EL MOKNI, RIDHA, Department of Pharmaceutical Sciences ‘A’, Laboratory of Botany, Cryptogamy and Plant Biology, Faculty of Pharmacy of Monastir, Avenue Avicenna, Monastir-5000, University of Monastir, Tunisia; Department of Forestry, Laboratory of Forest Ecology, National Research Institute of Rural Engineering, Water and Forests, Ariana-2080, IRESA, University of Carthage, Tunisia, Tunisia