ZUO, ZHENG-YU, School of Life Sciences, Yunnan University, Kunming, Yunnan, 650500, China; Germplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan, 650201, China; Kunming College of Life Science, University of Chinese Academy of Sciences, Kunming, Yunnan, 650201, China, China