QUANG, BUI HONG, Department of Botany, Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoang Quoc Viet, Cau Giay, 10000 Hanoi, Vietnam. Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoang Quoc Viet, Cau Giay, 10000 Hanoi, Vietnam., Viet Nam