EL MOKNI, RIDHA, University of Monastir, Department of Pharmaceutical Sciences “A”, Laboratory of Botany, Cryptogamy and Plant Biology, Faculty of Pharmacy of Monastir, Avenue Avicenna, 5000-Monastir, Tunisia. University of Jendouba, Department of Silvo-Pastoral Resources, Laboratory of Silvo-Pastoral Resources, Silvo-Pastoral Institute of Tabarka, BP. 345, 8110-Tabarka, Tunisia. University of Carthage, IRESA, Laboratory of Forest Ecology, National Research Institute of Rural Engineering, Water and Forests, Ariana 2080, Tunisia., Tunisia