ZHAO, YONGCHANG, Biotechnology and Germplasm Resources Institute, Yunnan Academy of Agricultural Sciences; Kunming, 650223, Yunnan, China. Yunnan Provincial Key Lab of Agricultural Biotechnology; Kunming, 650223, Yunnan, China. Key Lab of Southwestern Crop Gene Resources and Germplasm Innovation, Ministry of Agriculture, Kunming, 650223, Yunnan, China., China