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Issue: Vol. 739 No. 1: 23 Jan. 2026
Type: Article
Published: 2026-01-23
DOI: 10.11646/phytotaxa.739.1.2
Page range: 35-57
Abstract views: 110
PDF downloaded: 10

Synopsis of Acalypha (Euphorbiaceae) of the West Indies (Caribbean Region), with a focus on endemic species

Universidad Autónoma de Madrid, Faculty of Sciences, Department of Biology, Campus de Cantoblanco, C/ Darwin 2, 28049 Madrid, Spain; Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), C/ Darwin 2, 28049 Madrid, Spain
Universidad Autónoma de Madrid, Faculty of Sciences, Department of Biology, Campus de Cantoblanco, C/ Darwin 2, 28049 Madrid, Spain
Universidad de Alcalá, Faculty of Sciences, Department of Life Sciences Alcalá de Henares, 28805 Madrid, Spain
Universidad Autónoma de Madrid, Faculty of Sciences, Department of Biology, Campus de Cantoblanco, C/ Darwin 2, 28049 Madrid, Spain; Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), C/ Darwin 2, 28049 Madrid, Spain
Universidad Autónoma de Madrid, Faculty of Sciences, Department of Biology, Campus de Cantoblanco, C/ Darwin 2, 28049 Madrid, Spain; Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), C/ Darwin 2, 28049 Madrid, Spain
Jardín Botánico Nacional, Universidad de La Habana, Carretera del Rocío km 3, Calabazar, Boyeros, La Habana. 19230, Cuba
Universidad Complutense de Madrid, Faculty of Biological Sciences, Department of Biodiversity, Ecology and Evolution, C/ José Antonio Novais, 12, 28040 Madrid, Spain
biodiversity endemicity identification key nomenclatural review taxonomy typification Eudicots

Abstract

This contribution provides a critical synopsis of Acalypha species in the West Indies, with a focus on the region’s endemic species. It includes a comprehensive nomenclatural revision, primarily based on the study of type specimens. In total, 107 names associated with West Indian Acalypha are compiled, recognising 36 native species, 28 of which are endemic, including the nearly endemic A. chamaedrifolia. Additionally, five species are identified as introduced, while four are considered dubious and excluded. Lectotypes are designated for 41 names, and four names are proposed as new synonyms. We also present the first identification key for Acalypha at a regional scale, provide updated country-level distribution data for each species, and offer a comprehensive index of all published names, cross-referenced with the currently accepted names.

References

  1. Acevedo‑Rodríguez, P. & Strong, M.T. (2012) Catalogue of seed plants of the West Indies. Smithsonian Contributions to Botany 98: 1–1192.
  2. Adams, C.D., Proctor, G.R. & Read, R.W. (1972) Flowering Plants of Jamaica. University of the West Indies, Mona, 848 pp.
  3. Alain, F.S.C. (1953) Dicotiledóneas: Malpighiaceae a Myrtaceae. Contribuciones Ocasionales del Museo de Historia Natural del Colegio “De La Salle” 13 (3): 1–502.
  4. Almirall, A.L. (2013) Contribución al catálogo de flora cubana: endemismos de suelos derivados de ofiolitas. Botanica Complutensis 37: 139–156. https://doi.org/10.5209/rev_BOCM.2013.v37.42276
  5. Almirall, A.L. (2016) The Cuban endemic flora: Angiosperms and gymnosperms. National Museum of Natural History, La Habana. https://doi.org/10.13140/RG.2.1.2645.2729
  6. Anonymous (1978) Catálogo de plantas colectadas por el Padre Fuertes (facsimile). Academia de Ciencias de La República Dominicana, Santo Domingo.
  7. Baksh-Comeau, Y.S., Maharaj, S.S., Adams, C.D, Harris, S.A., Filer, D.L. & Hawthorne, W.D. (2016) An annotated checklist of the vascular plants of Trinidad and Tobago with analysis of vegetation types and botanical “hotspots.” Phytotaxa 250: 1–431. https://doi.org/10.11646/phytotaxa.250.1
  8. Barker, H.D. & Dardeau, S. (1930) Flore d’Haiti. Service technique du Départment de l’Agriculture, Port au Prince, 456 pp.
  9. Berazaín, R., Areces, F., Lazcano, J.C. & González-Torres, L.R. (2005) Lista roja de la flora vascular cubana. Jardín Botánico Atlántico, Gijón.
  10. Berazaín, R., Bécquer, E. & González-Torres, L.R. (2008) Euphorbiaceae y Phyllanthaceae. In: González-Torres, L.R., Rankin, R., Leiva, A.T & Palmarola, A. (Eds.) Categorización preliminar de taxones de la flora de Cuba―2008. Bissea 2 (número especial), pp. 41–53.
  11. Britton, N.L. (1912) Studies of West Indian Plants. Bulletin of the Torrey Botanical Club 39: 1–14.
  12. Britton, N.L. (1920) Description of Cuban plants new to science. Memoirs of the Torrey Botanical Club 16: 57–118.
  13. Britton, N.L. & Wilson, P. (1924) Botany of Porto Rico and the Virgin Islands. In: Scientific Survey of Porto Rico and the Virgin Islands 5 (4). New York Academy of Sciences, New York, pp. 475–676.
  14. Browne, P. (1756) The civil and natural history of Jamaica in three parts. Published by the author, London, 503 pp., 49 pl.
  15. Brummitt, R.K. (2001) World Geographical Scheme for Recording Plant Distributions, 2nd ed. Plant Taxonomic Database Standards, No. 2. International Working Group on Taxonomic Databases for Plant Sciences (TDWG). Hunt Institute for Botanical Documentation Carnegie Mellon University, Pittsburgh, 137 pp.
  16. Burman, N.L. (1768) Flora Indica: cui accedit series zoophytorum Indicorum, nec non prodromus florae Capensis. Cornelius Haak, Leiden & Johannes Schreuder, Amsterdam, 241 pp.
  17. Cardiel, J.M. (1999) Contribuciones a la flora de Venezuela: revisión del género Acalypha (Euphorbiaceae). Acta Botanica Venezuelica 22 (2): 255–324.
  18. Cardiel, J.M. (2015) Euphorbiaceae (Acalypha). In: Jørgensen, P.M., Nee, M.H. & Beck, S.G. (Eds.) Catálogo de las Plantas Vasculares de Bolivia. Missouri Botanical Garden Press, Saint Louis, pp. 587–589.
  19. Cardiel, J.M. (2020) Acalypha. Biology Department, Universidad Autónoma de Madrid. Occurrence dataset. Available from: https://doi.org/10.15468/yhsqkx (accessed 1 June 2024)
  20. Cardiel, J.M., Cordeiro de Sousa, A.A., Cordeiro, I., Rossi Caruzo, M.B., Marques da Silva, O.L., Muñoz-Rodríguez, P., López, A. & Montero-Muñoz, I. (2022) Updated synopsis of Acalypha L. (Euphorbiaceae, Acalyphoideae) from Brazil. Plant Systematics and Evolution 308: 24. https://doi.org/10.1007/s00606-022-01816-0
  21. Cardiel, J.M. & Montero-Muñoz, I. (2018) Synopsis of Acalypha (Euphorbiaceae) of West Tropical Africa, including Cameroon, Chad, Equatorial Guinea, Gabon, and São Tomé and Príncipe. Plant Systematics and Evolution 304: 93–110. https://doi.org/10.1007/s00606-017-1453-4
  22. Cardiel, J.M., Montero-Muñoz, I., Gamarra, R., Ortúñez, E. & Muñoz-Rodríguez, P. (2025b) A nomenclatural review of Acalypha L. (Euphorbiaceae) of mainland Africa. Taxon 74: 1217–1247. https://doi.org/10.1002/tax.13388
  23. Cardiel, J.M. & Muñoz-Rodríguez, P. (2012) Synopsis of Acalypha (Euphorbiaceae) of Continental Ecuador. PhytoKeys 17: 1–17. https://doi.org/10.3897/phytokeys.17.3190
  24. Cardiel, J.M., Muñoz-Rodríguez, P. & Muñoz-Garmendia, F. (2013a) Revised taxonomy and nomenclature of Acalypha sect. Communes (Euphorbiaceae), a complex group of species widespread in the north of the Southern Cone. Taxon 62: 1295–1303. https://doi.org/10.12705/626.11
  25. Cardiel, J.M. & Muñoz-Rodríguez, P. (2015) Synopsis of Acalypha (Euphorbiaceae) of Argentina, Paraguay, and Uruguay. Annals of the Missouri Botanical Garden 101: 384–405.
  26. Cardiel, J.M., Muñoz-Rodríguez, P., González-Berdasco, A. & Montero-Muñoz, I. (2023) Catalogue and Red List of Acalypha L. (Euphorbiaceae) from South America. European Journal of Taxonomy 886: 1–92. https://doi.org/10.5852/ejt.2023.886.2201
  27. Cardiel, J.M, Muñoz-Rodríguez, P., Montero-Muñoz, I., Gamarra, R. & Ortúñez, E. (2025a) Acalypha Taxonomic Information System. Universidad Autónoma de Madrid, Madrid. Available from: http://www.acalypha.es (accessed January 2025)
  28. Cardiel, J.M., Nee, M. & Muñoz-Rodríguez, P. (2013b) Synopsis of Acalypha L. (Euphorbiaceae) of Peru and Bolivia, with description of a new species. Anales del Jardin Botánico de Madrid 70: 152–177. https://doi.org/10.3989/ajbm.2366
  29. Carrington, C.M.S., Krupnick, G & Acevedo-Rodríguez, P. (2017) Herbarium-based preliminary conservation assessments of Lesser Antillean endemic seed plants reveal a flora at risk. Botanical Review 83: 107–151.
  30. Carrington, C.M.S., Edwards, R.D. & Krupnick, G. (2018) Assessment of the distribution of seed plants endemic to the Lesser Antilles in terms of habitat, elevation, and conservation status. Caribbean Naturalist 2: 30–47.
  31. Correll, D.S. & Correll, H.B. (1982) Flora of the Bahama Archipelago. Including the Turks and Caicos Islands. Cramer, J. (Ed.) Vaduz, 1692 pp.
  32. Coulter, J.M. (1894) Euphorbiaceae. In: Committee of the Botanical Club, American Association for the Advancement of Science (Eds.) List of Pteridophyta and Spermatophyta growing without Cultivation in Northeastern North America. Memoirs of the Torrey Botanical Club 5: pp. 213.
  33. Fawcett, W. & Rendle, A.B. (1919) Notes on Jamaica Plants. Euphorbiaceae II. Journal of Botany, British and Foreign 57: 312–314.
  34. Fawcett, W. & Rendle, A.B. (1920) Dicotyledons. Families Leguminosae to Callitrichaceae. In: Flora of Jamaica, containing descriptions of the flowering plants known from the island 4. British Museum (Natural History), London, 369 pp.
  35. Fitzpatrick, S.M. & Keegan, W.F. (2007) Human impacts and adaptations in the Caribbean Islands: An historical ecology approach. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 98: 29–45.
  36. Forster, J.G.A. (1786) Florulae insularum Australium: prodromus. Dietrich, Göttingen, 103 pp.
  37. Francisco-Ortega, F., Santiago-Valentín, E., Acevedo-Rodríguez, P., Lewis, C., Pipoly III, J., Meerow, A.W. & M. Maunder, M. (2007) Seed plant genera endemic to the Caribbean Island Biodiversity Hotspot: A review and a molecular phylogenetic perspective. The Botanical Review 73: 183–234.
  38. García-Mendoza, A.J. & Meave, J.A. (2012) Diversidad florística de Oaxaca: de musgos a angiospermas (colecciones y listas de especies), ed. 2. Instituto de Biología, Universidad Nacional Autónoma de México, 351 pp.
  39. González-Oliva, L., González-Torres, L.R., Palmarola, A., Barrios, D. & Testé, E. (2015) Categorización de taxones de la flora de Cuba―2015. Bissea 9 (4): 3–707.
  40. González‑Torres, L.R., Palmarola, A., González‑Oliva, L., Bécquer, E.R., Testé, E. & Barrios, D. (2016) Lista roja de la flora de Cuba. Bissea 10 (1): 1–352. https://doi.org/10.13140/RG.2.2.24056.65288
  41. Gooding, E.G.B., Loveless, A.R. & Proctor, G.R. (1965) Flora of Barbados. H. M. Stationery Office, London, 486 pp.
  42. Grisebach, A.H.R. (1859) Flora of the British West Indian Islands 1. Lovell Reeve & Co, London, pp. 1–192.
  43. Grisebach, A.H.R. (1865) Diagnosen neuer Euphorbiaceen aus Cuba. Nachrichten von der Königlichen Gesellschaft der Wissenschaften und von der Georg-Augusts-Universität 7: 171–181.
  44. Greuter, W. & Rankin‑Rodríguez, R. (2016) The spermatophyta of Cuba: a preliminary checklist. Part II: Checklist. Botanischer Garten & Botanisches Museum Berlin‑Dahlem; Jardín Botánico Nacional, Universidad de La Habana, 398 pp.
  45. Gutiérrez, J.E., Testé, E. & García Beltrán, J.A. (2024) Euphorbiaceae. In: GEPC, Catálogo de las Plantas de Cuba. Planta!, thePlantlife Conservation Society, Vancouver, pp. 463–497.
  46. Hornemann, J.W. (1819) Hortus regius Botanicus Hafniensis, in usum Tyronum et botanophilorum. Supplementum. E.A.H. Möller, Kopenhagen, 172 pp.
  47. Hosein, F.N., Austin, N., Maharaj, S., Johnson, S., Rostant, M., Ramdass, A.C. & Rampersad, S.N. (2016) Utility of DNA barcoding to identify rare endemic vascular plant species in Trinidad. Ecology and Evolution 7: 7311–7333.
  48. Howard, R.A. (1986) Three new species of Dicotyledoneae from St. Lucia, West Indies. Phytologia 61: 1–4.
  49. Howard, R.A. (1988) Charles Wright in Cuba, 1856–1867. Chadwyck-Healey, Alexandria, 90 pp.
  50. Howard, R.A. (1989) Flora of the Lesser Antilles. Leeward and Windward Islands 5: Dicotyledoneae—Part 2. Arnold Arboretum of Harvard University, Cambridge, 604 pp.
  51. Hurusawa, I. (1954) Eine nochmalige Durchsicht des herkömmlichen Systems der Euphorbiaceen im weiteren Sinne. Journal of the Faculty of Science: University of Tokyo, Section 3, Botany 6: 209–342.
  52. Iturralde‑Vinent, M.A. (2005) La paleogeografía del Caribe y sus implicaciones para la biogeografía histórica. Revista del Jardín Botánico Nacional (La Habana) 25–26: 49–78.
  53. Jacquin, N.J. von. (1760) Enumeratio Systematica Plantarum. Theodor Haak, Leiden, 41 pp.
  54. Jacquin, N.J. von (1790) Icones Plantarum Rariorum 3. C.F. Wappler, Wien, 24 pp.
  55. Jacquin, N.J. von (1797) Plantarum Rariorum Horti Caesarei Schoenbrunnensis. C.F. Wappler, Wien etc., 70 pp.
  56. Kelly, D.L. (1988) The threatened flowering plants of Jamaica. Biological Conservation 46: 201–216. https://doi.org/10.1016/0006-3207(88)90068-7
  57. Kuntze, O. (1891) Revisio Generum Plantarum 2. Arthur Felix, Leipzig, Dulau & Co., London, pp. 375–1011.
  58. Lamarck, J.–B. (1786) Encyclopédie méthodique. Botanique 2 (1). Chez Panckouke, Paris, Chez Plomteux, Liege, pp. 1–400.
  59. Levin, G.A. (2016) Acalypha (Euphorbiaceae). In: Flora of North America Editorial Committee (Eds.) Flora of North America North of Mexico, Magnoliophyta 12: Vitaceae to Garryaceae. Oxford University Press, New York, pp. 162–172.
  60. Levin, G.A., Cardinal-McTeage, W.M., Steinmann, V.W. & Sagun, V.G. (2022) Phylogeny, classification, and character evolution of Acalypha (Euphorbiaceae: Acalyphoideae). Systematic Botany 47: 477–497. https://doi.org/10.1600/036364422X16512572275034
  61. Levin, G.A., Steinmann, V.W. & Cardiel, J.M. (2025) Proposal to conserve the name Acalypha ostryifolia Riddell ex J.M.Coult. against A. persimilis Müll.Arg. (Euphorbiaceae). Taxon 74: 469–470. https://doi.org/10.1002/tax.13337
  62. Linnaeus, C. (1753) Species Plantarum. Laurentius Salvius, Stockholm, 1200 pp.
  63. Linnaeus, C. (1759) Systema Naturae, Editio Decima, 2. Laurentius Salvius, Stockholm, pp. 533–1327.
  64. Liogier, A.H. (1986) La Flora de la Española IV. Universidad Central Del Este, San Pedro de Macorís, 317 pp.
  65. Liogier, A.H. (1988) Descriptive Flora of Puerto Rico and adjacent islands. Spermatophyta 2. Editorial de la Universidad de Puerto Rico, Río Piedras, 481 pp.
  66. Liogier, A.H. & Martorell, L.F. (1982) Descriptive flora of Puerto Rico and adjacent islands: a systematic synopsis. Editorial de la Universidad de Puerto Rico, Río Piedras, 382 pp.
  67. Liogier, A.H. & Martorell, L.F. (2000) Flora of Puerto Rico and adjacent islands: A systematic synopsis, 2nd ed. Editorial de la Universidad de Puerto Rico, Río Piedras, 382 pp.
  68. McNeill, J. (2024) Holotype specimens and type citations: general issues. Taxon 63 (5): 1112–1113. https://doi.org/10.12705/635.7
  69. Mittermeier, R.A., Robles Gil, P., Hoffmann, M., Pilgrim, J., Brooks, T., Goettsch Mittermeier, C., Lamoreux, J. & Fonseca, G.A.B. da (2004) Hotspots revisited: Earth’s biologically richest and most endangered terrestrial ecoregions. CEMEX, Conservation International, Houston, 392 pp.
  70. Moench, C. (1802) Supplementum ad Methodum Plantas a Staminum situ Describendi. In officina nova libraria academiae, Marburg, 328 pp.
  71. Montero-Muñoz, I., Cardiel, J.M. & Levin, G.A. (2018) Nomenclatural review of Acalypha (Euphorbiaceae) of the Western Indian Ocean Region (Madagascar, the Comoros Archipelago, the Mascarene Islands and the Seychelles Archipelago). PhytoKeys 108: 85–116. https://doi.org/10.3897/phytokeys.108.27284
  72. Montero-Muñoz, I., Levin, G.A. & Cardiel, J.M. (2023) Monograph of Acalypha L. (Euphorbiaceae) of the Western Indian Ocean Region, with the description of a new species from Mayotte. Adansonia 45 (26): 395–496. https://doi.org/10.5252/adansonia2023v45a26
  73. Moscoso, R.M. (1943) Catalogus Florae Domingensis. Universidad de Santo Domingo, 732 pp.
  74. Muñoz-Rodríguez, P., Cardiel, J.M. & Atha, D. (2014) Acalypha subgenus Linostachys (Euphorbiaceae, Acalyphoideae): a global review. Phytotaxa 166: 199–221. https://doi.org/10.11646/phytotaxa.166.3.2
  75. Muñoz-Rodríguez, P., Carruthers, T., Wells, T., Sumadijaya, A., Wood, J.R.I. & Scotland, R.W. (2024) The research behind a taxonomic monograph: a case study from Ipomoea (Convolvulaceae). Kew Bulletin 79: 897–914. https://doi.org/10.1007/s12225-024-10184-6
  76. Müller Argoviensis, J. (1865) Euphorbiaceae. Vorläufige Mittheilungen aus dem für De Candolle’s Prodromus bestimmten Manuscript über diese Familie. Linnaea 34: 1–224.
  77. Müller Argoviensis, J. (1866) Euphorbiaceae excl. Euphorbieae. In: Candolle, A.L.P-P. de (Ed.) Prodromus Systematis Naturalis Regni Vegetabilis 15 (2). Victor Masson, Paris, pp. 189–1286.
  78. Pax, F.A. & Hoffmann, K. (1924) Euphorbiaceae-Crotonoideae-Acalypheae-Acalyphinae. In: Engler, A. (Ed.) Das Pflanzenreich IV. 147. XVI (Heft 85). Engelmann, Leipzig, pp. 1–231.
  79. Poiret, J.L.M. (1804) Encyclopédie Méthodique. Botanique 6 (1). Chez H. Agasse, Paris, 384 pp.
  80. Poeppig, E. (1841) Acalypha. In: Poeppig, E. & Endlicher, S. (Eds.) Nova Genera ac Species Plantarum 3. Sumptibus F. Hofmeister, Leipzig, pp. 21–22.
  81. Prain, D. & Hutchinson, J. (1913) Notes on some species of Acalyha. Bulletin of Miscellaneous Information, Kew 1913: 1–28.
  82. Proctor, G.R. (1984) Flora of the Cayman Islands. Kew Bulletin Additonal Series 11, 851 pp.
  83. Radcliffe-Smith, A. & Govaerts, R. (1997) New names & new combinations in the Euphorbiaceae-Acalyphoideae. Kew Bulletin 52: 477–481.
  84. Rafinesque, C.S. (1836) New flora and botany of North America. First part. Printed for the author and publisher, Philadelphia, 108 pp.
  85. Rafinesque, C.S. (1838) Sylva telluriana. Printed for the author and publisher, Philadelphia, 183 pp.
  86. Ricklefs, R. & Bermingham, E. (2007) The West Indies as a laboratory of biogeography and evolution. Philosophical Transactions of the Royal Society of London B 363: 2393–2413. https://doi.org/10.1098/rstb.2007.2068
  87. Riddell, J.L. (1835) A synopsis of the flora of the western states. E. Deming, Cincinnati, 116 pp.
  88. Rodriguez‑Silva, R. & Schlupp, I. (2021) Biogeography of the West Indies: a complex scenario for species radiations in terrestrial and aquatic habitats. Ecology and Evolution 11: 2416–2430. https://doi.org/10.1002/ece3.7236
  89. Roig, J.T. & Mesa, J. (1988) Diccionario botánico de nombres vulgares cubanos. Editorial Científico-Técnica, La Habana, 1142 pp.
  90. Sagra, R. de la (Ed.) (n.d.) Historia Física, Política y Natural de la Isla de Cuba 11. A. Bertrand, Paris, etc, 339 pp.
  91. Schlechtendal, D.F.L. von (1846) Plantae Leiboldianae. Linnaea 19: 234–312.
  92. Sloane, H. (1707) A voyage to the islands Madera, Barbados, Nieves, S. Christophers and Jamaica 1. Printed by B.M. for the author, London, 264 pp.
  93. Smith, M.L., Blair Hedges, S., Buck, W., Hemphill, A., Inchaustegui, S., Ivie, M.A., Martina, D., Maunder, M. & Ortega, J.F. (2004) Caribbean Islands. In: Mittermeier, R.A.P., Robles, M., Hoffmann, J., Pilgrim, T., Brooks, C., Mittermeier, C., Lamoreux, J. & Fonseca, G.A.B. da (Eds.) Hotspots revisited: Earth’s biologically richest and most endangered terrestrial ecoregions. CEMEX, Conservation International, Houston, pp. 112–118.
  94. Sprengel, K.P.J. (1826) Systema vegetabilium, editio decima sexta3. Dieterich, Göttingen, 900 pp.
  95. Swartz, O. (1788) Nova genera et species plantarum seu Prodromus. In bibliopoliis Acad. M. Swederi, Stockholm, 152 pp.
  96. Swartz, O. (1800) Flora Indiae Occidentalis: aucta atque illustrata sive descriptiones plantarum in Prodromo recensitarum 2 (2). Jacobi Palmii, Erlangen, 312 pp.
  97. Thiers, B. (2025) Index Herbariorum: a global directory of public herbaria and associated staff. The New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (accessed 1 May 2025)
  98. Thunberg, C.P. (1784) Flora japonica sistens plantas insularum japonicarum secundum systema sexuale emendatum redactas ad XX classes, ordines, genera et species cvm differentiis specificis, synonymis paucis, descriptionibus concinnis et XXXIX iconibus adjectis. J.G. Müller, Leipzig, 418 pp.
  99. Turland, N.J., Wiersema, J.H., Barrie, F.R., Gandhi, K.N., Gravendyck, J., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Klopper, R.R., Knapp, S., Kusber, W.-H., Li, D.-Z., May, T.W., Monro, A.M., Prado, J., Price, M.J., Smith, G.F. & Zamora Señoret, J.C. (2025) International Code of Nomenclature for algae, fungi, and plants (Madrid Code). Regnum Vegetabile 162. University of Chicago Press, Chicago. https://doi.org/10.7208/chicago/9780226839479.001.0001
  100. Urban, I. (1905) Flora portoricensis. In: Symbolae Antillanae 4 (2). Fratres Borntraeger, Lipsiae, pp. 193–352.
  101. Urban, I. (1908) VIII. Nova genera et species III. Symbolae Antillanae, seu, Fundamenta florae Indiae Occidentalis 5 (3–4): 287–531.
  102. Urban, I. (1919) Sertum antillanum 7. Repertorium Specierum Novarum Regni Vegetabilis 15: 397–415.
  103. Urban, I. (1920) Flora domingensis. In: Symbolae Antillanae seu fundamenta Florae Indiae Occidentalis, vol. 8 (1). Fratres Borntraeger, Lipsiae, 480 pp.
  104. Urban, I. (1921) Plantae haitienses novae vel rariores a cl. Er. L. Ekman 1917 lectae. Arkiv för Botanik 17 (7): 1–72.
  105. Urban, I. (1922) XXXVII. Sertum antillanum. XVI. Repertorium Specierum Novarum Regni Vegetabilis 18: 187–199.
  106. Urban, I. (1926) Plantae Haitienses novae vel rariores a cl. Er. L. Ekman 1917 lectae. Arkiv för Botanik 17 (7): 1–72.
  107. Urban, I. (1930) Sertum Antillanum XXX. Repertorium Specierum Novarum Regni Vegetabilis 28: 209–235. https://doi.org/10.1002/fedr.4870281402
  108. Urquiola, A., Gonzalez‑Oliva, L., Novo Carbó, R. & Acosta, Z. (2010) Libro Rojo de la Flora Vascular de la Provincia Pinar del Río. Servicio de Publicaciones, Universidad de Alicante, 458 pp.
  109. Vellozo, J.M. da C. (1831) Florae Fluminensis Icones 10. Ex off. lithogr. Senefelder, Paris, 143 tt.
  110. Willdenow, C.L. (1805) Species Plantarum. Ed. 4, 4 (1). Impensis G. C. Nauk, Berlin, 140 pp.

How to Cite

Cardiel, J.M., Sancho-García, I., Montero-Muñoz, I., Gamarra, R., Ortúñez, E., Gutiérrez, J. & Muñoz-Rodríguez, P. (2026) Synopsis of Acalypha (Euphorbiaceae) of the West Indies (Caribbean Region), with a focus on endemic species. Phytotaxa 739 (1): 35–57. https://doi.org/10.11646/phytotaxa.739.1.2