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Issue: Vol. 739 No. 2: 28 Jan. 2026
Type: Article
Published: 2026-01-28
DOI: 10.11646/phytotaxa.739.2.3
Page range: 145-182
Abstract views: 32
PDF downloaded: 2

A nomenclator of Ficus sect. Americanae (Moraceae). Part II: North and Central America, and the Antilles

University of Wisconsin-Madison, Department of Botany, 430 Lincoln Dr, Madison, WI, 53706, U.S.A.; Inkaterra Asociación, Victor Larco Herrera, 130 Miraflores, Lima, Peru
University of Wisconsin-Madison, Department of Botany, 430 Lincoln Dr, Madison, WI, 53706, U.S.A.; Inkaterra Asociación, Victor Larco Herrera, 130 Miraflores, Lima, Peru
Case Western Reserve University, Department of Biology, 2080 Adelbert Road, Cleveland, OH 44106, U.S.A.
Universidade Federal Fluminense, Instituto de Ciência da Sociedade e Desenvolvimento Regional, Rua Santiago Carvalhido Filho, 80-Centro, Campos dos Goytacazes-RJ, 28035-160, Brazil; Programa de Pós-Graduação em Botânica, Escola Nacional de Botânica Tropical, Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, R. Pacheco Leão 2040, 22460-036, Rio de Janeiro - RJ, Brazil
Antilles lectotype Mexico Mixtiflores Neotropics nomenclature Rosales Spherosuke systematics typification Eudicots

Abstract

Typification underpins the stability of nomenclature, thereby making coherent taxonomic discourse possible. As part of ongoing systematic studies in Ficus sect. Americanae (Moraceae), we present the second installment of a two-part nomenclatural treatment addressing all published names within the section. The first part encompassed species described from South America, while the present contribution focuses on names from North and Central America and the Antilles. Across both treatments, we examined the relevant literature and critically assessed type vouchers for taxa described from the Linnaean era to the present. In this work, we designate 28 lectotypes, two neotypes, and two epitypes. Nomenclatural adjustments include the correction of holotype to lectotype for 24 names (ICN, Art. 9.12) and type to lectotype for four names (ICN, Art. 9.17), with the term type corrected to neotype for F. lentiginosa var. imrayana and F. eximia var. cubensis. We also designate epitypes for F. americana and F. citrifolia. These revisions provide the necessary nomenclatural stability to support the accurate application of names to phylogenetically delimited clades in forthcoming integrative treatments of Ficus sect. Americanae. Together, both indexes constitute an authoritative reference that establishes a framework for taxonomic and systematic stability and accuracy in Ficus, both regionally and globally.

References

  1. Aiton, W.T. (1789) Hortus Kewensis; Or a Cataloque of the Plants Cultivated. in the Royal Botanic Garden at Kew, vol. 3. George Nicol, London, 547 pp. https://doi.org/10.5962/bhl.title.116053
  2. Aublet, J.B.C.F. (1775) Histoire des plantes de la Guiane Françoise, 2. P.F. Didot, London, Paris, 621 pp. https://doi.org/10.5962/bhl.title.48831
  3. Berg, C.C. (1981) The Reinstatement of the West African Ficus lutea Vahl (Moraceae). Kew Bulletin 36 (3): 597–598. https://doi.org/10.2307/4117591
  4. Berg, C.C. (1986) The Ficus species (Moraceae) of Madagascar and the Comoro Islands. Bulletin du Muséum National d'Histoire Naturelle Section B, Adansonia sér. 4 8 (1): 17–55.
  5. Berg, C.C. & Simonis, J.E. (1981) The Ficus flora of Venezuela: five species complexes discussed and two new species described. Ernstia 6: 1–12.
  6. Berg, C.C. (2007) Proposals for treating four species complexes in Ficus subgenus Urostigma section Americanae (Moraceae). Blumea 52 (2): 295–312. https://doi.org/10.3767/000651907X609034
  7. Barrie, F.R. (2006) Report of the General Committee: 9. Taxon 55 (3): 795–800. https://doi.org/10.2307/25065657
  8. Bentham, G. (1861) Flora Hongkongensis: a description of the flowering plants and ferns of the island of Hongkong. L. Reeve and co., London, 482 pp. https://doi.org/10.5962/bhl.title.23084
  9. Bentham, G. (1873) Flora australiensis: a description of the plants of the Australian territory vol. 6. L. Reeve and co., London, 475 pp. https://doi.org/10.5962/bhl.title.141
  10. Benz, B.F., LópezMestas Camberos, M.L. & Ramos de la Vega, J. (2006) Organic offerings, paper, and fibers from the Huitzilapa shaft tomb, Jalisco, Mexico. Ancient Mesoamerica 17 (2): 283–296. [http://www.jstor.org/stable/26309480]
  11. Berg, C.C. (1981) A new species of Ficus (Moraceae) from Costa Rica. Acta Botanica Neerlandica 30 (3): 231–233. https://doi.org/10.1111/j.1438-8677.1981.tb00404.x
  12. Berg, C.C. (1989) Classification and distribution of Ficus. The comparative biology of figs. Experientia 45: 605–611. https://doi.org/10.1007/BF01975677
  13. Berg, C.C. (1992) Moraceae. In: Gorts-Van Rijn, A.R.A. & Jansen-Jacobs, M.J. (Eds.) Flora of the Guianas. Serie A, Phanerogams, Koeltz Scientific Books, Germany, pp. 10–92.
  14. Berg, C.C. (2003) (1587–1590) Proposals to Conserve the Names Ficus citrifolia against F. caribaea, F. maxima with a Conserved Type, F. aurea against F. ciliolosa, and F. americana against F. perforata (Moraceae). Taxon 52 (2): 368–370. https://doi.org/10.2307/3647421
  15. Berg, C.C. (2004) A new species of Ficus (Moraceae) of uncertain provenance. Brittonia 56 (1): 54–57. https://doi.org/10.1663/0007-196X(2004)056[0054:ANSOFM]2.0.CO;2
  16. Berg, C.C. (2007) Proposals for treating four species complexes in Ficus subgenus Urostigma section Americanae (Moraceae). Blumea 52 (2): 295–312. https://doi.org/10.3767/000651907X609034
  17. Berg, C.C. (2009) Moraceae (Ficus). In: Harling, G. & Persson, C. (Eds.) Flora of Ecuador. Vol. 85. University of Gothenburg, Göteborg, pp. 68–94.
  18. Berg, C.C. (2011) Ficus laurifolia (Moraceae) finally in position. Blumea 56 (2): 159. https://doi.org/10.3767/000651911X592119
  19. Berg, C.C. (2012) Moraceae. In: Davidse, G., Sousa, S.M., Knapp, S. & Chiang, F. (Eds.) Flora Mesoamericana 2 (2), Moraceae. Missouri Botanical Press, St. Louis, pp. 1–91.
  20. Berg, C.C. & DeWolf, G.P. (1975) Moraceae. In: Lanjouw, J. & Lanjouw, A.L. (Eds.) Flora of Suriname. E.J. Brill, Leiden, pp. 173–267.
  21. Berg, C.C. & Roselli, P.F. (1996) New taxa and combinations in Moraceae and Cecropiaceae from Central and South America. Novon 6 (3): 230–252. https://doi.org/10.2307/3392086
  22. Berg, C.C. & Simonis, J.E. (2000) Moraceae. In: Riina, R. (Ed.) Flora de Venezuela. Fundación Instituto Botánico de Venezuela Dr. Tobías Lasser, Caracas, pp. 5–189.
  23. Berg, C.C. & Villavicencio, X. (2004) Taxonomic studies on Ficus (Moraceae) in the West Indies, extra-Amazonian Brazil and Bolivia. Ilicifolia 5: 1–173.
  24. Berg, C.C., Vázquez-Ávila, M. & Kooy, F. (1986 [1984]) Ficus species of Brazilian Amazonia and the Guianas. Acta Amazonica 14 (1/2): Suplemento: 159–194. https://doi.org/10.1590/1809-43921984145194
  25. Berg, C.C., Carauta, J.P.P. & Machado, A.F.P. (2013) Ficus goiana, a replacement name for a Brazilian species of fig (Moraceae). Phytotaxa 108 (1): 54–56. https://doi.org/10.11646/phytotaxa.108.1.4
  26. Boldingh, J. (1914) The Flora of the Dutch West Indian Islands. II. The Flora of Curacao, Aruba and Bonaire. E. J. Brill, Leyden, 197 pp. https://doi.org/10.5962/bhl.title.2750
  27. Britton, N.L. & Millspaugh, C.F. (1920) The Bahama flora. Published by the authors, New York. https://doi.org/10.5962/bhl.title.54084
  28. Britton, N.L. (1922 [1921]) Studies of West Indian plants—X. Bulletin of the Torrey Botanical Club 48 (12): 327–352.
  29. Brummitt, R.K. (2005a) Report of the Committee for Spermatophyta: 56. Taxon 54 (2): 527–536. https://doi.org/10.2307/25065389
  30. Brummitt, R.K. (2005b) Report of the Committee for Spermatophyta: 57. Taxon 54 (4): 1093–1103. https://doi.org/10.2307/25065499
  31. Burger, W.C. (1973) Notes on the flora of Costa Rica, 3: new species in the Moraceae. Phytologia 26 (6): 421–434. https://doi.org/10.5962/bhl.part.13728
  32. Burman, J. (1757) Plantarum americanarum, fasc. 6. Amstelaedami [Amsterdam]: sumtibus autoris ... apud viduam et filium S. Schouten; etc. https://doi.org/10.5962/bhl.title.899
  33. Burman, N.L. (1768) Flora Indica: cui accedit series zoophytorum indicorum, nec non Prodromus Florae Capensis. Cornelium Haek. & Johannem Schreuderum, Amsterdam, 241 pp. https://doi.org/10.5962/bhl.title.60581
  34. Callmander, M.W., Vogt, R., Donatelli, A., Buerki, S. & Nepi, C. (2021) Otto Warburg and his contributions to the screw pine family (Pandanaceae). Willdenowia 51 (1): 5–31. https://doi.org/10.3372/wi.51.51101
  35. Carauta, J.P.P. (1989) Ficus (Moraceae) no Brasil: conservação e taxonomia. Albertoa 2: 1–365.
  36. Carvajal, S. (2001 [2000]) Ficus americana Aublet or F. perfoliata L. Ibugana. Boletín del Instituto de Botánica, Universidad de Guadalajara 8 (1–2): 169–173.
  37. Carvajal, S. (2012) Sistema para la familia Moraceae en México. Ibugana 3: 3–103.
  38. Carvajal, S. & Cuevas-Figueroa, X.M. (2005 [2003]) Ficus rzedowskiana (Moraceae, subgen. Urostigma), una especie nueva de México. Ibugana. Boletín del Instituto de Botánica, Universidad de Guadalajara 11 (2): 37–40.
  39. Carvajal, S. & Peña-Pinela, C. (1997 [1996]) Ficus jacquelineae (Moraceae, subgen.: Urostigma), a new name for a Mexican fig. Ibugana. Boletín del Instituto de Botánica, Universidad de Guadalajara 4 (1–3): 57–59.
  40. Carvajal, S., Rivera-Espinosa, R. & Palacios-Juárez, H. (2001 [2000]) Nuevas combinaciones en especies del género Ficus L. subgen. Urostigma (Moraceae) de México. Ibugana. Boletín del Instituto de Botánica, Universidad de Guadalajara 8 (1–2): 127–136.
  41. D’Arcy, W.G. (1973) New names for neotropical plants. Phytologia 25 (3): 116.
  42. Delbourgo, J. (2012) Collecting Hans Sloane. In: Walker, A., MacGregor, A. & Hunter, M. (Eds.) From books to bezoars: Hans Sloane and his collections. The British Library, London, pp. 9–23.
  43. Desfontaines, R.L. (1804) Tableau de l’École de Botanique du Museum d’Historie Naturelle, 1st edition. J.A. Brosson, Paris, 238 pp. https://doi.org/10.5962/bhl.title.13828
  44. Desfontaines, R.L. (1829) Catalogus Plantarum Horti Regii Parisiensis cum annotationibus de plantis novis aut minus cognitis, 3rd edition. J.S. Chaude, Paris, 484 pp. https://doi.org/10.5962/bhl.title.69101
  45. Domin, K. (1930) Ficus lentiginosa Vahl in Dominica. Acta botanica Bohemica 9: 46–48.
  46. Dorr, L.J., Pilz, G.E. & Boggan, J. (2009) The transfer of types from the Escuela Agrícola Panamericana to the US National Herbarium in 1956 by Louis O. Williams. Taxon 58 (3): 993–1001. https://doi.org/10.1002/tax.583030
  47. Dugand, A. (1943) Nuevas nociones sobre el género Ficus en Colombia. Caldasia 2 (8): 265–283.
  48. Dugand, A. (1944) Nuevas nociones sobre el género Ficus en Colombia III. Caldasia 2: 441.
  49. Fawcett, W. & Rendle, A.B. (1912) New plants from Jamaica. Journal of Botany, British and Foreign 50: 177–182.
  50. Fawcett, W. & Rendle, A.B. (1914) Dicotyledons: families Piperaceae to Connaracea. Flora of Jamaica 3: 1–332.
  51. Felger, R.S. & Lowe, C.H. (1970) New combinations for plant taxa in Northwestern Mexico and Southwestern United States. Journal of the Arizona Academy of Science 6 (1): 82–84. https://doi.org/10.2307/40022855
  52. Freire-Fierro, A. & Landale, A. (2011) Lectotypification of Thomas Nuttall's names applied to North American Polygala (Polygalaceae). Journal of the Botanical Research Institute of Texas 5 (1): 109–124. [https://www.jstor.org/stable/41972494]
  53. Gardner, E.M., Bruun-Lund, S., Niissalo, M., Chantarasuwan, B., Clement, W.L., Geri, C., Harrison, R.D., Hipp, A.L., Holvoet, M., Khew, G., Kjellberg, F., Liao, S., Pederneiras, L.C., Peng, Y.-Q., Pereira, J.T., Phillipps, Q., Puad, A.S.A., Rasplus, J.-Y., Sang, J., Schou, S.J., Velautham, E., Weiblen, G.D., Zerega, N.J.C., Zhang, Q. & Rønsted, N. (2023) Echoes of ancient introgression punctuate stable genomic lineages in the evolution of figs. Proceedings of the National Academy of Sciences 120 (28): e2222035120. https://doi.org/10.1073/pnas.2222035120
  54. Gasparrini, G. (1844) Nova Genera, quae super nonnullis Fici speciebus struebat. Tipographia Francisco Del Vechio, Napoli, pp. 1–11.
  55. Govaerts, R., Goyder, D. & Leeuwenberg, A. (2021) World Checklist of Moraceae. Available from: https://powo.science.kew.org/ (accessed 29 August 2025)
  56. González-Ramírez, J. & Poveda-Álvarez, L. (2003) Ficus lasiosyce (Moraceae), una nueva especie del subgénero Urostigma en el Neotrópico. Lankesteriana 8: 13–15.
  57. Greenman, J.M. (1906 [1905]) I. Descriptions of Spermatophytes from the Southwestern United States, Mexico, and Central America. Proceedings of the American Academy of Arts and Sciences 41 (9): 235–270. https://doi.org/10.2307/20013502
  58. Griffith, W. (1854) Notulae ad Plantas Asiaticas. Part IV: Posthumous Papers bequeathed to the Honourable, the East India Company, and printed by order of the government of Bengal. Charles A. Serrao, Calcutta, 764 pp.
  59. Grisebach, A.H.R. (1857) Plantae caribaeae. Abhandlungen der Königlichen Gesellschaft der Wissenschaften zu Göttingen 7: 161–286.
  60. Grisebach, A.H.R. (1859) Flora of the British West Indian Islands. Lovell Reeve & Co., London, pp. 1–152. https://doi.org/10.5962/bhl.title.143
  61. Hemsley, W.B. (1883) Biologia Centrali-Americana. Botany 3 (15): 143–148.
  62. Hernández, F. (1942) Historia de las plantas de la Nueva España. III Volúmenes. Instituto de Biología/Universidad Nacional Autónoma de México, Mexico City, 1105 pp. https://doi.org/10.5962/bhl.title.144936
  63. Hernández‐Esquivel, K.B., Piedra‐Malagón, E.M., Cornejo‐Tenorio, G., Mendoza‐Cuenca, L., González‐Rodríguez, A., Ruiz‐Sanchez, E. & Ibarra‐Manríquez, G. (2020) Unraveling the extreme morphological variation in the neotropical Ficus aurea complex (subg. Spherosuke, sect. Americanae, Moraceae). Journal of Systematics and Evolution 58 (3): 263–281. https://doi.org/10.1111/jse.12564
  64. Holmgren, P.K., Holmgren, N.H., Rodríguez, R.R. & Greuter, W. (2001) Humboldt, Willdenow, and Polygala (Polygalaceae). Taxon 50 (4): 1231–1247. https://doi.org/10.1002/j.1996-8175.2001.tb02618.x
  65. Hooker, W.J. & Walker-Arnott, G.A. (1841 [1838]) The botany of Captain Beechey’s voyage; comprising an acount of the plants collected by Messrs. Lay and Collie, and other officers of the expedition, during the voyage to the Pacific and Behring’s Strait, performed in His Majesty’s ship Blossom, under the command of Captain F.W. Beechey, R.N, F.R. & A.S., in the years 1825, 26, 27 and 28. Part 7. Henry G. Bohn, London, 485 pp. https://doi.org/10.5962/bhl.title.246
  66. Howard, R.A., Garay, L.A., Sweet, H.R. & Proctor, G.R. (1974) Flora of the Lesser Antilles: Leeward and Windward Islands, vols. 1–6. Arnold Arboretum, Harvard University, Massachusetts. https://doi.org/10.5962/t.187734
  67. Howard, R.A. (1988) Dicotyledoneae, vol. 4. In: Howard, R.A., Garay, L.A., Sweet, H.R. & Proctor, G.R. (Eds.) Flora of the Lesser Antilles: Leeward and Windward Islands. Arnold Arboretum, Harvard University, Massachusetts, pp. 4. https://doi.org/10.5962/t.187734
  68. Ibarra-Manríquez, G., Cornejo-Tenorio, G., González-Castañeda, N., Piedra-Malagón, E.M. & Luna, A. (2012) El género Ficus L. (Moraceae) en México. Botanical Sciences 90 (4): 389–452. https://doi.org/10.17129/botsci.472
  69. Jacquin, N.J. (1767) Observationum botanicarum iconibus ab auctore delineatis illustratarum, Pars II. Ex officina Krausiana, Vindobonae [Wien], 32 pp.
  70. Johnston, I.M. (1949) The botany of San Jose Island (Gulf of Panama). Sargentia 8: 1–306. https://doi.org/10.5962/p.265317
  71. Kunth, C.S. (1817) Urticeae. In: F.W.H.A. von Humboldt, A.J.A. Bonpland& C.S. Kunth (Eds.) Nova genera et species plantarum, vol. 2. Libreria Grecchi, Paris, pp. 28–50.
  72. Kunth, C.S. (1847) Enumeratio Synoptica Ficus specierum cum novarum tum congnitarum Hortii Regii Botanici Berolinenss. Annales des Sciences Naturelles Ser. 3, t. 7: 231–245.
  73. Lamarck, J.B.A.P. de (1788) Encyclopedie Methodique. Botanique, Tome second. Panckoucke, Paris, 774 pp. https://doi.org/10.5962/bhl.title.824
  74. Lamont, E.E., Sinon, S. & Jackson, E.M. (2020) John Torrey and the Torrey Botanical Society: Two hundred years studying the flora of the New York City region. Memoirs of the Torrey Botanical Society 29: 48–69. [https://www.jstor.org/stable/27003463]
  75. Liebmann, F. (1851) Mexicos og Central-Americas, neldeagtige Planter, ordo Urticaceae. Det Kongelige Danske videnskabernes selskabs skrifter 5 (2): 287–343.
  76. Linnaeus, C. (1759) Systema Naturae, ed. 10. 2. Laurentii Salvii, Stockholm, 559 pp.
  77. Linnaeus, C. (1775) D.D. Plantæ Surinamenses, Qvas, Venia Experient. Facult. Medicæ, In Reg. Acad. Upsaliensi, Præside Viro Nobiliss. atque Generoso, D:no Doct. Carolo Linné, ... Publice Ventilundam Offert Stip. Reg. Jacob Alm, Uplandus. In Auditorio Carolino Majori, Die XXIII. Juni, Anno MDCCLXXV. Horis, ante meridiem, solitis. Typis Edmannianis, Upsaliæ [Uppsala], 18 pp. https://doi.org/10.5962/bhl.title.59868
  78. Linnaeus filius, C. von (1782 [1781]) Supplementum Plantarum Systematis Vegetabilium Editionis Decimae Tertiae, generum Plantarum Editiones Sextae, et Specierum Plantarum Editionis Secundae. Editum a Carolo a Linné. Impensis Orphanotrophei, Brunsvigae, pp. 441–442. https://doi.org/10.5962/bhl.title.555
  79. López-Binnqüist, C., Quintanar-Isaías, A. & Vander Meeren, M. (2012) Mexican Bark Paper: Evidence of History of Tree Species Used and Their Fiber Characteristics. Economic Botany 66 (2): 138–148. [http://www.jstor.org/stable/41494197]
  80. Lundell, C.L. (1942) Studies of American Spermatophytes—II. Contributions from the University of Michigan Herbarium 7: 3–54. https://doi.org/10.3998/mpub.12948020
  81. Lundell, C.L. (1961) Plantae Mayanae—II. Wrightia 2 (3): 116–117.
  82. Lundell, C.L. (1975) Studies of American Plants—X. Wrightia 5 (6): 151–186.
  83. Miller, P. (1768) The Gardeners Dictionary, ed. 8. Printed by the author, London.
  84. Miquel, F.A.W. (1847) Prodromus monographiae Ficuum. London Journal of Botany 6: 527–550.
  85. Miquel, F.A.W. (1854) Flora of the isthmus of Panama. In: Seemann, B. (Ed.) The Botany of the Voyage of H.M.S. Herald: under the command of Captain Henry Kellett, R.N., C.B., during the years 1845–51. Reeve & Co., London, pp. 195–196.
  86. Miquel, F.A.W. (1862) Over de geographische verspreiding der Ficeae. Verslagen en Mededeelingen der Koninklijke Akademie van Wetenschappen, Afdeeling Natuurkunde 13: 382–415.
  87. Miquel, F.A.G. (1867) Annotationes de Ficus speciebus. Annales Musei Botanici Lugduno-Batavi 3 (10): 260–315.
  88. Mitidieri-Rivera, N. & Damián-Parizaca, A. (2025) Ficus sytsmae (Moraceae), a new species from South America and the reinstatement of Ficus schiedeana. Phytotaxa 708 (2): 167–181. https://doi.org/10.11646/phytotaxa.708.2.4
  89. Mitidieri-Rivera, N., Damián-Parizaca, A., Gardner, E.M., Plata-Castro, A.D., Zerega, N.J. & Pederneiras, L.C. (2025a) A nomenclator of Ficus sect. Americanae (Moraceae). Part I: South America. Phytotaxa 711 (3): 215–262. https://doi.org/10.11646/phytotaxa.711.3.1
  90. Mitidieri-Rivera, N., Wajer, W. & Pederneiras, L.C. (2025b) (3127) Proposal to reject the name Ficus calyculata (Moraceae). Taxon 74 (6): 1584–158. https://doi.org/10.1002/tax.70084
  91. Moench, C. (1794) Methodus Plantas Horti Botanici et Agri Marburgensis: a staminum situ describendi 1. Officina Nova Libraria Academiae, Marburgi Cattorum, 368 pp. https://doi.org/10.5962/bhl.title.304
  92. Murray, A.E. (1983) Notae spermatophytae no. 2. Kalmia 13: 1–24.
  93. Nobis, M., Klichowska, E., Wolanin, M., Nobis, A. & Nowak, A. (2022) Typification of five plant names described based on specimens collected by Józef Warszewicz in Central and South America. PhytoKeys 192: 45–61. https://doi.org/10.3897/phytokeys.192.78409
  94. Nuttall, T. (1846) The North American sylva 2. J. Dobson, Philadelphia, 123 pp.
  95. Pederneiras, L.C. (2023) Moraceae. In: Zuloaga, F.O. & Zanotti, C.A. (Eds.) Flora Vascular de la República Argentina, vol. 16. Estudio Sigma S.R.L., Buenos Aires, pp. 135–151.
  96. Pederneiras, L.C., Romaniuc-Neto, S. & Mansano, V.D.F. (2018) Nomenclatural revision of Ficus sect. Americanae (Moraceae): typification of Ficus americana and allied species. Phytotaxa 361 (2): 244–250. https://doi.org/10.11646/phytotaxa.361.2.9
  97. Pederneiras, L.C., Dussán, C.M., Mitidieri, N., Romaniuc-Neto, S., Costa, A.F. & Mansano, V.D.F. (2019) Nomenclatural revision of Ficus sect. Americanae (Moraceae): typification of Ficus crocata and allied species. Phytotaxa 414: 262–268. https://doi.org/10.11646/phytotaxa.414.5.4
  98. Pederneiras, L.C., Costa, A.F., Medeiros, H., Rivera, N.M., Forzza, R.C., Romaniuc-Neto, S. & Mansano, V.D.F. (2020a) Species diversity of Ficus L. sect. Americanae (Moraceae) in Acre, Brazil. Brittonia 72: 215–231. https://doi.org/10.1007/s12228-020-09620-1
  99. Pederneiras, L.C., Dussán, C.M., Mitidieri-Rivera, N., Romaniuc-Neto, S. & Mansano, V.D.F. (2020b) Nomenclatural revision of the Ficus sect. Americanae (Moraceae): Typification of F. citrifolia and allied species. Phytotaxa 474: 145–153. https://doi.org/10.11646/phytotaxa.474.2.4
  100. Pederneiras, L.C., Mitidieri-Rivera, N., Romaniuc-Neto, S. & Mansano, V.D.F. (2021b) Typification of Ficus sect. Americanae (Moraceae): F. aurea and F. pertusa complexes. Phytotaxa 514 (2): 149–157. https://doi.org/10.11646/phytotaxa.514.2.6
  101. Pederneiras, L.C., Zamengo, H.B., Zapata-C, D.A., Mitidieri-Rivera, N., Romaniuc-Neto, S. & Mansano, V.D.F. (2023a) Ficus subgen. Spherosuke sect. Americanae ser. Dugandii (Moraceae): A Curved Ostiolar Margin Group of Neotropical Fig Trees. Systematic Botany 48 (2): 263–282. https://doi.org/10.1600/036364423X16847773873215
  102. Pederneiras, L.C., Zamengo, H.B., Plata-Castro, A.D., Romaniuc-Neto, S. & Mansano, V.D.F. (2023b) Ficus sect. Americanae ser. Kinuppii: a ramiflorous group of Neotropical fig trees. Brittonia 75 (3): 249–268. https://doi.org/10.1007/s12228-023-09750-2
  103. Pelissari, G., Gaglioti, A. & Romaniuc-Neto, S. (2019) Sinopse de Ficus subg. Spherosuke seção Americanae: citrifolia/pertusa. Novas Edições Acadêmicas, International Book Market Service Ltd., Beau Bassin, Mauritius.
  104. Plumier, C. (1689–1697) Botanicon Americanum seu historia plantarum in Americanis insulis nascentium (unpublished), vols. 1–30. Bibliothèque Centrale, Muséum d’Histoire Naturelle, Paris, 1335 pp.
  105. Rafinesque, C.S. (1838) Sylva Telluriana. Mantisa Synoptica. New Genera and Species of Trees and Shrubs of North America. Published for the author and publisher, Philadelphia, 184 pp.
  106. Richard, A. (1850) Urticaceae. In: de la Sagra, R. (Ed.) Historia Física, Política y Natural de la Isla de Cuba, segunda parte, vol. 11, Botanica. Arthus Bertrand, Paris, pp. 218–226.
  107. Richard, A. (1850) Atlas de Botánica. In: de la Sagra, R. (Ed.) Historia Física, Política y Natural de la Isla de Cuba, segunda parte, vol. 12, Botanica Atlas. Arthus Bertrand, Paris, plates 1–89.
  108. Rossberg, G. (1935) Zur Kenntnis der westindischen Moraceen II. Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem 12: 565–587. https://doi.org/10.2307/3994852
  109. Sauvalle, F.A. (1871) Flora Cubana—Revisio Catalogi Grisebachiani vel Index Plantarum Cubensium Continua. Anales de la Academia de Ciencias Médicas Físicas y Naturales de la Habana 7: 510–514.
  110. Scott, B., Pickering, V., Coulton, R., Nyhan, J. & Carine, M. (2025) Collecting and cataloguing the world: the botanical collections of Hans Sloane (1660–1753). Systematics and Biodiversity 23 (1): 2455440. https://doi.org/10.1080/14772000.2025.2455440
  111. Seibert, R.J. & Woodson, R.E. (1937) Contributions Toward a Flora of Panama. I. Collections in the Provinces of Chiriqui, Cocle, and Panama, by R.J. Seibert during the Summer of 1935. Annals of the Missouri Botanical Garden 24 (2): 175–210. https://doi.org/10.2307/2394204
  112. Sosa, V. & Gómez-Pompa, A. (1994) Lista Florística. Flora de Veracruz, Fascículo 82. Instituto de Ecología A.C., Xalapa, 245 pp. https://doi.org/10.21829/fv.398.1994.82
  113. Stafleu, F.A. & Cowan, R.S. (1981) Taxonomic literature: a selective guide to botanical publications and collections with dates, commentaries and types, Ed. 2, Vol. 3 (Lh-O). Bohn, Scheltema & Holkema, Utrecht, 980 pp. https://doi.org/10.5962/t.207599
  114. Stafleu, F.A. & Cowan, R.S. (1983) Taxonomic literature: a selective guide to botanical publications and collections with dates, commentaries and types, Ed. 2, Vol. 4 (P-Sak). Bohn, Scheltema & Holkema, Utrecht, 1214 pp. https://doi.org/10.5962/bhl.title.48631
  115. Standley, P.C. (1917) The Mexican and central American species of Ficus. Contributions from the United States National Herbarium 20 (1): 1–35.
  116. Standley, P.C. (1923) New species of plants from Salvador. Journal of the Washington Academy of Sciences 13 (16): 363–369. https://doi.org/10.5962/bhl.part.5298
  117. Standley, P.C. (1929a) Studies of American plants—I. Field Museum of Natural History, Botanical Series 4 (8): 197–299. https://doi.org/10.5962/bhl.title.2342
  118. Standley, P.C. (1929b) Studies of American plants—II. Field Museum of Natural History, Botanical Series 4 (8): 301–345. https://doi.org/10.5962/bhl.title.2342
  119. Standley, P.C. (1930) Studies of American Plants—III. Field Museum of Natural History, Botanical Series 8 (1): 3–73. https://doi.org/10.5962/bhl.title.2342
  120. Standley, P.C. (1935) New plants from the Yucatan Peninsula. Carnegie Institution of Washington 461 (4): 51–91.
  121. Standley, P.C. (1937) Ficus. In: Standley, P.C. & Dahlgren, B.E. (Eds.) Flora of Costa Rica, part I. Field Museum of Natural History, Botanical Series 18 (1): pp. 384–388. https://doi.org/10.5962/bhl.title.2251
  122. Standley, P.C. (1940) Studies of American Plants—IX. Field Museum of Natural History, Botanical Series 22 (1): 3–62. https://doi.org/10.5962/bhl.title.2346
  123. Standley, P.C. & Steyermark, J.A. (1940) Studies of American Plants—I. Field Museum of Natural History, Botanical Series 22 (4): 221–321. https://doi.org/10.5962/bhl.title.2272
  124. Standley, P.C. & Williams, L.O. (1950) Plantas nuevas. Ceiba 1 (2): 78–79.
  125. Standley, P.C. & Williams, L.O. (1951) Plantae Centrali-Americanae, II. Ceiba 1 (4): 231–254.
  126. Standley, P.C. & Williams, L.O. (1953) Plantae Centrali-Americanae, V. Ceiba 3 (3): 187–223.
  127. Stehlé, H., Stehlé, M. & Quentin, L. (1938) Catalogue des Phanérogames et fougères. In: Stehlé, H. (Ed.) Flore de la Guadeloupe et dépendances 2, 1. Basse-Terre.
  128. Stevens, W.D., Ulloa, C., Pool, A. & Montiel, O.M. (2001) Flora de Nicaragua, vol. 85. Missouri Botanical Garden Press, St. Louis, 943 pp.
  129. Steudel, E.T. (1840) Nomenclator Botanicus ed. 2, 1. J.G. Cottae, Stuttgartiae et Tubingae, 852 pp.
  130. Swartz, O. (1788) Monandri, classis I. Monogynia. Brosimum. In: Swartz, O. (Ed.) Nova Genera et Species Plantarum seu Prodromus. Academia M. Swederi, Holmiae [Stockholm], pp. 12.
  131. Turland, N.J., Wiersema, J.H., Barrie, F.R., Gandhi, K.N., Gravendyck, J., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Klopper, R.R., Knapp, S., Kusber, W.-H., Li, D.-Z., May, T.W., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. & Zamora Señoret, J.C. (2025) International Code of Nomenclature for algae, fungi, and plants (Madrid Code). Regnum Vegetabile 162. University of Chicago Press, Chicago and London, 303 pp. https://doi.org/10.7208/chicago/9780226839479.001.0001
  132. Urban, I. (1900) Species novae, praesertim portoricences. Supplementum. In: Urban, I. (Ed.) Symbolae Antillanae, vol. 1, no. 3. Berlin, Paris, London, pp. 472–481.
  133. Urban, I. (1905) Flora Portoricensis. Moraceae. In: Urban, I. (Ed.) Symbolae Antillanae, vol. 4. Leipzig, Paris, London, pp. 195–199.
  134. Urban, I. (1917) XVII. Sertum antillanum. IV. Repertorium Specierum Novarum Regni Vegetabilis 15: 98–112. https://doi.org/10.1002/fedr.19170150407
  135. Vahl, M. (1805) Enumeratio plantarum vel ab aliis, vel ab ipso observatarum, cum earum differentiis specificis, synonymis selectis et descriptionibus succinctis 2. Impensis auctoris & prostat apud J.H. Schubothe, Hauniae [Copenhagen], pp. 181–204. https://doi.org/10.5962/bhl.title.7564
  136. Warburg, O. (1903) VIII. Ficus. In: Urban, I. (Ed.) Symbolae Antillanae, seu, Fundamenta florae Indiae Occidentalis, vol. 3. Fratres Borntraeger, Lipsiae [Leipzig], pp. 453–491.
  137. Watson, S. (1891a [1889]) VI. Contributions to American Botany. Proceedings of the American Academy of Arts and Sciences 24: 36–87. https://doi.org/10.2307/20021550
  138. Watson, S. (1891b) XI. Contributions to American Botany. Proceedings of the American Academy of Arts and Sciences 26: 124–163. https://doi.org/10.2307/20013481
  139. Willdenow, C.L. (1806) Caroli a Linné Species plantarum, vol. 4. G.C. Nauk, Berlin, 1157 pp.

How to Cite

Mitidieri-Rivera, N., Damián-Parizaca, A., Gardner, E. & Pederneiras, L.C. (2026) A nomenclator of Ficus sect. Americanae (Moraceae). Part II: North and Central America, and the Antilles. Phytotaxa 739 (2): 145–182. https://doi.org/10.11646/phytotaxa.739.2.3