EL MADIHI, MOHAMMED, Research Centre of Plant and Microbial Biotechnologies, Biodiversity and Environment, Faculty of Sciences, Mohammed V University in Rabat, 4 avenue Ibn Battouta, B.P. 1014 RP, Rabat, Morocco; Tour du Valat, Research Institute for the conservation of Mediterranean wetlands, Le Sambuc, 13200, Arles, France, Morocco