Abstract
Lopholejeunea (Stephani 1890: 285) is a pantropical genus with 42 species and one variety (Söderström et al., 2016), of which 12 species have been recorded from Madagascar (Marline et al., 2012). During examination of various Lopholejeunea materials from Madagascar, a few nomenclatural changes became necessary. As a result, ten species are accepted for Madagascar: Lopholejeunea borbonica Stephani (1896: 109), L. erugata Thiers (1984: 174), L. eulopha (Taylor 1846: 391.) Schiffner (1893: 129), L. grandicrista Stephani (1893: 34), L. leioptera Sass-Gyarmati (2001: 80), L. lepidoscypha Kiaer et Pearson (Pearson 1892: 5), L. nigricans (Lindenberg in Gottsche et al. 1845: 316) Schiffner (1898: 293), L. onraedtii Vanden Berghen (1984: 452), L. subfusca (Nees 1830: 36) Schiffner (1897: 593) and L. utriculata Stephani (1912: 69).
References
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<a href="https://doi.org/10.5962/bhl.title.15221">https://doi.org/10.5962/bhl.title.15221</a>
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Sass-Gyarmati, A. (2001) Lopholejeunea leioptera Gyarmati (Lejeuneaceae, subfam. Ptychanthoideae), une nouvelle espèce récoltée dans la Réserve Spéciale de Manongarivo (Nord-Ouest de Madagascar). Candollea 56 (1): 79–83.
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Schiffner, V. (1897) Revision der Gattungen Omphalanthus und Lejeunea im Herbarium des Berliner Museums. Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 23 (5): 578–600.
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<a href="https://doi.org/10.11646/phytotaxa.65.1.5">https://doi.org/10.11646/phytotaxa.65.1.5</a>
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<a href="https://doi.org/10.3897/phytokeys.59.6261">https://doi.org/10.3897/phytokeys.59.6261</a>
Stephani, F. (1890) Hepaticae africanae novae in insulis Bourbon, Maurice et Madagascar lectae. Botanical Gazette 15 (11): 281–292.
<a href="https://doi.org/10.1086/326585">https://doi.org/10.1086/326585</a>
Stephani, F. (1893) Hepaticae. In: Renauld, F. & Cardot, J. (Eds.) Musci exotici novi vel minus cogniti. IV. Bulletin de la Société Royale de Botanique de Belgique, Comptes-rendus des Séances 32 (2): 8–40.
Stephani, F. (1896) Hepaticarum species novae IX. Hedwigia 35 (3): 73–140.
Stephani, F. (1912) Species hepaticarum 5. George & Cie, Genève & Bale, pp. 1–176.
<a href="https://doi.org/10.5962/bhl.title.95494">https://doi.org/10.5962/bhl.title.95494</a>
Taylor, T. (1846) New hepaticae. London Journal of Botany 5: 365–417.
Thiers, B.M. (1984) Studies in Lejeuneaceae, III. Lopholejeunea erugata, a new name for Ptychocoleus inermis. Brittonia 36 (2): 174–177.
<a href="https://doi.org/10.2307/2806626">https://doi.org/10.2307/2806626</a>
Vanden Berghen, C. (1984) Le genre Lopholejeunea (Spruce) Schiffn. (Lejeuneaceae, Hepaticae) en Afrique. Bulletin du Jardin Botanique National de Belgique 54: 393–464.
<a href="https://doi.org/10.2307/3667852">https://doi.org/10.2307/3667852</a>
Zhu, R.-L. & Gradstein, S.R. (2005) A monograph of the genus Lopholejeunea (Hepaticae) in Asia. Systematic Botany Monographs 74: 1–98.
<a href="https://doi.org/10.5962/bhl.title.15221">https://doi.org/10.5962/bhl.title.15221</a>
Marline, L., Andriamiarisoa, R.L., Bardat, J., Chuah-Petiot, M., Reeb, C., Strasberg, D., Wilding, N. & Ah-Peng, C. (2012) Checklist of the Bryophytes of Madagascar. Cryptogamie, Bryologie 33 (3): 199–255.
<a href="https://doi.org/10.7872/cryb.v33.iss3.2012.199">https://doi.org/10.7872/cryb.v33.iss3.2012.199</a>
Nees, C.G. (1830) Enumeratio plantarum cryptogamicarum Javae et insularum adiacentium. Fasciculus prior, hepaticas complectens, ab editore illustratas. Grass, Barth & Co., Breslau, 86 pp.
Pearson, W.H. (1892) Lejeuneae madagascariensis. Forhandlinger i Videnskabs-Selskabet i Kristiania 1892 (8): 3–9.
Sass-Gyarmati, A. (2001) Lopholejeunea leioptera Gyarmati (Lejeuneaceae, subfam. Ptychanthoideae), une nouvelle espèce récoltée dans la Réserve Spéciale de Manongarivo (Nord-Ouest de Madagascar). Candollea 56 (1): 79–83.
Schiffner, V. (1893) Hepaticae. In: Engler, A. & Prantl, K. (Eds.) Die Natürlichen Pflanzenfamilien, Teil. I, Abt. 3. Engelmann, Leipzig, pp. 1–144.
Schiffner, V. (1897) Revision der Gattungen Omphalanthus und Lejeunea im Herbarium des Berliner Museums. Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 23 (5): 578–600.
Schiffner, V. (1898) Conspectus hepaticarum archipelagi indici. Staatsdruckerei, Batavia, 382 pp.
Söderström, L., Hagborg, A. & von Konrat, M. (2012) Notes on Early Land Plants Today. Phytotaxa 65: 41–42.
<a href="https://doi.org/10.11646/phytotaxa.65.1.5">https://doi.org/10.11646/phytotaxa.65.1.5</a>
Söderström, L., Hagborg, A., von Konrat M., Bartholomew-Began, S., Bell, D., Briscoe, L., Brown, E., Cargill, D.C., Costa, D.P., Crandall-Stotler, B.J., Cooper, E.D., Dauphin, G., Engel, J.J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Ilkiu-Borges, A.L., Heinrichs, J., Hentschel, J., Katagiri, T., Konstantinova, N.A., Larraín, J., Long, D.G., Nebel, M., Pócs, T., Puche, F., Reiner-Drehwald, M.E.,Renner, M.A.M., Sass-Gyarmati, A., Schäfer-Verwimp, A., Segarra Moragues, J.G., Stotler, R.E., Sukkharak, P., Thiers, B.M., Uribe, J., Vá?a, J., Villarreal, J.C., Wigginton, M., Zhang, L. & Zhu, R.-L. (2016) World Checklist of Hornworts and Liverworts. PhytoKeys 59: 1–828.
<a href="https://doi.org/10.3897/phytokeys.59.6261">https://doi.org/10.3897/phytokeys.59.6261</a>
Stephani, F. (1890) Hepaticae africanae novae in insulis Bourbon, Maurice et Madagascar lectae. Botanical Gazette 15 (11): 281–292.
<a href="https://doi.org/10.1086/326585">https://doi.org/10.1086/326585</a>
Stephani, F. (1893) Hepaticae. In: Renauld, F. & Cardot, J. (Eds.) Musci exotici novi vel minus cogniti. IV. Bulletin de la Société Royale de Botanique de Belgique, Comptes-rendus des Séances 32 (2): 8–40.
Stephani, F. (1896) Hepaticarum species novae IX. Hedwigia 35 (3): 73–140.
Stephani, F. (1912) Species hepaticarum 5. George & Cie, Genève & Bale, pp. 1–176.
<a href="https://doi.org/10.5962/bhl.title.95494">https://doi.org/10.5962/bhl.title.95494</a>
Taylor, T. (1846) New hepaticae. London Journal of Botany 5: 365–417.
Thiers, B.M. (1984) Studies in Lejeuneaceae, III. Lopholejeunea erugata, a new name for Ptychocoleus inermis. Brittonia 36 (2): 174–177.
<a href="https://doi.org/10.2307/2806626">https://doi.org/10.2307/2806626</a>
Vanden Berghen, C. (1984) Le genre Lopholejeunea (Spruce) Schiffn. (Lejeuneaceae, Hepaticae) en Afrique. Bulletin du Jardin Botanique National de Belgique 54: 393–464.
<a href="https://doi.org/10.2307/3667852">https://doi.org/10.2307/3667852</a>
Zhu, R.-L. & Gradstein, S.R. (2005) A monograph of the genus Lopholejeunea (Hepaticae) in Asia. Systematic Botany Monographs 74: 1–98.