytology 39: 65–71.
Dey, M. & Singh, D.K. (2012) Epiphyllous liverworts of Eastern Himalaya. Botanical survey of India, Kolkata. 364 pp.
Dey, M. & Singh, D.K. (2016) Three foliicolous taxa of Lejeuneaceae (Marchantiophyta) new to India from Great Nicobar Biosphere Reserve, Andaman and Nicobar Islands. Acta Botanica Hungarica 58: 49–68. https://doi.org/10.1556/034.58.2016.1-2.2
Dey, M., Singh, D. & Singh, D.K. (2013) The genus Drepanolejeunea (Marchantiophyta: Lejeuneaceae) in India with a note on three new additions from Eastern Himalaya. Tropical Biology 35:14–25. https://doi.org/10.11646/bde.35.1.2
Gradstein, S.R. (2018) Amphi-Pacific tropical disjunctions in the bryophyte floras of Asia and the New World. Philippine Journal of Systematic Biology 12: 1–11. https://doi.org/10.26757/pjsb.2018a12012
Grolle, R. (1965) Lebermoose aus Neuguinea. I. Journal of the Hattori Botanical Laboratory 28: 43–54.
Grolle, R. & Zhu, R.-L. (2000) A study of Drepanolejeunea subg. Rhaphidolejeunea (Herzog) Grolle & R.L. Zhu, stat. nov. (Hepaticae, Lejeuneaceae) in China with notes on its species elsewhere. Nova Hedwigia 70: 373–395.
He, Q., Zhu, R.-L., Chantanaorrapint, S., Kornochalert, S. & Printarakul, N. (2012) Drepanolejeunea laciniata (Lejeuneaceae), a new species from northern Thailand. Cryptogamie, Bryologie 33: 291–298. http://dx.doi.org/10.7872/cryb.v33.iss3.2012.291
Herzog, T. (1943) Rhaphidolejeunea Herz., eine neue Lejeuneaceengattung der Indomalaya. Mittheilungen des Thüringischen botanischen Vereins 50: 100–105.
Inuthai, J., Zhu, R.-L. & Chantanaorrapint, S. (2014) Drepanolejeunea actinogyna (Lejeuneaceae), a new species from southern Thailand. The Bryologist 117: 165–169. https://doi.org/10.1639/0007-2745-117.2.165
Inuthai, J., Zhu, R.-L. & Chantanaorrapint, S. (2015) Taxonomic notes on Drepanolejeunea pleiodictya (Marchantiophyta, Lejeuneaceae), a little-known Asiatic species. Polish Botanical Journal 60(1): 13–17. https://doi.org/10.1515/pbj-2015-0012
Kumar, S., Pócs, T. & Singh, S.K. (2015) Two Drepanolejeunea [Marchantiophyta] new to bryoflora of Meghalaya. Indian Journal of Forestry 38 (4): 343–349.
Lai, M.-J., Zhu, R.-L. & Chantanaorrapint, S. (2008) Liverworts and hornworts of Thailand: an updated checklist and bryofloristic accounts. Annales Botanici Fennici 45: 321–341. https://doi.org/10.5735/085.045.0501
Long, D.G. & Rubasinghe, S.C.K. (2014) Liverworts and Hornworts of Sri Lanka: a revised checklist. Ceylon Journal of Science (Biological Sciences) 43: 1–36. http://dx.doi.org/10.4038/cjsbs.v43i1.7280
Mizutani, M. (1966) Epiphyllous species of Lejeuneaceae from Sabah (North Borneo). The Journal of the Hattori Botanical Laboratory 29: 153–170.
Mizutani, M. (1970) Lejeuneaceae, subfamily Lejeunoideae and Cololejeunoideae from Sabah (North Borneo). The Journal of the Hattori Botanical Laboratory 33: 225–265.
Pócs, T. (2012) New or little known epiphyllous liverworts, XVI. A small collection from Laos. Acta Biologica Plantarum Agriensis 2: 5–10.
Pócs, T., Luong, T.-T. & Ho, B.-C. (2013) New or little known epiphyllous liverworts, XVIII. Records from The Bidoup-Núi Bà National Park, Vietnam, with the description of Drepanolejeunea bidoupensis sp. nov. Cryptogamie, Bryologie 34: 287–298. https://doi.org/10.7872/cryb.v34.iss3.2013.287
Pócs, T., Mizutani, M. & Koponen, T. (2019) Bryophyte flora of the Huon Peninsula, Papua New Guinea. LXXX. Cheilolejeunea and Drepanolejeunea, with contributions to Ceratolejeunea, Cololejeunea, Diplasiolejeunea, Lejeunea, Leptolejeunea, Metalejeunea and Microlejeunea (Lejeuneaceae, Marchantiophyta). Acta Bryolichenologica Asiatica 8: 41–84.
Pócs, T., Mohamed, H., Yong, K.-T. & Cheah, Y.-H. (2014) Data to the Malaysian liverwort flora, I. Polish Botanical Journal 59:215–220. https://doi.org/10.2478/pbj-2014-0042
Pócs, T & Podani, J. (2015) Southern Thailand Bryophytes II: Epiphylls from the Phang-Nga area. Acta Botanica Hungarica 57:183–198. http://dx.doi.org/10.1556/ABot.57.2015.1-2.14
Sande Lacoste, C.M. van der (1864) Hepaticae, Jungermannieae Archipelagi Indici. adjectis quibusdam speciebus Japonicis. Annales Musei botanici lugduno-batavi 1: 287–314.
Schiffner, V. (1898) Conspectus Hepaticarum Archipelagi Indici. Batavia, 382 pp.
Schuster, R.M. (1963) An annotated synopsis of the genera and subgenera of Lejeuneaceae. I. Introduction; annotated keys to subfamilies and genera. Nova Hedwigia Beihefte 9: 1–203.
Singh, D., Dey, M. & Singh, D.K. (2010) A Synoptic Flora of Liverworts and Hornworts of Manipur. Nelumbo 52: 9–52.
Singh, D.K. (1996) Hepaticae (Bryophyta). In: P.K. Hajra (Ed.) A contribution to the flora of Namdapha Arunachal Pradesh. Botanical Survey of India, New Delhi, pp. 46–67.
Singh, D.K. & Singh, D. (2016) Epiphyllous Liverworts of India: An Overview. Plant Science Today 3: 157–174. https://doi.org/10.14719/pst.2016.3.2.228
Singh, S.K. & Barbhuiya, H.A. (2012) A compendium to Marchantiophyta and Anthocerotophyta of Assam, India. Archive for Bryology 149: 1–30.
Singh, S.K. & Kumar, S. (2021) Notes on Extended Distribution of Drepanolejeunea spicata and Lejeunea stevensiana (Lejeuneaceae: Marchantiophyta) two Scarcely known Species in India. Acta Botanica Hungarica 63: 213–221. https://doi.org/10.1556/ 034.63.2021.1-2.12
Söderström, L., Hagborg, A., Pócs, T. & von Konrat, M. (2020) Listing the unknown—checklist of liverworts and hornworts of Laos. Bryophyte Diversity and Evolution 42: 19–31. https://doi.org/10.11646/bde.42.1.2
Söderström, L., Hagborg, A., von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Cargill, D.C., da Costa, D.P., Crandall-Stotler, B.J., Cooper, E.D., Dauphin, G., Engel, J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A.L., Katagiri, T., Konstantinova, N.A., Larraín, J., Long, D., Nebel, M., Pócs, T., Puche, F., Reiner-Drehwald, E., Renner, M., Sass-Gyarmati, A., Schäfer-Verwimp, A., Segarra-Moragues, J.G., Stotler, R.E., Sukkharak, P., Thiers, B., Uribe, J., Vá?a, J., Villarreal, J., Wigginton, M., Zhang, L. & Zhu, R.-L. (2016) World checklist of hornworts and liverworts. PhytoKeys 59: 1–821. https://doi.org/10.3897/phytokeys.59.6261
Spruce, R. (1884) Hepaticae Amazonicae et Andinae. I. Transactions and Proceedings of the Botanical Society of Edinburgh 15: 1–308.
Stephani, F. (1891a) Hepaticae africanae. Hedwigia 30: 201–217.
Stephani, F. (1891b) Hepaticae africanae. Hedwigia 30: 265–272.
Stephani, F. (1896) Hepaticarum species novae IX. Hedwigia 35: 73–140.
Stephani, F. (1913) Species Hepaticarum 5. Genève & Bale, George & Cie, pp. 177–448.
Thiers, B. (2021) Index herbariorum: A global directory of public herbaria and associates staff. New York Botanical Garden’s Virtual Herbarium. [http://sweetgum.nybg.org/science/ ih/]
Tixier, P. (1970a) Bryophytae Indosinicae. Bryophytes of Thai Tenasserim. The Natural History Bulletin of the Siam Society 23: 541–560.
Tixier, P. (1970b) Bryophytae Indosinicae. Bryophytes du Laos méridional (Paksé et Bolovens). Annales de la Faculté des Sciences, Université de Phnom Penh 3: 153–172.
Tixier, P. (1973) Bryophytae Indosinicae. Liverworts collected in Thailand. The Natural History Bulletin of the Siam Society 24: 449–456.
Tixier, P. (1979) Bryogéographie du Mont Bokor (Cambodge) (Bryophyta indosinica XXIV). Bryophytorum Bibliotheca 18: 5–121.
Udar, R. & Awasthi, U.S. (1984) The genus Rhaphidolejeunea Hork. in India. Yushania 1: 15–17.
Verma, P.K., Alam, A. & Rawat, K.K. (2013) Assessment of liverwort and hornwort flora of Nilgiri Hills, Western Ghats (India). Polish Botanical Journal 58: 525–537. https://doi.org/10.2478/pbj-2013-0038
Yang, J.-D. & Lin, S.H. (2012) Two Species of Drepanolejeunea (Family Lejeuneaceae) Newly Recorded to Taiwan. Taiwan Journal of Biodiversity 14: 63–73.
Zartman, C.E., Sierra, A.M. & Zhu, R.-L. (2019) Drepanolejeunea robinsonii (Lejeuneaceae: Drepanolejeuneinae), a new epiphyllous species from the Amazon forest. The Bryologist 122: 292–296. https://doi.org/10.1639/0007-2745-122.2.292
Zhu, R.-L. (2006) New Checklist of Chinese liverworts, hornworts, and takakiophytes. 1<sup>st </sup>version in 2000; 3<sup>rd</sup> version in Jan 2006 (online). Available from: https://www.yumpu.com/it/document/read/4359880/new-checklist-of-chinese-liverworts-hornworts-and-takakiophytes (accessed 16 June 2021)
Zhu, R.-L., Long, D.G. & Yamaguchi, T. (1999) Cololejeunea yulensis (Steph.) Benedix and Rhaphidolejeunea spicata (Steph.) Grolle new to Japan, with taxonomic notes on Cololejeunea obliqua (Nees & Mont.) Schiffn. Journal of Bryology 21: 293–297. https://doi.org/10.1179/jbr.1999.21.4.293
Zhu, R.-L. & So, M.-L. (2001) Epiphyllous liverworts of China. Nova Hedwig Beihefte 121: 1–418.
Zhu, R.-L. & So, M.-L. (2003) Liverworts and hornworts of Shangsi County of Guangxi (Kwangsi), with an updated checklist of the hepatic flora of Guangxi Province of China. Cryptogamie, Bryologie, 24: 319–334.
Bai, Y., Chen, S.W., Qian, H.Y., Yu, S.H., Xu, Y.M., Zhang, Z.X., Shen, C., Chen, Y.Q., Zhang, M.Q., Yu, J.P. & Zhu, R.L. (2020) Species diversity of epiphyllous liverworts in Qianjiangyuan National Park, Zhejiang. Biodiversity Science 28: 231–237. https://doi.org/10.17520/biods.2019273
Bakalin, V.A. & Nguyen, V.S. (2016) The checklist of liverworts (Hepaticae) and hornworts (Anthocerotae) of Vietnam updated based on literature survey. Tap Chi Sinh Hoc 38: 480–491. https://doi.org/10.15625/0866-7160/v38n4.8771
Bischler, H. (1968) Monographie du genre Rhaphidolejeunea Herzog. Revue Bryologique et Lichenologique 36: 56–104.
But, P.P.-H. & Gao, C.-H. (1991) Epiphyllous Hepaticae of Kowloon Peninsula (Hong Kong). Tropical Bryology 4: 17–22.
Chandini, V.K., Mufeed, B., Manju, C.N. & Rajesh, K.P. (2020) The genus Drepanolejeunea (Spruce) Schiffn. (Lejeuneaceae; Marchantiophyta) in the Western Ghats with special reference to Kerala. Plant Science Today 7: 476–493. https://doi.org/10.14719/pst.2020.7.3.789
Chantanaorrapint, S., Boonkerd, T. & Thaithong, O. (2004) Checklist of bryophytes at the summit of Khao Luang, Huai Yang waterfall National Park, Prachuap Khiri Khan Province, Thailand. National History Bulletin of the Siam Society 52: 163–179.
Chen, P.C. (1955) Bryophyta nova Sinica. Feddes Repertorium Specierum Novarum Regni Vegetabilis 58: 23–52.
Dandotiya, D., Govindapyari, H., Sumam, S. & Uniyal, P.L. (2011) Checklist of the bryophytes of India. Archive for Bryology 88: 1–126.
Dey, M. & Singh, D.K. (2010) Drepanolejeunea fleischeri (Steph.) Grolle & R.L. Zhu (Hepaticae: Lejeuneaceae): an addition to Indian Himalaya. Geophytology 39: 65–71.
Dey, M. & Singh, D.K. (2012) Epiphyllous liverworts of Eastern Himalaya. Botanical survey of India, Kolkata. 364 pp.
Dey, M. & Singh, D.K. (2016) Three foliicolous taxa of Lejeuneaceae (Marchantiophyta) new to India from Great Nicobar Biosphere Reserve, Andaman and Nicobar Islands. Acta Botanica Hungarica 58: 49–68. https://doi.org/10.1556/034.58.2016.1-2.2
Dey, M., Singh, D. & Singh, D.K. (2013) The genus Drepanolejeunea (Marchantiophyta: Lejeuneaceae) in India with a note on three new additions from Eastern Himalaya. Tropical Biology 35:14–25. https://doi.org/10.11646/bde.35.1.2
Gradstein, S.R. (2018) Amphi-Pacific tropical disjunctions in the bryophyte floras of Asia and the New World. Philippine Journal of Systematic Biology 12: 1–11. https://doi.org/10.26757/pjsb.2018a12012
Grolle, R. (1965) Lebermoose aus Neuguinea. I. Journal of the Hattori Botanical Laboratory 28: 43–54.
Grolle, R. & Zhu, R.-L. (2000) A study of Drepanolejeunea subg. Rhaphidolejeunea (Herzog) Grolle & R.L. Zhu, stat. nov. (Hepaticae, Lejeuneaceae) in China with notes on its species elsewhere. Nova Hedwigia 70: 373–395.
He, Q., Zhu, R.-L., Chantanaorrapint, S., Kornochalert, S. & Printarakul, N. (2012) Drepanolejeunea laciniata (Lejeuneaceae), a new species from northern Thailand. Cryptogamie, Bryologie 33: 291–298. http://dx.doi.org/10.7872/cryb.v33.iss3.2012.291
Herzog, T. (1943) Rhaphidolejeunea Herz., eine neue Lejeuneaceengattung der Indomalaya. Mittheilungen des Thüringischen botanischen Vereins 50: 100–105.
Inuthai, J., Zhu, R.-L. & Chantanaorrapint, S. (2014) Drepanolejeunea actinogyna (Lejeuneaceae), a new species from southern Thailand. The Bryologist 117: 165–169. https://doi.org/10.1639/0007-2745-117.2.165
Inuthai, J., Zhu, R.-L. & Chantanaorrapint, S. (2015) Taxonomic notes on Drepanolejeunea pleiodictya (Marchantiophyta, Lejeuneaceae), a little-known Asiatic species. Polish Botanical Journal 60(1): 13–17. https://doi.org/10.1515/pbj-2015-0012
Kumar, S., Pócs, T. & Singh, S.K. (2015) Two Drepanolejeunea [Marchantiophyta] new to bryoflora of Meghalaya. Indian Journal of Forestry 38 (4): 343–349.
Lai, M.-J., Zhu, R.-L. & Chantanaorrapint, S. (2008) Liverworts and hornworts of Thailand: an updated checklist and bryofloristic accounts. Annales Botanici Fennici 45: 321–341. https://doi.org/10.5735/085.045.0501
Long, D.G. & Rubasinghe, S.C.K. (2014) Liverworts and Hornworts of Sri Lanka: a revised checklist. Ceylon Journal of Science (Biological Sciences) 43: 1–36. http://dx.doi.org/10.4038/cjsbs.v43i1.7280
Mizutani, M. (1966) Epiphyllous species of Lejeuneaceae from Sabah (North Borneo). The Journal of the Hattori Botanical Laboratory 29: 153–170.
Mizutani, M. (1970) Lejeuneaceae, subfamily Lejeunoideae and Cololejeunoideae from Sabah (North Borneo). The Journal of the Hattori Botanical Laboratory 33: 225–265.
Pócs, T. (2012) New or little known epiphyllous liverworts, XVI. A small collection from Laos. Acta Biologica Plantarum Agriensis 2: 5–10.
Pócs, T., Luong, T.-T. & Ho, B.-C. (2013) New or little known epiphyllous liverworts, XVIII. Records from The Bidoup-Núi Bà National Park, Vietnam, with the description of Drepanolejeunea bidoupensis sp. nov. Cryptogamie, Bryologie 34: 287–298. https://doi.org/10.7872/cryb.v34.iss3.2013.287
Pócs, T., Mizutani, M. & Koponen, T. (2019) Bryophyte flora of the Huon Peninsula, Papua New Guinea. LXXX. Cheilolejeunea and Drepanolejeunea, with contributions to Ceratolejeunea, Cololejeunea, Diplasiolejeunea, Lejeunea, Leptolejeunea, Metalejeunea and Microlejeunea (Lejeuneaceae, Marchantiophyta). Acta Bryolichenologica Asiatica 8: 41–84.
Pócs, T., Mohamed, H., Yong, K.-T. & Cheah, Y.-H. (2014) Data to the Malaysian liverwort flora, I. Polish Botanical Journal 59:215–220. https://doi.org/10.2478/pbj-2014-0042
Pócs, T & Podani, J. (2015) Southern Thailand Bryophytes II: Epiphylls from the Phang-Nga area. Acta Botanica Hungarica 57:183–198. http://dx.doi.org/10.1556/ABot.57.2015.1-2.14
Sande Lacoste, C.M. van der (1864) Hepaticae, Jungermannieae Archipelagi Indici. adjectis quibusdam speciebus Japonicis. Annales Musei botanici lugduno-batavi 1: 287–314.
Schiffner, V. (1898) Conspectus Hepaticarum Archipelagi Indici. Batavia, 382 pp.
Schuster, R.M. (1963) An annotated synopsis of the genera and subgenera of Lejeuneaceae. I. Introduction; annotated keys to subfamilies and genera. Nova Hedwigia Beihefte 9: 1–203.
Singh, D., Dey, M. & Singh, D.K. (2010) A Synoptic Flora of Liverworts and Hornworts of Manipur. Nelumbo 52: 9–52.
Singh, D.K. (1996) Hepaticae (Bryophyta). In: P.K. Hajra (Ed.) A contribution to the flora of Namdapha Arunachal Pradesh. Botanical Survey of India, New Delhi, pp. 46–67.
Singh, D.K. & Singh, D. (2016) Epiphyllous Liverworts of India: An Overview. Plant Science Today 3: 157–174. https://doi.org/10.14719/pst.2016.3.2.228
Singh, S.K. & Barbhuiya, H.A. (2012) A compendium to Marchantiophyta and Anthocerotophyta of Assam, India. Archive for Bryology 149: 1–30.
Singh, S.K. & Kumar, S. (2021) Notes on Extended Distribution of Drepanolejeunea spicata and Lejeunea stevensiana (Lejeuneaceae: Marchantiophyta) two Scarcely known Species in India. Acta Botanica Hungarica 63: 213–221. https://doi.org/10.1556/ 034.63.2021.1-2.12
Söderström, L., Hagborg, A., Pócs, T. & von Konrat, M. (2020) Listing the unknown—checklist of liverworts and hornworts of Laos. Bryophyte Diversity and Evolution 42: 19–31. https://doi.org/10.11646/bde.42.1.2
Söderström, L., Hagborg, A., von Konrat, M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Cargill, D.C., da Costa, D.P., Crandall-Stotler, B.J., Cooper, E.D., Dauphin, G., Engel, J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A.L., Katagiri, T., Konstantinova, N.A., Larraín, J., Long, D., Nebel, M., Pócs, T., Puche, F., Reiner-Drehwald, E., Renner, M., Sass-Gyarmati, A., Schäfer-Verwimp, A., Segarra-Moragues, J.G., Stotler, R.E., Sukkharak, P., Thiers, B., Uribe, J., Vá?a, J., Villarreal, J., Wigginton, M., Zhang, L. & Zhu, R.-L. (2016) World checklist of hornworts and liverworts. PhytoKeys 59: 1–821. https://doi.org/10.3897/phytokeys.59.6261
Spruce, R. (1884) Hepaticae Amazonicae et Andinae. I. Transactions and Proceedings of the Botanical Society of Edinburgh 15: 1–308.
Stephani, F. (1891a) Hepaticae africanae. Hedwigia 30: 201–217.
Stephani, F. (1891b) Hepaticae africanae. Hedwigia 30: 265–272.
Stephani, F. (1896) Hepaticarum species novae IX. Hedwigia 35: 73–140.
Stephani, F. (1913) Species Hepaticarum 5. Genève & Bale, George & Cie, pp. 177–448.
Thiers, B. (2021) Index herbariorum: A global directory of public herbaria and associates staff. New York Botanical Garden’s Virtual Herbarium. [http://sweetgum.nybg.org/science/ ih/]
Tixier, P. (1970a) Bryophytae Indosinicae. Bryophytes of Thai Tenasserim. The Natural History Bulletin of the Siam Society 23: 541–560.
Tixier, P. (1970b) Bryophytae Indosinicae. Bryophytes du Laos méridional (Paksé et Bolovens). Annales de la Faculté des Sciences, Université de Phnom Penh 3: 153–172.
Tixier, P. (1973) Bryophytae Indosinicae. Liverworts collected in Thailand. The Natural History Bulletin of the Siam Society 24: 449–456.
Tixier, P. (1979) Bryogéographie du Mont Bokor (Cambodge) (Bryophyta indosinica XXIV). Bryophytorum Bibliotheca 18: 5–121.
Udar, R. & Awasthi, U.S. (1984) The genus Rhaphidolejeunea Hork. in India. Yushania 1: 15–17.
Verma, P.K., Alam, A. & Rawat, K.K. (2013) Assessment of liverwort and hornwort flora of Nilgiri Hills, Western Ghats (India). Polish Botanical Journal 58: 525–537. https://doi.org/10.2478/pbj-2013-0038
Yang, J.-D. & Lin, S.H. (2012) Two Species of Drepanolejeunea (Family Lejeuneaceae) Newly Recorded to Taiwan. Taiwan Journal of Biodiversity 14: 63–73.
Zartman, C.E., Sierra, A.M. & Zhu, R.-L. (2019) Drepanolejeunea robinsonii (Lejeuneaceae: Drepanolejeuneinae), a new epiphyllous species from the Amazon forest. The Bryologist 122: 292–296. https://doi.org/10.1639/0007-2745-122.2.292
Zhu, R.-L. (2006) New Checklist of Chinese liverworts, hornworts, and takakiophytes. 1<sup>st </sup>version in 2000; 3<sup>rd</sup> version in Jan 2006 (online). Available from: https://www.yumpu.com/it/document/read/4359880/new-checklist-of-chinese-liverworts-hornworts-and-takakiophytes (accessed 16 June 2021)
Zhu, R.-L., Long, D.G. & Yamaguchi, T. (1999) Cololejeunea yulensis (Steph.) Benedix and Rhaphidolejeunea spicata (Steph.) Grolle new to Japan, with taxonomic notes on Cololejeunea obliqua (Nees & Mont.) Schiffn. Journal of Bryology 21: 293–297. https://doi.org/10.1179/jbr.1999.21.4.293
Zhu, R.-L. & So, M.-L. (2001) Epiphyllous liverworts of China. Nova Hedwig Beihefte 121: 1–418.
Zhu, R.-L. & So, M.-L. (2003) Liverworts and hornworts of Shangsi County of Guangxi (Kwangsi), with an updated checklist of the hepatic flora of Guangxi Province of China. Cryptogamie, Bryologie, 24: 319–334.