LEI, LEI, Key Laboratory for Plant Diversity and Biogeography of East Asia, Yunnan Key Laboratory of Fungal Diversity and Green Development, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China. Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand. School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand., China