Abstract
As part of our ongoing studies on the Brazilian Stachytarpheta (Verbenaceae), we describe Stachytarpheta oblongifolia P.H.Cardoso & Salimena from Central Brazil, propose the new combination Stachytarpheta chascanoides (Moldenke) P.H.Cardoso, and reestablish Stachytarpheta mollis Moldenke. In addition, several taxa are here treated differently from previous classifications, namely Stachytarpheta atriflora S.Atkins is synonymized with Stachytarpheta glauca (Pohl) Walp.; Stachytarpheta caatingensis S.Atkins and Stachytarpheta villosa var. bahiensis Moldenke are synonymized with Stachytarpheta brasiliensis Moldenke; Stachytarpheta canescens var. elliptica Moldenke is synonymized with Stachytarpheta hirsutissima Link; Stachytarpheta gesnerioides var. simplex (Hayek) Moldenke is synonymized with Stachytarpheta gesnerioides Cham. var. gesnerioides; Stachytarpheta galactea S.Atkins is synonymized with Stachytarpheta coccinea Schauer; and, Stachytarpheta spathulata Moldenke subsp. spathulata and Stachytarpheta spathulata subsp. mogolensis S.Atkins are synonymized with Stachytarpheta chascanoides. Lectotypes are designated for 11 names, Stachytarpheta gesnerioides var. simplex, Stachytarpheta glauca, Stachytarpheta hatschbachii Moldenke, Stachytarpheta integrifolia (Pohl) Walp., Stachytarpheta longispicata (Pohl) S.Atkins subsp. longispicata, Stachytarpheta longispicata subsp. brevibracteata (Moldenke) S.Atkins, Stachytarpheta longispicata var. parvifolia (Moldenke) S.Atkins, Stachytarpheta lythrophylla Schauer, Stachytarpheta martiana Schauer, Stachytarpheta obovata Hayek, and Stachytarpheta speciosa Pohl ex Schauer.
References
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Bachman, S., Moat, J., Hill, A.W., de la Torre, J. & Scott, B. (2011) Supporting Red List threat assessments with GeoCAT: geospatial conservation assessment tool. ZooKeys 150: 117–126.
<a href="https://doi.org/10.3897/zookeys.150.2109">https://doi.org/10.3897/zookeys.150.2109</a>
Barbosa, A.G. (2008) As estratégias de conservação da biodiversidade na Chapada dos Veadeiros: conflitos e oportunidades. Universidade de Brasília, Brasília, 128 pp.
BFG - The Brazil Flora Group (2018) Brazilian Flora 2020: Innovation and collaboration to meet Target 1 of the Global Strategy for Plant Conservation (GSPC). Rodriguésia 69 (4): 1513–1527.
<a href="https://doi.org/10.1590/2175-7860201869402">https://doi.org/10.1590/2175-7860201869402</a>
Briquet, J. (1895) Verbenaceae. In: Engler, A. & Prantl, K. (Eds.) Die naturlichen Pflanzenfamilien 4, vol. 3a, pp. 132–182.
Cardoso, P.H., Alves, R.J.V., Menini Neto, L. & Salimena, F.R.G. (2019a) Stachytarpheta sobraliana (Verbenaceae), a new species from the Serra de São José, Minas Gerais, Brazil. Phytotaxa 415 (5): 286–292.
<a href="https://doi.org/10.11646/phytotaxa.415.5.5">https://doi.org/10.11646/phytotaxa.415.5.5</a>
Cardoso, P.H., Lima, L.V., Dittrich, V.A.O. & Salimena, F.R.G. (2019b) Typifications and additional taxonomic notes in Brazilian Stachytarpheta (Verbenaceae). Phytotaxa 411 (3): 223–229.
<a href="https://doi.org/10.11646/phytotaxa.411.3.6">https://doi.org/10.11646/phytotaxa.411.3.6</a>
Cardoso, P.H., Menini Neto, L. & Salimena, F.R.G. (2019c) A new species of Stachytarpheta (Verbenaceae) from an inselberg of Espírito Santo state, Brazil. Phytotaxa 400 (5): 273–278.
<a href="https://doi.org/10.11646/phytotaxa.400.5.3">https://doi.org/10.11646/phytotaxa.400.5.3</a>
Cardoso, P.H., Menini Neto, L. & Salimena, F.R.G. (2019d) Novos sinônimos em Stachytarpheta (Verbenaceae) do Brasil. Hoehnea 46: e082019.
<a href="https://doi.org/10.1590/2236-8906-08/2019">https://doi.org/10.1590/2236-8906-08/2019</a>
Cardoso, P.H., Menini Neto, L. & Salimena, F.R.G. (2019e) Stachytarpheta grandiflora, a new species of Verbenaceae from the Parque Nacional da Serra da Canastra, Minas Gerais, Brazil. Phytotaxa 413 (1): 61–66.
<a href="https://doi.org/10.11646/phytotaxa.413.1.7">https://doi.org/10.11646/phytotaxa.413.1.7</a>
Cardoso, P.H., Menini Neto, L., Bernacci, L.C. & Salimena, F.R.G. (2019f) Stachytarpheta kriegeriana (Verbenaceae), a new species from São Paulo State, Brazil. Phytotaxa 422 (3): 248–254.
<a href="https://doi.org/10.11646/phytotaxa.422.3.5">https://doi.org/10.11646/phytotaxa.422.3.5</a>
Cardoso, P.H., Menini Neto, L. & Salimena, F.R.G. (2020) Taxonomic notes on Stachytarpheta (Verbenaceae) from Brazil. Kew Bulletin 75: 17.
<a href="https://doi.org/10.1007/s12225-020-9875-5">https://doi.org/10.1007/s12225-020-9875-5</a>
Chamisso, L.K.A.von (1832) De plantis in expeditione Rommanzoffiana et in Herbariis Regiis, Verbenaceae. Linnaea 7: 213–272.
Fiedler, N.C., Merlo, D.A. & Medeiros, M.B.D. (2006) Ocorrência de incêndios florestais no Parque Nacional da Chapada dos Veadeiros, Goiás. Ciência Florestal 16: 153–161.
<a href="https://doi.org/10.5902/198050981896">https://doi.org/10.5902/198050981896</a>
Gonçalves, E.G. & Lorenzi, H. (2007) Morfologia vegetal: organografia e dicionário ilustrado de morfologia das plantas vasculares. Instituto Plantarum, Nova Odessa, 416 pp.
Harley, R.M., Antar, G.M. & Giulietti, A.M. (2019) Stachytarpheta atkinsiae, a new species of Verbenaceae endemic to the Chapada Diamantina, Bahia, Brazil. Phytotaxa 401 (2): 127–132.
<a href="https://doi.org/10.11646/phytotaxa.401.2.4">https://doi.org/10.11646/phytotaxa.401.2.4</a>
Hayek, A. (1907) Verbenaceae novae herbarii Vindobonensis. Repertorium Specierum Novarum Regni Vegetabilis 3: 273–274.
<a href="https://doi.org/10.1002/fedr.19070034402">https://doi.org/10.1002/fedr.19070034402</a>
Harris, J.G. & Harris, M.W. (2003) Plant identification terminology: an illustrated glossary. 2ed. Spring Lake Publ., Spring Lake, 216 pp.
IUCN Standards and Petitions Committee (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Available from: <a href="http://www.iucnredlist.org/documents/RedListGuidelines.pdf">http://www.iucnredlist.org/documents/RedListGuidelines.pdf</a> (accessed 30 June 2020)
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<a href="https://doi.org/10.5962/bhl.title.66">https://doi.org/10.5962/bhl.title.66</a>
Marx, H., O’leary, N., Yuan, Y., Lu-Irving, P., Tank, D., Múlgura, M.E. & Olmstead, R. (2010) A molecular phylogeny and classification of Verbenaceae. American journal of Botany 97: 1647–1663.
<a href="https://doi.org/10.3732/ajb.1000144">https://doi.org/10.3732/ajb.1000144</a>
McNeill, J. (2014) Holotype specimens and type citations: general issues. Taxon 63: 1112–1113.
<a href="https://doi.org/10.12705/635.7">https://doi.org/10.12705/635.7</a>
Melo, J.I.M., Medeiros-Alves, I., Sousa, R.T.M., Barbosa, L.M.M.A. & Andrade, W.M. (2010) Verbenaceae sensu lato em um trecho da ESEC Raso da Catarina, Bahia, Brasil. Revista Caatinga 23: 41–47.
Moldenke, H.N. (1934) Some New and Neglected Species and Varieties of the Verbenaceae. Repertorium Specierum Novarum Regni Vegetabilis 37: 14–20.
<a href="https://doi.org/10.1002/fedr.19340371402">https://doi.org/10.1002/fedr.19340371402</a>
Moldenke, H. N. (1947a) Notes on new and noteworthy plants I. Phytologia 2: 213–242.
Moldenke, H.N. (1947b) Notes on new or noteworthy plants III. Phytologia 2: 363–372.
Moldenke, H.N. (1949) Notes on new or noteworthy plants VII. Phytologia 3: 58–64.
Moldenke, H.N. (1974a) Notes on new and noteworthy plants LXXI. Phytologia 29: 75–78.
<a href="https://doi.org/10.5962/bhl.part.13103">https://doi.org/10.5962/bhl.part.13103</a>
Moldenke, H.N. (1974b) Notes on new and noteworthy plants LXVII. Phytologia 28: 192–295.
<a href="https://doi.org/10.5962/bhl.part.13103">https://doi.org/10.5962/bhl.part.13103</a>
Moldenke, H.N. (1980) Notes on New and Noteworthy Plants CXXXIII. Phytologia 45: 36–40.
<a href="https://doi.org/10.5962/bhl.part.28295">https://doi.org/10.5962/bhl.part.28295</a>
Moldenke, H.N. (1984) Notes on new and noteworthy plants CLXXIV. Phytologia 55: 232–234.
O’Leary, N., Calviño, C.I., Martínez, S., Lu-Irving, P., Olmstead, R.G. & Múlgura, M.E. (2012) Evolution of morphological traits in Verbenaceae. American Journal of Botany 99: 1778–1792.
<a href="https://doi.org/10.3732/ajb.1200123">https://doi.org/10.3732/ajb.1200123</a>
Patricio, R.L. (2009) Avaliação de métodos de revegetação de áreas degradadas utilizados na mineração de níquel em Niquelândia Goiás. Universidade Católica de Brasília, Brasília, 41pp.
Pohl, J.B.E. (1827) Plantarum Brasiliae Icones et Descriptiones 1. A.Strauss, Wien, pp. 1–152
Santos, J.S., Oliveira, L.D.S.D., van den Berg, C., & Sales, M.F. (2017) Lectotipificações de Stachytarpheta Vahl (Verbenaceae). Iheringia. Série Botânica 72: 441–444.
<a href="https://doi.org/10.21826/2446-8231201772315">https://doi.org/10.21826/2446-8231201772315</a>
Salimena, F.R.G., Ferreira, S.C., Cardoso, P.H. & Valério, V.I.R. (2016) Verbenaceae. In: Rizzo, J.A. (Ed.) Flora dos estados de Goiás e Tocantins. Coleção Rizzo. v. 47. Universidade Federal de Goiás, Goiânia, 157 pp.
Salimena, F.R.G. & Múlgura, M.E. (2015) Notas taxonômicas em Verbenaceae do Brasil. Rodriguésia 66: 191–197.
<a href="https://doi.org/10.1590/2175-7860201566110">https://doi.org/10.1590/2175-7860201566110</a>
Santos, J.S. (2015) Filogenia e Diversidade do gênero Stachytarpheta Vahl (Verbenaceae). Universidade Federal Rural de Pernambuco, Tese de Doutorado, 237 pp.
Schauer, J.C. (1847) Verbenaceae. In: De Candolle, A.P. (Ed.) Prodromus Systematis Naturalis Regni Vegetabilis 11. Sumptibus Sociorum Treuttel et Wu?rtz, Paris, pp. 522–700.
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Silva, M.S., Gurgel, H. Laques, A., Silveira, B.D. & Siqueira, R.V. (2018) 30 anos de dinâmica espaçotemporal (1984-2015) da região de influência do Parque Nacional da Chapada dos Veadeiros Goiás. Revista franco-brasilera de geografia, Confins: 35.
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<a href="https://doi.org/10.3897/zookeys.150.2109">https://doi.org/10.3897/zookeys.150.2109</a>
Barbosa, A.G. (2008) As estratégias de conservação da biodiversidade na Chapada dos Veadeiros: conflitos e oportunidades. Universidade de Brasília, Brasília, 128 pp.
BFG - The Brazil Flora Group (2018) Brazilian Flora 2020: Innovation and collaboration to meet Target 1 of the Global Strategy for Plant Conservation (GSPC). Rodriguésia 69 (4): 1513–1527.
<a href="https://doi.org/10.1590/2175-7860201869402">https://doi.org/10.1590/2175-7860201869402</a>
Briquet, J. (1895) Verbenaceae. In: Engler, A. & Prantl, K. (Eds.) Die naturlichen Pflanzenfamilien 4, vol. 3a, pp. 132–182.
Cardoso, P.H., Alves, R.J.V., Menini Neto, L. & Salimena, F.R.G. (2019a) Stachytarpheta sobraliana (Verbenaceae), a new species from the Serra de São José, Minas Gerais, Brazil. Phytotaxa 415 (5): 286–292.
<a href="https://doi.org/10.11646/phytotaxa.415.5.5">https://doi.org/10.11646/phytotaxa.415.5.5</a>
Cardoso, P.H., Lima, L.V., Dittrich, V.A.O. & Salimena, F.R.G. (2019b) Typifications and additional taxonomic notes in Brazilian Stachytarpheta (Verbenaceae). Phytotaxa 411 (3): 223–229.
<a href="https://doi.org/10.11646/phytotaxa.411.3.6">https://doi.org/10.11646/phytotaxa.411.3.6</a>
Cardoso, P.H., Menini Neto, L. & Salimena, F.R.G. (2019c) A new species of Stachytarpheta (Verbenaceae) from an inselberg of Espírito Santo state, Brazil. Phytotaxa 400 (5): 273–278.
<a href="https://doi.org/10.11646/phytotaxa.400.5.3">https://doi.org/10.11646/phytotaxa.400.5.3</a>
Cardoso, P.H., Menini Neto, L. & Salimena, F.R.G. (2019d) Novos sinônimos em Stachytarpheta (Verbenaceae) do Brasil. Hoehnea 46: e082019.
<a href="https://doi.org/10.1590/2236-8906-08/2019">https://doi.org/10.1590/2236-8906-08/2019</a>
Cardoso, P.H., Menini Neto, L. & Salimena, F.R.G. (2019e) Stachytarpheta grandiflora, a new species of Verbenaceae from the Parque Nacional da Serra da Canastra, Minas Gerais, Brazil. Phytotaxa 413 (1): 61–66.
<a href="https://doi.org/10.11646/phytotaxa.413.1.7">https://doi.org/10.11646/phytotaxa.413.1.7</a>
Cardoso, P.H., Menini Neto, L., Bernacci, L.C. & Salimena, F.R.G. (2019f) Stachytarpheta kriegeriana (Verbenaceae), a new species from São Paulo State, Brazil. Phytotaxa 422 (3): 248–254.
<a href="https://doi.org/10.11646/phytotaxa.422.3.5">https://doi.org/10.11646/phytotaxa.422.3.5</a>
Cardoso, P.H., Menini Neto, L. & Salimena, F.R.G. (2020) Taxonomic notes on Stachytarpheta (Verbenaceae) from Brazil. Kew Bulletin 75: 17.
<a href="https://doi.org/10.1007/s12225-020-9875-5">https://doi.org/10.1007/s12225-020-9875-5</a>
Chamisso, L.K.A.von (1832) De plantis in expeditione Rommanzoffiana et in Herbariis Regiis, Verbenaceae. Linnaea 7: 213–272.
Fiedler, N.C., Merlo, D.A. & Medeiros, M.B.D. (2006) Ocorrência de incêndios florestais no Parque Nacional da Chapada dos Veadeiros, Goiás. Ciência Florestal 16: 153–161.
<a href="https://doi.org/10.5902/198050981896">https://doi.org/10.5902/198050981896</a>
Gonçalves, E.G. & Lorenzi, H. (2007) Morfologia vegetal: organografia e dicionário ilustrado de morfologia das plantas vasculares. Instituto Plantarum, Nova Odessa, 416 pp.
Harley, R.M., Antar, G.M. & Giulietti, A.M. (2019) Stachytarpheta atkinsiae, a new species of Verbenaceae endemic to the Chapada Diamantina, Bahia, Brazil. Phytotaxa 401 (2): 127–132.
<a href="https://doi.org/10.11646/phytotaxa.401.2.4">https://doi.org/10.11646/phytotaxa.401.2.4</a>
Hayek, A. (1907) Verbenaceae novae herbarii Vindobonensis. Repertorium Specierum Novarum Regni Vegetabilis 3: 273–274.
<a href="https://doi.org/10.1002/fedr.19070034402">https://doi.org/10.1002/fedr.19070034402</a>
Harris, J.G. & Harris, M.W. (2003) Plant identification terminology: an illustrated glossary. 2ed. Spring Lake Publ., Spring Lake, 216 pp.
IUCN Standards and Petitions Committee (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Available from: <a href="http://www.iucnredlist.org/documents/RedListGuidelines.pdf">http://www.iucnredlist.org/documents/RedListGuidelines.pdf</a> (accessed 30 June 2020)
Link, J.H.F. (1821) Enumeratio Plantarum Horti Regii Berolinensis Altera. pt. 1. Paris, 458 pp.
<a href="https://doi.org/10.5962/bhl.title.66">https://doi.org/10.5962/bhl.title.66</a>
Marx, H., O’leary, N., Yuan, Y., Lu-Irving, P., Tank, D., Múlgura, M.E. & Olmstead, R. (2010) A molecular phylogeny and classification of Verbenaceae. American journal of Botany 97: 1647–1663.
<a href="https://doi.org/10.3732/ajb.1000144">https://doi.org/10.3732/ajb.1000144</a>
McNeill, J. (2014) Holotype specimens and type citations: general issues. Taxon 63: 1112–1113.
<a href="https://doi.org/10.12705/635.7">https://doi.org/10.12705/635.7</a>
Melo, J.I.M., Medeiros-Alves, I., Sousa, R.T.M., Barbosa, L.M.M.A. & Andrade, W.M. (2010) Verbenaceae sensu lato em um trecho da ESEC Raso da Catarina, Bahia, Brasil. Revista Caatinga 23: 41–47.
Moldenke, H.N. (1934) Some New and Neglected Species and Varieties of the Verbenaceae. Repertorium Specierum Novarum Regni Vegetabilis 37: 14–20.
<a href="https://doi.org/10.1002/fedr.19340371402">https://doi.org/10.1002/fedr.19340371402</a>
Moldenke, H. N. (1947a) Notes on new and noteworthy plants I. Phytologia 2: 213–242.
Moldenke, H.N. (1947b) Notes on new or noteworthy plants III. Phytologia 2: 363–372.
Moldenke, H.N. (1949) Notes on new or noteworthy plants VII. Phytologia 3: 58–64.
Moldenke, H.N. (1974a) Notes on new and noteworthy plants LXXI. Phytologia 29: 75–78.
<a href="https://doi.org/10.5962/bhl.part.13103">https://doi.org/10.5962/bhl.part.13103</a>
Moldenke, H.N. (1974b) Notes on new and noteworthy plants LXVII. Phytologia 28: 192–295.
<a href="https://doi.org/10.5962/bhl.part.13103">https://doi.org/10.5962/bhl.part.13103</a>
Moldenke, H.N. (1980) Notes on New and Noteworthy Plants CXXXIII. Phytologia 45: 36–40.
<a href="https://doi.org/10.5962/bhl.part.28295">https://doi.org/10.5962/bhl.part.28295</a>
Moldenke, H.N. (1984) Notes on new and noteworthy plants CLXXIV. Phytologia 55: 232–234.
O’Leary, N., Calviño, C.I., Martínez, S., Lu-Irving, P., Olmstead, R.G. & Múlgura, M.E. (2012) Evolution of morphological traits in Verbenaceae. American Journal of Botany 99: 1778–1792.
<a href="https://doi.org/10.3732/ajb.1200123">https://doi.org/10.3732/ajb.1200123</a>
Patricio, R.L. (2009) Avaliação de métodos de revegetação de áreas degradadas utilizados na mineração de níquel em Niquelândia Goiás. Universidade Católica de Brasília, Brasília, 41pp.
Pohl, J.B.E. (1827) Plantarum Brasiliae Icones et Descriptiones 1. A.Strauss, Wien, pp. 1–152
Santos, J.S., Oliveira, L.D.S.D., van den Berg, C., & Sales, M.F. (2017) Lectotipificações de Stachytarpheta Vahl (Verbenaceae). Iheringia. Série Botânica 72: 441–444.
<a href="https://doi.org/10.21826/2446-8231201772315">https://doi.org/10.21826/2446-8231201772315</a>
Salimena, F.R.G., Ferreira, S.C., Cardoso, P.H. & Valério, V.I.R. (2016) Verbenaceae. In: Rizzo, J.A. (Ed.) Flora dos estados de Goiás e Tocantins. Coleção Rizzo. v. 47. Universidade Federal de Goiás, Goiânia, 157 pp.
Salimena, F.R.G. & Múlgura, M.E. (2015) Notas taxonômicas em Verbenaceae do Brasil. Rodriguésia 66: 191–197.
<a href="https://doi.org/10.1590/2175-7860201566110">https://doi.org/10.1590/2175-7860201566110</a>
Santos, J.S. (2015) Filogenia e Diversidade do gênero Stachytarpheta Vahl (Verbenaceae). Universidade Federal Rural de Pernambuco, Tese de Doutorado, 237 pp.
Schauer, J.C. (1847) Verbenaceae. In: De Candolle, A.P. (Ed.) Prodromus Systematis Naturalis Regni Vegetabilis 11. Sumptibus Sociorum Treuttel et Wu?rtz, Paris, pp. 522–700.
Schauer, J.C. (1851) Verbenaceae. In: Martius, C.F.P. von & Eichler, A.W. (Eds.) Flora brasiliensis. F. Fleischer, Leipzig, pp. 169?308.
Schrader, H.A. (1821) Illustrationes super plantis quibusdam novis et minus cognitis, a Principe Serenissimo Maximiliano Neowidensi in Brasilia observatis: “Plantarum rariorum a Principe Serenissimo Maximiliano Neowidensi in itinere per Brasiliam observatarum Fascicul. I”. Göttingische Gelehrte Anzeigen unter der Aufsicht der Königl. Gesellschaft der Wissenschaften: 705–720.
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<a href="https://doi.org/10.4000/confins.14851">https://doi.org/10.4000/confins.14851</a>
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