YAN, JIAWEI, State Key Laboratory of Plant Diversity and Specialty Crops/Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, 510650, Guangzhou, China, South China National Botanical Garden, 510650, Guangzhou, China, Department of Horticulture and Landscape Architecture, Zhongkai University of Agriculture and Engineering, 510225, Guangzhou, China, China