Abstract
Niutoushan National Forest Park, situated in Zhejiang Province, China, is recognized as one of the province's most significant forest parks due to its rich woody vegetation. Despite this, the diversity of macrofungi in the area has not been thoroughly documented. Between 2023 and 2025, a systematic survey of macrofungi was conducted within the park, resulting in the identification of 121 species, including four ascomycetes and 117 basidiomycetes. Notably, a new species, Skeletocutis sinuosa, is described here, accompanied by detailed morphological illustrations and phylogenetic analysis. The macrofungal assemblage recorded in Niutoushan National Forest Park comprises temperate, warm-temperate, subtropical, and tropical elements.
References
- Ahmadni, A.S., Kamar, N.A., Amir Hadi, M. & Terhem, R. (2023) Assessment of macrofungi diversity in Perlis State Park, Perlis, Malaysia. Journal of Wildlife and Biodiversity 8 (1): 298–307. https://doi.org/10.5281/zenodo.10206965
- Alem, D., Dejene, T., Oria-de-Rueda, J.A. & Martín-Pinto, P. (2021) Survey of macrofungal diversity and analysis of edaphic factors influencing the fungal community of church forests in dry Afromontane areas of northern Ethiopia. Forest Ecology and Management 496: 119391. https://doi.org/10.1016/j.foreco.2021.119391
- Anonymous. (1969) Flora of British Fungi. Colour identification chart. Her Majesty´s Stationery Office, London.
- Bau, T., Li, H.J., Bao, H.Y. & Li, Y. (2024) A revised checklist of poisonous mushrooms in China. Journal of Fungi 22: 301–321.
- Bian, L.S., An, Q., Wang, X.H., Chen, L. & Han, M.L. (2023) Diversity of wood-inhabiting macrofungi in the Taihang Mountains, northern China. Mycosystema 42 (11): 2188–2202. https://doi.org/10.13346/j.mycosystema.230088
- Chen, J.J., Cui, B.K. & Dai, Y.C. (2016) Global diversity and molecular systematics of Wrightoporia s.l. (Russulales, Basidiomycota). Persoonia—Molecular Phylogeny and Evolution of Fungi 37 (1): 21–36. https://doi.org/10.3767/003158516X689666
- Corrales, A., Koch, R.A., Vasco-Palacios, A.M., Smith, M.E., Ge, Z.W. & Henkel, T.W. (2022) Diversity and distribution of tropical ectomycorrhizal fungi. Mycologia 114 (6): 919–933. https://doi.org/10.1080/00275514.2022.2115284
- Cui, B.K., Li, H.J., Ji, X., Zhou, J.L., Song, J., Si, J. & Dai, Y.C. (2019) Species diversity, taxonomy and phylogeny of Polyporaceae (Basidiomycota) in China. Fungal diversity 97: 137–392. https://doi.org/10.1007/s13225-019-00427-4
- Cui, Y.J., Wu, Y.D., Jiang, Y.H., Zhu, A.H., Wu, F., Liu, H.G., Dai, Y.C. & Yuan, Y. (2025) Diversity of macrofungi in southeast Xizang 1. the wood-decay fungi. Mycology 16 (2): 635–669. https://doi.org/10.1080/21501203.2024.2379476
- Dai, Y.C. (2010) Hymenochaetaceae (Basidiomycota) in China. Fungal Diversity 45: 131–343. https://doi.org/10.1007/s13225-010-0066-9
- Dai, Y.C., Yang, Z.L., Cui, B.K., Wu, G., Yuan, H.S., Zhou, L.W., He, S.H., Ge, Z.W., Wu, F., Wei, Y.L., Yuan, Y. & Si, J. (2021) Diversity and systematics of the important macrofungi in Chinese forests. Mycosystema 40: 770–805. https://doi.org/10.13346/j.mycosystema.210036
- David, A. (1982) Étude monographique du genre Skeletocutis (Polyporaceae). Naturaliste Canadien 109: 235–272.
- Dennis, R.W.G., Reid, D.A. & Spooner, B.M. (1977) The fungi of the Azores. Kew Bulletin 32 (1): 85–136. https://doi.org/10.2307/4117263
- Dong, J.H., Gu, J.Y. & Zhao, C.L. (2023) Diversity of wood-decaying fungi in Wenshan Area, Yunnan Province, China. Mycosystema 42 (3): 638–662. https://doi.org/10.13346/j.mycosystema.220205
- Dong, J.H., Chen, M.L., Chen, M., Li, Q., Zhu, Y.J., Zhang, X.C., Zhou, C.Q., Li, W., Muhammad, A., Zhou, H.M., Jabeen, S. & Zhao, C.L. (2025) Notes, outline, taxonomy and phylogeny of wood-inhabiting Agaricales. Mycosphere 16 (1): 2599–2711. https://doi.org/10.5943/mycosphere/16/1/16
- Dong, J.H., Li, Q., Yuan, Q., Luo, Y.X., Zhang, X.C., Dai, Y.F., Zhou, Q., Liu, X.F., Deng, Y.L., Zhou, H.M., Muhammad, A. & Zhao, C.L. (2024) Species diversity, taxonomy, molecular systematics and divergence time of wood-inhabiting fungi in Yunnan-Guizhou Plateau, Asia. Mycosphere 15 (1): 1110–1293. https://doi.org/10.5943/mycosphere/15/1/10
- Du, R. & Dai, Y.C. (2020) A new species and three new combinations of Skeletocutis (Incrustoporiaceae, Basidiomycota). Mycosystema 39 (4): 637–644. https://doi.org/10.13346/j.mycosystema.200006
- Du, R. & Ji, X.H. (2019) A new species of Skeletocutis (Polyporales, Basidiomycota) from Vietnam. Journal of Microbiology & Experimentation 7 (1): 20–25. https://doi.org/10.15406/jmen.2019.07.00234
- Hall, F.R. (1999) Precision Farming: technologies and information as risk-reduction tools. https://doi.org/10.1021/bk-1999-0734.ch008
- Hu, Y.Q., Li, L.Y., Liu, X.Z., Lang, N., Qi, L.L. & Li, Y. (2024) Diversity of macrofungi in the karst area of Sino-Vietnam border in Guangxi, South China. Mycosystema 43 (10): 240106. https://doi.org/10.13346/j.mycosystema.240106
- Jayawardena, R.S., Hyde, K.D., Wang, S., Sun, Y.-R., Suwannarach, N., Sysouphanthong, P., Abdel-Wahab, M.A., Abdel-Aziz, F.A., Abeywickrama, P.D., Abreu, V.P., Armand, A., Aptroot, A., Bao, D.-F., Begerow, D., Bellanger, J.-M., Bezerra, J.D.P., Bundhun, D., Calabon, M.S., Cao, T., Cantillo, T., Carvalho, J.L.V.R., Chaiwan, N., Chen, C.-C., Courtecuisse, R., Cui, B.-K., Damm, U., Denchev, C.M., Denchev, T.T., Deng, C.Y., Devadatha, B., de Silva, N.I., dos Santos, L.A., Dubey, N.K., Dumez, S., Ferdinandez, H.S., Firmino, A.L., Gafforov, Y., Gajanayake, A.J., Gomdola, D., Gunaseelan, S., He, S.-C., Htet, Z.H., Kaliyaperumal, M., Kemler, M., Kezo, K., Kularathnage, N.D., Leonardi, M., Li, J.-P., Liao, C., Liu, S., Loizides, M., Luangharn, T., Ma, J., Madrid, H., Mahadevakumar, S., Maharachchikumbura, S.S.N., Manamgoda, D.S., Martín, M.P., Mekala, N., Moreau, P.-A., Mu, Y.-H., Pahoua, P., Pem, D., Pereira, O.L., Phonrob, W., Phukhamsakda, C., Raza, M., Ren, G.-C., Rinaldi, A.C., Rossi, W., Samarakoon, B.C., Samarakoon, M.C., Sarma, V.V., Senanayake, I.C., Singh, A., Souza, M.F., Souza-Motta, C.M., Spielmann, A.A., Su, W., Tang, X., Tian, X., Thambugala, K.M., Thongklang, N., Tennakoon, D.S., Wannathes, N., Wei, D.-P., Welti, S., Wijesinghe, S.N., Yang, H., Yang, Y., Yuan, H.-S., Zhang, H., Zhang, J., Balasuriya, A., Bhunjun, C.S., Bulgakov, T.S., Cai, L., Camporesi, E., Chomnunti, P., Deepika, Y.S., Doilom, M., Duan, W.-J., Han, S.-L., Huanraluek, N., Jones, E.B.G., Lakshmidevi, N., Li, Y., Lumyong, S., Luo, Z.-L., Khuna, S., Kumla, J., Manawasinghe, I.S., Mapook, A., Punyaboon, W., Tibpromma, S., Lu, Y.-Z., Yan, J.-Y. & Wang, Y. (2022) Fungal diversity notes 1512–1610: taxonomic and phylogenetic contributions on genera and species of fungal taxa. Fungal Diversity 117 (1): 1–272. https://doi.org/10.1007/s13225-022-00513-0
- Jörgensen, K., Clemmensen, K.E., Wallander, H. & Lindahl, B.D. (2024) Ectomycorrhizal fungi are more sensitive to high soil nitrogen levels in forests exposed to nitrogen deposition. New Phytologist 242: 1725–1738. https://doi.org/10.1111/nph.19509
- Katoh, K., Rozewicki, J. & Yamada, K.D. (2019) MAFFT online service: multiple sequence alignment, interactive sequence choice and visualization. Brief Bioinform 20 (4): 1160–1166. https://doi.org/10.1093/bib/bbx108
- Kotlaba, F. & Pouzar, Z. (1958) Polypori novi vel minus cogniti Cechoslovakiae 3. Ceská Mykologie 12: 95–104.
- Kotiranta, H. (1984) Skeletocutis jelicii: a new member of the Finnish polypore flora. Karstenia 24: 73–76. https://doi.org/10.29203/ka.1984.229
- Li, M.Q., Yan, X.F., Ren, Y.F., Zhou, L.B. & Deng, X.J. (2023) Diversity of macrofungi in Liupanshan National Nature Reserve, Ningxia. Mycosystema 42 (9): 1889–1905. https://doi.org/10.13346/j.mycosystema.220501
- Li, Y.C. & Yang, Z.L. (2021) The Boletes of China: Tylopilus s.l. Science Press & Springer Nature Singapore Pte Ltd., 418 pp. https://doi.org/10.1007/978-981-16-2986-0
- Li, Y., Li, T.H., Yang, Z.L., Tu, L.G.E. & Dai, Y.C. (2015) Atlas of Chinese macrofungal resources. Zhongyuan Farmers Press, Zhengzhou, pp. 1–1351 (in Chinese).
- Liu, Z.B., Yuan, Y., Dai, Y.C., Liu, H.G., Vlasák, J., Zeng, G.Y., He, S.H. & Wu, F. (2025a) Global diversity and systematics of Hymenochaetaceae with non-poroid hymenophore. Fungal Diversity 131: 1–97. https://doi.org/10.1007/s13225-025-00552-3
- Liu, Z.B., Liu, H.G., Vlasák, J., Gates, G.M., Dai, Y.C. & Yuan, Y. (2025b) Global diversity and phylogeny of Incrustoporiaceae (Polyporales, Basidiomycota) with an emphasis on Skeletocutis. Mycology 16 (3): 1083–1140. https://doi.org/10.1080/21501203.2024.2448145
- Lu, W., Alfagham, A.T., Tibpromma, S., Suwannarach, N., Kumla, J., Dai, D.Q., Elgorban, A.M., Chukeatirote, E. & Karunarathna, S.C. (2025) Hidden in the daylight: A polyphasic approach reveals two bioluminescent Mycenaceae species from Yunnan Province, China (Basidiomycota, Agaricales). Biodiversity Data Journal 13: p.e168858. https://doi.org/10.3897/BDJ.13.e168858
- Ma, H.X., Si, J., Dai, Y.C., Zhu, A.H., Cui, B.K., Fan, Y.G., Yuan, Y. & He, S.H. (2022) Diversity of wood-inhabiting macrofungi in Hainan Province, South China. Mycosystema 41: 695–712. https://doi.org/10.13346/j.mycosystema.210424
- Maddison, W.P. & Maddison, D.R. (2023) Mesquite: a modular system for evolutionary analysis, version 3.81. [http://www.mesquiteproject.org]
- Mao, W.L., Wu, Y.D., Liu, H.G., Yuan, Y. & Dai, Y.C. (2023) A contribution to Porogramme (Polyporaceae, Agaricomycetes) and related genera. IMA Fungus 14: 5. https://doi.org/10.1186/s43008-023-00110-z
- Miettinen, O. & Niemelä, T. (2018) Two new temperate polypore species of Skeletocutis (Polyporales, Basidiomycota). Annales Botanici Fennici 55 (4–6): 195–206. https://doi.org/10.5735/085.055.0401
- Miller, M.A., Schwartz, T., Pickett, B.E., He, S., Klem, E.B., Scheuermann, R.H., Passarotti, M., Kaufman, S. & O’Leary, M.A. (2015) A RESTful API for access to phylogenetic tools via the CIPRES science gateway. Evol Bioinform Online 11: 43–48. https://doi.org/10.4137/EBO.S21501
- Mueller, G.M., Schmit, J.P., Leacock, P.R., Buyck, B., Cifuentes, J., Desjardin, D.E., Halling, R.E., Hjortstam, K., Iturriaga, T., Larsson, K.H., Lodge, D.J., May, T.W., Minter, D., Rajchenberg, M., Redhead, S.A., Ryvarden, L., Trappe, J.M., Watling, R. & Wu, Q.X. (2007) Global diversity and distribution of macrofungi. Biodiversity and Conservation 16: 37–48. https://doi.org/10.1007/s10531-006-9108-8
- Niemelä, T. (1998) The Skeletocutis subincarnata complex (Basidiomycetes), a revision. Acta Botanica Fennica 161: 1–35.
- Nylander, J. (2004) MrModeltest V2. program distributed by the author. Bioinformatics 24: 581–583. https://doi.org/10.1093/bioinformatics/btm388
- Petersen, J.H. (1996) The Danish mycological society´s colour-chart. Foreningen til Svampekundskabens Fremme, Greve.
- Posada, D. & Crandall, K.A. (1998) MODELTEST: testing the model of DNA substitution. Bioinformatics 14 (9): 817–818. https://doi.org/10.1093/bioinformatics/14.9.817
- Qin, G.F., Qin, W.M., Wang, H.C., Zhao, J., Korhonen, K., Chen, J., Dai, Y.C. & Yuan, Y. (2024) Phylogeny and species diversity of Armillaria in China based on morphological, mating test, and GCPSR criteria. Mycology 16 (2): 777–811. https://doi.org/10.1080/21501203.2024.2404121
- Rambaut, A. (2018) Molecular evolution, phylogenetics and epidemiology. FigTree ver. 1.4.4 software. [http://tree.bio.ed.ac.uk/software/figtree/]
- Rathnayaka, A.R., Tennakoon, D.S., Jones, G.E., Wanasinghe, D.N., Bhat, D.J., Priyashantha, A.H., Stephenson, S.L., Tibpromma, S. & Karunarathna, S.C. (2025) Significance of precise documentation of hosts and geospatial data of fungal collections, with an emphasis on plant-associated fungi. New Zealand Journal of Botany 63 (2–3): 462–489. https://doi.org/10.1080/0028825X.2024.2381734
- Ren, J., Wang, K., Niu, C.Y., Kang, X., Li, S.M., Wang, L.A., Liu, T.Z., Wei, T.Z. & Li, G.J. (2024) Species diversity and funga of macrofungi in Yanshan area, North China. Mycosystema 43 (11): 240184. https://doi.org/10.13346/j.mycosystema.240184
- Shaw, S., Puthal, C., Shil, R., Saha, R.P., Majumder, R. & Ghosh, S. (2025) Chapter 4—Geographic distribution of forest fungi and their associated plant taxa. In: Forest Fungi. Academic Press, pp. 63–81. https://doi.org/10.1016/B978-0-443-18870-1.00002-0
- Si, J., Meng, G., Wu, Y., Ma, H.F., Cui, B.K. & Dai, Y.C. (2019) Medium composition optimization, structural characterization, and antioxidant activity of exopolysaccharides from the medicinal mushroom Ganoderma lingzhi. International Journal of Biological Macromolecules 124: 1186–1196. https://doi.org/10.1016/j.ijbiomac.2018.11.274
- Stamatakis, A. (2014) RAxML version 8: a tool for phylogenetic analysis and post-analysis of large phylogenies. Bioinformatics 30 (9): 1312–1313. https://doi.org/10.1093/bioinformatics/btu033
- Swofford, D.L. (2002) PAUP*: phylogenetic analysis using parsimony (*and other methods), version 4.0b10. Sinauer Associates, Sunderland. https://doi.org/10.1002/0471650129
- Tian, X., Bao, D., Han, J., Dai, D., Lu, Y., Han, M., Zheng, D., Tarafder, E., Acharya, K., Dovana, F. & Boccardo, F. (2026) Morphology and multi-locus phylogeny reveal 23 fungal taxa from terrestrial habitats. Mycology. https://doi.org/10.1080/21501203.2025.2594942
- Wang, C.G., Dai, Y.C., Kout, J., Gates, G.M., Liu, H.G., Yuan, Y. & Vlasák, J. (2024b) Multi-gene phylogeny and taxonomy of Physisporinus (Polyporales, Basidiomycota). Mycosphere 15: 1455–1521. https://doi.org/10.1080/21501203.2024.2379476
- Wang, C.G., Wu, Y.D., Zhang, X., Dai, Y.C., Li, Z.H. & Yuan, Y. (2025) Studies in phylogeny and divergence times of Irpicaceae and Meripilaceae (Polyporales, Basidiomycota), with an emphasis on Ceriporia and Meripilus including ten new species. IMA Fungus 16: e161336. https://doi.org/10.3897/imafungus.16.161336
- Wang, C.G., Zhao, H., Liu, H.G., Zeng, G.Y., Yuan, Y. & Dai, Y.C. (2023) A multi‑gene phylogeny clarifies species diversity, taxonomy, and divergence times of Ceriporia and other related genera in Irpicaceae (Polyporales, Basidiomycota). Mycosphere 14 (1): 1665–1729. https://doi.org/10.5943/mycosphere/14/1/19
- Wang, K., Zhao, M.J. & Cai, L. (2024a) Annual review on fungi nomenclature novelties in China and around the world (2023). Biodiversity Science 32: 24361. https://doi.org/10.17520/biods.2024361
- Wang, K., Zhao, M.J. & Cai, L. (2026) The Rise of China in Fungal Taxonomic Studies and Biodiversity Cataloging in the Past Decade. Journal of Fungi 12 (2): 101. https://doi.org/10.3390/jof12020101
- White, T.J., Bruns, T., Lee, S. & Taylor, J. (1990) Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. In: Innis, M.A., Gelfand, D.H., Sninsky, J.J. & White, T.J. (Eds.) PCR protocols, a guide to methods and applications. Academic Press, New York, pp. 315–322. https://doi.org/10.1038/srep34568
- Wu, F., Zhou, L.W., Yang, Z.L., Bau, T., Li, T.H. & Dai, Y.C. (2019) Resource diversity of Chinese macrofungi: edible, medicinal and poisonous species. Fungal Diversity 98: 1–76. https://doi.org/10.1007/s13225-019-00432-7
- Wu, F., Man, X.W., Tohtirjap, A. & Dai, Y.C. (2022) A comparison of polypore funga and species composition in forest ecosystems of China, North America, and Europe. Forest Ecosystems 9: 100051. https://doi.org/10.1016/j.fecs.2022.100051
- Xin, Q., Hsiang, T., Li, Y., Zhu, Z.X. & Liu, S.Y. (2023) Species diversity of macrofungi in Wangtian’e Nature Reserve, Jilin Province. Mycosystema 42 (9): 1876–1888. https://doi.org/10.13346/j.mycosystema.220514
- Xu, W.Q., Li, Y., Li, H.J., Liu, D.M., Yang, N., Zhang, Q. & He, S.H. (2023) Species diversity and resource evaluation of macrofungi in Beijing. Biodiversity Science 31: 23196. https://doi.org/10.17520/biods.2023196
- Yang, C.L., Xu, X.L., Zeng, Q., Liu, L.J., Liu, F., Deng, Y., Wang, F.H., Sun, Q.R., Jeewon, R., Hyde, K.D., Jayawardena, R.S., McKenzie, E.H.C., Wanasinghe, D.N., Liu, Y.G., Xiao, Q.G., Han, S., Yang, H., Li, S.J., Liu, L. & Xie, J.L. (2024) Bambusicolous mycopathogens in China with an update on taxonomic diversity, novel species, pathogenicity, and new insights. Mycosphere 15 (1): 4788–4918. https://doi.org/10.5943/mycosphere/15/1/21
- Yang, Y., Xu, Y., Wang, L., Jiang, Q.Q., Su, J.Q., Li, R., Zhou, H.M. & Zhao, C.L. (2025) Multigene phylogeny of seven wood-inhabiting fungal orders in Basidiomycota, and proposal of a new genus and thirteen new species. Mycosphere 16 (1): 245–295. https://doi.org/10.5943/mycosphere/16/1/4
- Yu, X.D., Zhang, S.N., Liang, X.D., Zhu, J.T., Hyde, K.D. & Liu, J.K. (2024) Bambusicolous fungi from Southwestern China. Mycosphere 15 (1): 5038–5145. https://doi.org/10.5943/mycosphere/15/1/24
- Yuan, Y., Bian, L.S., Wu, Y.D., Chen, J.J., Wu, F., Liu, H.G., Zeng, G.Y. & Dai, Y.C. (2023) Species diversity of pathogenic wood-rotting fungi (Agaricomycetes, Basidiomycota) in China. Mycology 14 (3): 204–226. https://doi.org/10.1080/21501203.2023.2238779
- Zhang, Q.Y., Liu, H.G., Papp, V., Zhou, M., Dai, Y.C. & Yuan, Y. (2023) New insights into the classification and evolution of Favolaschia (Agaricales, Basidiomycota) and its potential distribution, with descriptions of eight new species. Mycosphere 14: 777–814. https://doi.org/10.5943/mycosphere/14/1/10
- Zhao, H., Nie, Y., Zong, T.K., Wang, K., Lv, M.L., Cui, Y.J., Tohtirjap, A., Chen, J.J., Zhao, C.L., Wu, F., Cui, B.K., Yuan, Y., Dai, Y.C. & Liu, X.Y. (2023) Species diversity, updated classification and divergence times of the phylum Mucoromycota. Fungal Diversity 123 (1): 49–157. https://doi.org/10.1007/s13225-023-00525-4
- Zhao, H., Wu, Y.D., Yang, Z.R., Liu, H.G., Wu, F., Dai, Y.C. & Yuan, Y. (2024) Polypore funga and species diversity in tropical forest ecosystems of Africa, America and Asia, and a comparison with temperate and boreal regions of the Northern Hemisphere. Forest Ecosystems 11: 100200. https://doi.org/10.1016/j.fecs.2024.100200
- Zheng, D., Tibpromma, S., Lu, W., Han, M., Suwannarach, N., Elgorban, A.M., Alfagham, A.T., Yu, F., Dai, D., Zhang, L. & Karunanayake, C. (2026) Two novel species belonging to Psathyrella sections Atomatae and Hydrophilae (Psathyrellaceae, Agaricales) from Yunnan Province, China, based on morphological and multi-locus phylogenetic analyses. MycoKeys 129: 1–22. https://doi.org/10.3897/mycokeys.129.179449
- Zheng, H.F., Shao, Y.Y., Huang, H.S., Wei, Q.L., Huang, F.C. & Liu, B. (2024) Macrofungal diversity of White-headed Langurs Nature Reserve in Chongzuo, Guangxi, South China. Mycosystema 43 (10): 240043. https://doi.org/10.13346/j.mycosystema.240043
- Zhou, M., Cui, Y.J., Vlasák, J.V., Wu, Y.D., Li, C.X., Zeng, G.Y., Zhuang, L. & Yuan, Y. (2026) Global phylogeny, divergence times and evolutionary history of Trichaptum s.l. (Hymenochaetales, Basidiomycota): two new families and six new species. Mycosphere. https://doi.org/10.48130/mycosphere-0026-0005
- Zhou, M., Dai, Y.C., Vlasák, J., Liu, H.G. & Yuan, Y. (2023a) Updated systematics of Trichaptum s.l. (Hymenochaetales, Basidiomycota). Mycosphere 14 (1): 815–917. https://doi.org/10.5943/mycosphere/14/1/11
- Zhou, L.J., Geng, B.B., Zhang, G.L., Zhang, Y.H. & Tian, X.M. (2023b) Diversity and resources of wood-rotting fungi in Shandong Province. Mycosystema 42 (12): 2331–2355. https://doi.org/10.13346/j.mycosystema.230111
