JU, WEN-BIN, Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration Biodiversity Conservation, Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China; Key Laboratory of Bio-Resources and Eco-Environment of Ministry of Education, College of Life Sciences, Sichuan University, 610065, Chengdu, Sichuan, People’s Republic of China; University of Chinese Academy of Sciences, Beijing 100049, China, China