WANASINGHE, DHANUSHKA N., Center of Excellence, Mae Fah Luang University, Chiang Rai, Thailand Key Laboratory for Plant Biodiversity and Biogeography of East Asia (KLPB), Kunming Institute of Botany, Chinese Academy of Science, Kunming 650201, Yunnan China World Agro Forestry Centre, East and Central Asia, 132 Lanhei Road, Kunming 650201, Yunnan China School of Science, Mae Fah Luang University, Chiang Rai, 57100, Thailand, Thailand