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Issue: Vol. 765 No. 3: 2 Jul. 2026
Type: Article
Published: 2026-07-02
DOI: 10.11646/phytotaxa.765.3.3
Page range: 246-256
Abstract views: 0
PDF downloaded: 0

A new clavarioid wood-inhabiting fungus, Artomyces tsugae (Russulales Basidiomycota), from Guangxi, southern China

School of Ecology and Nature Conservation; Beijing Forestry University; Beijing 100083; China
School of Ecology and Nature Conservation; Beijing Forestry University; Beijing 100083; China; Harbin Academy of Agricultural Sciences; Harbin 150029; Heilongjiang; China
School of Ecology and Nature Conservation; Beijing Forestry University; Beijing 100083; China; Harbin Academy of Agricultural Sciences; Harbin 150029; Heilongjiang; China
Yunnan Key Laboratory for Fungal Diversity and Green Development; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650201; China
School of Ecology and Nature Conservation; Beijing Forestry University; Beijing 100083; China
Fungi Agaricomycetes clavarioid fungi subtropical China wood-inhabiting fungi

Abstract

A new lignicolous species, Artomyces tsugae, is described from the high mountains of Guangxi, subtropical China, based on morphological examination and phylogenetic analysis. The new species is characterized by annual, stipitate, well-branched and snow white basidomata when fresh, a monomitic hyphal system with clamp connections on generative hyphae, the presence of gloeoplerous hyphae and gloeocystidia, ellipsoid, hyaline, thin-walled, verrucose, amyloid basidiospores measuring 4.6–5.2 × 3.1–3.6 µm, and growing on rotten wood of Tsuga tchekiangensis. Phylogenetic analysis based on ITS sequences indicated that it was closely related to A. brunneoalbus, A. yunnanensis, and five other species. A description, illustrations, and phylogenetic analysis results of the new species are provided. In addition, the new species is compared with closely related and similar species.

References

  1. Cai, Q., He, S., Li, G.M., Fan, X.P., Li, Y.C. & Yang, Z.L. (2025) The genus Artomyces (Auriscalpiaceae, Basidiomycota) from the Gaoligong Mountains and adjacent areas, southwestern China. Mycologia 117: 516–531. https://doi.org/10.1080/00275514.2025.2476936
  2. Dai, Y.C. & Penttilä, R. (2006) Polypore diversity of Fenglin Nature Reserve, northeastern China. Annales Botanici Fennici 43: 81–96.
  3. Dai, Y.C. (2000) A checklist of polypores from Northeast China. Karstenia 40: 23–29. https://doi.org/10.29203/ka.2000.347
  4. Dai, Y.C., Wang, Z., Binder, M. & Hibbett, D.S. (2006) Phylogeny and a new species of Sparassis (Polyporales, Basidiomycota): evidence from mitochondrial atp6, nuclear rDNA and rpb2 genes. Mycologia 98: 584–592. https://doi.org/10.1080/15572536.2006.11832661
  5. Dai, Y.C., Yang, Z.L., Cui, B.K., Wu, G., Yuan, H.S., Zhou, L.W., He, S.H., Ge, Z.W., Wu, F., Wei, Y.L., Yuan, Y. & Si, J. (2021) Diversity and systematics of the important macrofungi in Chinese forests. Mycosystema 40 (4): 770–805. https://doi.org/10.13346/j.mycosystema.210036
  6. Dai, Y.C., Yu, C.J. & Wang, H.C. (2007) Polypores from eastern Xizang (Tibet), western China. Annales Botanici Fennici 44: 135–145.
  7. Dong, J.H., Li, Q., Yuan, Q., Luo, Y.X., Zhang, X.C., Dai, Y.F., Zhou, Q., Liu, X.F., Deng, Y.L., Zhou, H.M., Muhammad, A. & Zhao, C.L. (2024) Species diversity, taxonomy, molecular systematics and divergence time of wood-inhabiting fungi in Yunnan-Guizhou Plateau, Asia. Mycosphere 15 (1): 1110–1293. https://doi.org/10.5943/mycosphere/15/1/10
  8. Gilbertson, R.L. & Ryvarden, L. (1986) North American polypores 2. Megasporoporia - Wrightoporia. Fungiflora, Oslo.
  9. Gilbertson, R.L. & Ryvarden, L. (1987) North American polypores 2. Megasporoporia - Wrightoporia. Fungiflora, Oslo, pp. 434–885.
  10. Haelewaters, D., Dirks, A.C., Kappler, L.A., Mitchell, J.K., Quijada, L., Vandegrift, R., Buyck, B. & Pfister, D.H. (2018) A preliminary checklist of fungi at the Boston Harbor Islands. Northeastern Naturalist 25: 45–76. https://doi.org/10.1656/045.025.s904
  11. Hall, T.A. (1999) BioEdit: a user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. Nucleic Acids Symposium Series 41: 95–98.
  12. He, M.Q., Zhao, R.L., Hyde, K.D., Begerow, D., Kemler, M., Yurkov, A., McKenzie, E.H.C., Raspé, O., Kakishima, M., Sánchez-Ramírez, S., Vellinga, E.C., Halling, R., Papp, V., Zmitrovich, I.V., Buyck, B., Ertz, D., Wijayawardene, N.N., Cui, B.K., Schoutteten, N., Liu, X.Z., Li, T.H., Yao, Y.J., Zhu, X.Y., Liu, A.Q., Li, G.J., Zhang, M.Z., Ling, Z.L., Cao, B., Antonín, V., Boekhout, T., da Silva, B.D.B., De Crop, E., Decock, C., Dima, B., Dutta, A.K., Fell, J.W., Geml, J., Ghobad-Nejhad, M., Giachini, A.J., Gibertoni, T.B., Gorjón, S.P., Haelewaters, D., He, S.H., Hodkinson, B.P., Horak, E., Hoshino, T., Justo, A., Lim, Y.W., Menolli Jr., N., Mešić, A., Moncalvo, J.M., Mueller, G.M., Nagy, L.G., Nilsson, R.H., Noordeloos, M., Nuytinck, J., Orihara, T., Ratchadawan, C., Rajchenberg, M., Silva-Filho, A.G.S., Sulzbacher, M.A., Tkalčec, Z., Valenzuela, R., Verbeken, A., Vizzini, A., Wartchow, F., Wei, T.Z., Weiß, M., Zhao, C.L. & Kirk, P.M. (2019) Notes, outline and divergence times of Basidiomycota. Fungal Diversity 99: 105–367. https://doi.org/10.1007/s13225-019-00435-4
  13. Hyde, K.D., Noorabadi, M.T., Thiyagaraja, V., He, M.Q., Johnston, P.R., Wijesinghe, S.N., Armand, A., Biketova, A.Y., Chethana, K.W.T., Erdoğdu, M., Ge, Z.W., Groenewald, J.Z., Hongsanan, S., Kušan, I., Leontyev, D.V., Li, D.W., Lin, C.G., Liu, N.G., Maharachchikumbura, S.S.N., Matočec, N., May, T.W., McKenzie, E.H.C., Mešić, A., Perera, R.H., Phukhamsakda, C., Piątek, M., Samarakoon, M.C., Selcuk, F., Senanayake, I.C., Tanney, J.B., Tian, Q., Vizzini, A., Wanasinghe, D.N., Wannasawang, N., Wijayawardene, N., Zhao, R.L., Abdel-Wahab, M.A., Abdollahzadeh, J., Abeywickrama, P.D., Abhinav, Absalan, S., Acharya, K., Afshari, N., Afshan,. NS, Afzalinia, S., Ahmadpour, S.A., Akulov, O., Alizadeh, A., Alizadeh, M., Al-Sadi, A.M., Alves, A., Alves, V.C.S., Alves-Silva, G., Antonín, V., Aouali, S., Aptroot, A., Apurillo, C.C.S., Arias, R.M., Asgari, B., Asghari, R., Assis, D.M.A., Assyov, B., Atienza, V., Aumentado, H.D.R., Avasthi, S., Azevedo, E., Bakhshi, M., Bao, D.F., Baral, H.O., Barata, M., Barbosa, K.D., Barbosa, R.N., Barbosa, F.R., Baroncelli, R., Barreto, G.G., Baschien, C., Bennett, R.M., Bera, I., Bezerra, J.D.P., Bhunjun, C.S., Bianchinotti, M.V., Błaszkowski, J., Boekhout, T., Bonito, G.M., Boonmee, S., Boonyuen, N., Bortnikov, F.M., Bregant, C., Bundhun, D., Burgaud, G., Buyck, B., Caeiro, M.F., Cabarroi-Hernández, M., Cai, M.F., Cai, L., Calabon, M.S., Calaça, F.J.S., Callalli, M., Câmara, M.P.S., Cano-Lira, J., Cao, B., Carlavilla, J.R., Carvalho, A., Carvalho, T.G., Castañeda-Ruiz, R.F., Catania, M.D.V., Cazabonne, J., Cedeño-Sanchez, M., Chaharmiri-Dokhaharani, S., Chaiwan, N., Chakraborty, N., Cheewankoon, R., Chen, C., Chen, J., Chen, Q., Chen, Y.P., Chinaglia, S., Coelho-Nascimento, C.C., Coleine, C., CostaRezende, D.H., Cortés-Pérez, A., Crouch, J.A., Crous, P.W., Cruz, R.H.S.F., Czachura, P., Damm, U., Darmostuk, V., Daroodi, Z., Das, K., Davoodian, N., Davydov, E.A., da Silva, G.A., da Silva, I.R., da Silva, R.M.F., da Silva, S.A.C., Dai, D.Q., Dai, Y.C., de Groot, M.D., De Kesel, A., De Lange, R., de Medeiros, E.V., de Souza, C.F.A., de Souza, F.A., dela Cruz, T.E.E., Decock, C., Delgado, G., Denchev, C.M., Denchev, T.T., Deng, Y.L., Dentinger, B.T.M., Devadatha, B., Dianese, J.C., Dima, B., Doilom, M., Dissanayake, A.J., Dissanayake, D.M.L.S., Dissanayake, L.S., Diniz, A.G., Dolatabadi, S., Dong, J.H., Dong, W., Dong, Z.Y., Drechsler-Santos, E.R., Druzhinina, I.S., Du, T.Y., Dubey, M.K., Dutta, A.K., Elliott, T.F., Elshahed, M.S., Egidi, E., Eisvand, P., Fan, L., Fan, X., Fan, X.L., Fedosova, A.G., Ferro, L.O., Fiuza, P.O., Flakus, A.W., Fonseca, E.O., Fryar, S.C., Gabaldón, T., Gajanayake, A.J., Gannibal, P.B., Gao, F., GarcíaSánchez, D., García-Sandoval, R., Garrido-Benavent, I., Garzoli, L., Gasca-Pineda, J., Gautam, A.K., Gené, J., Ghobad-Nejhad, M., Ghosh, A., Giachini, A.J., Gibertoni, T.B., Gentekaki, E., Gmoshinskiy, V.I., GóesNeto, A., Gomdola, D., Gorjón, S.P., Goto, B.T., Granados-Montero, M.M., Griffith, G.W., Groenewald, M., Grossart, H.-P., Gu, Z.R., Gueidan, C., Gunarathne, A., Gunaseelan, S., Guo, S.L., Gusmão, L.F.P., Gutierrez, A.C., Guzmán-Dávalos, L., Haelewaters, D., Haituk, H., Halling, R.E., He, S.C., Heredia, G., HernándezRestrepo, M., Hosoya, T., Hoog, S.D., Horak, E., Hou, C.L., Houbraken, J., Htet, Z.H., Huang, S.K., Huang, W.J., Hurdeal, V.G., Hustad, V.P., Inácio, C.A., Janik, P., Jayalal, R.G.U., Jayasiri, S.C., Jayawardena, R.S., Jeewon, R., Jerônimo, G.H., Jin, J., Jones, E.B.G., Joshi, Y., Jurjević, Ž., Justo, A., Kakishima, M., Kaliyaperumal, M., Kang, G.P., Kang, J.C., Karimi, O., Karunarathna, S.C., Karpov, S.A., Kezo, K., Khalid, A.N., Khan, M.K., Khuna, S., Khyaju, S., Kirchmair, M., Klawonn, I., Kraisitudomsook, N., Kukwa, M., Kularathnage, N.D., Kumar, S., Lachance, M.A., Lado, C., Latha, K.P.D., Lee, H.B., Leonardi, M., Lestari, A.S., Li, C., Li, H., Li, J., Li, Q., Li, Y., Li, Y.C., Li, Y.X., Liao, C.F., Lima, J.L.R., Lima, J.M.S., Lima, N.B., Lin, L., Linaldeddu, B.T., Linn, M.M., Liu, F., Liu, J.K., Liu, J.W., Liu, S., Liu, S.L., Liu, X.F., Liu, X.Y., Longcore, J.E., Luangharn, T., Luangsaard, J.J., Lu, L., Lu, Y.Z., Lumbsch, H.T., Luo, L., Luo, M., Luo, Z.L., Ma, J., Madagammana, A.D., Madhushan, A., Madrid, H., Magurno, F., Magyar, D., Mahadevakumar, S., Malosso, E., Malysh, J.M., Mamarabadi, M., Manawasinghe, I.S., Manfrino, R.G., Manimohan, P., Mao, N., Mapook, A., Marchese, P., Marasinghe, D.S., Mardones, M., Marin-Felix, Y., Masigol, H., Mehrabi, M., MehrabiKoushki, M., Meiras-Ottoni, A., de Melo, R.F.R., Mendes-Alvarenga, R.L., Mendieta, S., Meng, Q.F., Menkis, A., Menolli, J.N., Mikšík, M., Miller, S.L., Moncada, B., Moncalvo, J.M., Monteiro, J.S., Monteiro, M., Mora-Montes, H.M., Moroz, E.L., Moura, J.C., Muhammad, U., Mukhopadhyay, S., Nagy, G.L., Najamul, S.A., Najafiniya, M., Nanayakkara, C.M., Naseer, A., Nascimento, E.C.R., Nascimento, S.S., Neuhauser, S., Neves, M.A., Niazi, A.R., Nie, Yong., Nilsson, R.H., Nogueira, P.T.S., Novozhilov, Y.K., Noordeloos, M., Norphanphoun, C., Nuñez, O.N., O’Donnell, R.P., Oehl, F., Oliveira, J.A., Oliveira, J.I., Oliveira, N.V.L., Oliveira, P.H.F., Orihara, T., Oset, M., Pang, K.L., Papp, V., Pathirana, L.S., Peintner, U., Pem, D., Pereira, O.L., Pérez-Moreno, J., Pérez-Ortega, S., Péter, G., Pires-Zottarelli, C.L.A., Phonemany, M., Phongeun, S., Pošta, A., Prazeres, J.F.S.A., Quan, Y., Quandt, C.A., Queiroz, M.B., Radek, R., Rahnama, K., Raj, K.N.A., Rajeshkumar, K.C., Rajwar, S., Ralaiveloarisoa, A.B., Rämä, T., Ramírez-Cruz, V., Rambold, G., Rathnayaka, A.R., Raza, M., Ren, G.C., Rinaldi, A.C., Rivas-Ferreiro, M., Robledo, G.L., Ronikier, A., Rossi, W., Rusevska, K., Ryberg, M., Safi, A., Salimi, F., Salvador-Montoya, C.A., Samant, B., Samaradiwakara, N.P., Sánchez-Castro, I., Sandoval-Denis, M., Santiago, A.L.C.M.A., Santos, A.C.D.S., Santos, L.A., Sarma, V.V., Sarwar, S., Savchenko, A., Savchenko, K., Saxena, R.K., Schoutteten, N., Selbmann, L., Ševčíková, H., Sharma, A., Shen, H.W., Shen, Y.M., Shu, Y.X., Silva, H.F., Silva-Filho, A.G.S., Silva, V.S.H., Simmons, D.R., Singh, R., Sir, E.B., Sohrabi, M., Souza, F.A., Souza-Motta, C.M., Sriindrasutdhi, V., Sruthi, O.P., Stadler, M., Stemler, J., Stephenson, S.L., Stoyneva-Gaertner, M.P., Strassert, J.F.H., Stryjak-Bogacka, M., Su, H., Sun, Y.R., Svantesson, S., Sysouphanthong, P., Takamatsu, S., Tan, T.H., Tanaka, K., Tang, C., Tang, X., Taylor, J.E., Taylor, P.W.J, Tennakoon, D.S., Thakshila, S.A.D., Thambugala, K.M., Thamodini, G.K., Thilanga, D., Thines, M., Tiago, P.V., Tian, X.G., Tian, W.H., Tibpromma, S., Tkalčec, Z., Tokarev, Y.S., Tomšovský, M., Torruella, G., Tsurykau, A., Udayanga, D., Ulukapı, M., Untereiner, W.A., Usman, M., Uzunov, B.A., Vadthanarat, S., Valenzuela, R., Van den Wyngaert, S., Van Vooren, N., Velez, P., Verma, R.K., Vieira, L.C., Vieira, W.A.S., Vinzelj, J.M., Tang, A.M.C., Walker, A., Walker, A.K., Wang, Q.M., Wang, Y., Wang, X.Y., Wang, Z.Y., Wannathes, N., Wartchow, F., Weerakoon, G., Wei, D.P., Wei, X., White, J.F., Wijesundara, D.S.A., Wisitrassameewong, K., Worobiec, G., Wu, H.X., Wu, N., Xiong, Y.R., Xu, B., Xu, J.P., Xu, R., Xu, R.F., Xu, R.J., Yadav, S., Yakovchenko, L.S., Yang, H.D., Yang, X., Yang, Y.H., Yang, Y., Yang, Y.Y., Yoshioka, R., Youssef, N.H., Yu, F.M., Yu, Z.F., Yuan, L.L., Yuan, Q., Zabin, D.A., Zamora, J.C., Zapata, C.V., Zare, R., Zeng, M., Zeng, X.Y., Zhang, J.F., Zhang, J.Y., Zhang, S., Zhang, X.C., Zhao, C.L., Zhao, H., Zhao, Q., Zhao, H., Zhao, H.J., Zhou, H.M., Zhu, X.Y., Zmitrovich, I.V., Zucconi, L. & Zvyagina, E. (2024) The 2024 Outline of Fungi and fungus-like taxa. Mycosphere 15 (1): 5146–6239. https://doi.org/10.5943/mycosphere/15/1/25
  14. Katoh, K. & Standley, D.M. (2013) MAFFT multiple sequence alignment software version 7: improvements in performance and usability. Molecular Biology and Evolution 30: 772–780. https://doi.org/10.1093/molbev/mst010
  15. Kim, C.S., Jo, J.W., Kwag, Y.N., Sung, G.H., Lee, S.G., Kim, S.Y., Shin, C.H. & Han, S.K. (2015) Mushroom flora of Ulleung-gun and a newly recorded bovista species in the Republic of Korea. Mycobiology 43: 239–257. https://doi.org/10.5941/Myco.2015.43.3.239
  16. Kneal, R. & Smith, M.E. (2015) Artomyces nothofagi sp. nov. a clavarioid fungus from a Chilean Nothofagus forest. Mycotaxon 130: 653–660. https://doi.org/10.5248/130.653
  17. Lickey, E.B., Hughes, K.W. & Petersen, R.H. (2002) Biogeographical patterns in Artomyces pyxidatus. Mycologia 94: 461–471. https://doi.org/10.1080/15572536.2003.11833211
  18. Lickey, E.B., Hughes, K.W. & Petersen, R.H. (2003a) Phylogenetic and taxonomic studies in Artomyces and Clavicorona (Homobasidiomycetes: Auriscalpiaceae). Sydowia 55: 181–254.
  19. Lickey, E.B., Hughes, K.W. & Petersen, R.H. (2003b) Variability and phylogenetic incongruence of an SSU nrDNA Group I intron in Artomyces, Auriscalpium, and Lentinellus (Auriscalpiaceae: Homobasidiomycetes). Molecular Biology and Evolution 20: 1909–1916. https://doi.org/10.1093/molbev/msg210
  20. Liu, X.Y., Liu, S.L., Wei, H.W., Wang, X.W., Yu, J., Shen, S. & Zhou, L.W. (2023) Preliminary species diversity and community phylogenetics of wood-inhabiting basidiomycetous fungi in the Dabie Mountains, Central China reveal unexpected richness. IMA Fungus 14: 23. https://doi.org/10.1186/s43008-023-00130-9
  21. Liu, L.L., Ren, Y.L., Habib, K., Lu, C.T., Wu, Y.P., Long, S.H., Lin, Y., Zhang, X., Kang, Y.Q., Wijayawardene, N.N., Wang, F., Elgorban, A.M., Al-Rejaie, S., Samarakoon, M.C., Shen, X.C. & Li, Q.R. (2025) New taxa of Xylariales from Karst Ecosystems in Southwestern China. Mycosphere 16 (1): 1–78. https://doi.org/10.5943/mycosphere/16/1/1
  22. Nylander, J.A.A. (2004) MrModeltest v2. Program distributed by the author, Evolutionary Biology Centre, Uppsala University.
  23. Núñez, M. & Ryvarden, L. (2000) East Asian polypores 1. Ganodermataceae and Hymenochaetaceae. Synopsis Fungorum 13: 1–168.
  24. Núñez, M. & Ryvarden, L. (2001) East Asian polypores 2. Polyporaceae s. lato. Synopsis Fungorum 14: 170–522.
  25. Petersen, J.H. (1996) The Danish mycological society´s colour-chart. Foreningen til Svampekundskabens Fremme, Greve.
  26. Rathnayaka, A.R., Tennakoon, D.S., Jones, G.E., Wanasinghe, D.N., Bhat, D.J., Priyashantha, A.H., Stephenson, S.L., Tibpromma, S. & Karunarathna, S.C. (2025) Significance of precise documentation of hosts and geospatial data of fungal collections, with an emphasis on plant-associated fungi. New Zealand Journal of Botany 63 (2–3): 462–489. https://doi.org/10.1080/0028825x.2024.2381734
  27. Ronquist, F., Teslenko, M., Van Der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. Systematic Biology 61 (3): 539–542. https://doi.org/10.1093/sysbio/sys029
  28. Ryvarden, L. & Melo, I. (2014) Poroid fungi of Europe. Synopsis Fungorum 31. Fungiflora, 455 pp.
  29. Song, C.G., Xu, T.M., Xu, Y.H., Wang, D., Zeng, L., Fan, X.P., Sun, Y.F. & Cui, B.K. (2025) Systematic revision, molecular phylogeny and divergence times of Thelephorales (Basidiomycota). Mycosphere 16 (1): 296–422. https://doi.org/10.5943/mycosphere/16/1/5
  30. Spirin, V., Malysheva, V., Viner, I., Alvarenga, R.L.M., Grebenc, T., Gruhn, G., Savchenko, A., Grootmyers, D., Ryvarden, L., Vlasák, J., Larrson, K. & Nilsson, R.H. (2025) Additions to the taxonomy of the Auriculariales (Basidiomycota) with pedunculate basidia. MycoKeys 120: 339–392. https://doi.org/10.3897/mycokeys.120.155492
  31. Stamatakis, A. (2014) RAxML version 8: A tool for phylogenetic analysis and post‐analysis of large phylogenies. Bioinformatics 30: 1312–1313. https://doi.org/10.1093/bioinformatics/btu033
  32. Wang, C.G., Dai, Y.C., Kout, J., Gates, G.M., Liu, H.G., Yuan, Y. & Vlasák, J. (2024) Multi-gene phylogeny and taxonomy of Physisporinus (Polyporales, Basidiomycota). Mycosphere 15: 1455–1521. https://doi.org/10.5943/mycosphere/15/1/12
  33. White, T.J., Bruns, T., Lee, S. & Taylor, J. (1990) Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. In: Innis, M.A., Gelfand, D.H., Sninsky, J.J. & White, T.J. (Eds.) PCR protocols, a guide to methods and applications. Academic Press, New York, pp. 315–322. https://doi.org/10.1016/b978-0-12-372180-8.50042-1
  34. Wu, F., Man, X.W., Tohtirjap, A. & Dai, Y.C. (2022) A comparison of polypore funga and species composition in forest ecosystems of China, North America, and Europe. Forest Ecosystems 9: 100051. https://doi.org/10.1016/j.fecs.2022.100051
  35. Yang, Y., Xu, Y., Wang, L., Jiang, Q.Q., Su, J.Q., Li, R., Zhou, H.M. & Zhao, C.L. (2025) Multigene phylogeny of seven wood-inhabiting fungal orders in Basidiomycota, and proposal of a new genus and thirteen new species. Mycosphere 16 (1): 245–295. https://doi.org/10.5943/mycosphere/16/1/4
  36. Zhao, H., Cui, Y.J., Guan, Q.X., Wang, K., Zhuang, L., Zeng, G.Y., Wei, Y.L., Wu, F. & Yuan, H.S. (2025) Global species diversity and distribution patterns within the order Hymenochaetales (Agaricomycetes, Basidiomycota). Mycosphere 16 (1): 3257–3280. https://doi.org/10.5943/mycosphere/16/1/24
  37. Zhao, H., Wu, Y.D., Yang, Z.R., Liu, H.G., Wu, F., Dai, Y.C. & Yuan, Y. (2024) Polypore funga and species diversity in tropical forest ecosystems of Africa, America and Asia, and a comparison with temperate and boreal regions of the Northern Hemisphere. Forest Ecosystems 11: 100200. https://doi.org/10.1016/j.fecs.2024.100200
  38. Zhou, L.W., Nakasone, K.K., Burdsall, J.H., Ginns, J., Vlasák, J., Miettinen, O., Spirin, V., Niemelä, T., Yuan, H.S., He, S.H., Cui, B.K., Xing, J.H. & Dai, Y.C. (2016) Polypore diversity in North America with an annotated checklist. Mycological Progress 15: 771–790. https://doi.org/10.1007/s11557-016-1207-7
  39. Zhuang, L., Ma, Y.P., Zhang, X., Luo, K.Y., Li, X., Yuan, Y. & Zhao, H. (2026) A checklist of wood-decaying fungi in Zhangguangcai Ling Mountains, northeast China. Mycosystema 45: 250338. https://doi.org/10.13346/j.mycosystema.250338

How to Cite

Zhao, Y.-L., Li, M.-R., Zhaung, L., Cai, Q. & Dai, Y.-C. (2026) A new clavarioid wood-inhabiting fungus, Artomyces tsugae (Russulales Basidiomycota), from Guangxi, southern China. Phytotaxa 765 (3): 246–256. https://doi.org/10.11646/phytotaxa.765.3.3