Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2024-07-23
Page range: 225-242
Abstract views: 33
PDF downloaded: 2

A nomenclatural and taxonomic review of the names and new combinations published between 1881 and 1930 in the genus Kitchingia, basionym of Kalanchoe subg. Kitchingia (Crassulaceae subfam. Kalanchooideae), by John Gilbert Baker (1834–1920), Otto Stapf (1857–1933), and Alwin Berger (1871–1931)

Ria Olivier Herbarium, Department of Botany, Nelson Mandela University, P.O. Box 77000, Gqeberha, 6031 South Africa
Madagascar typification Eudicots

Abstract

During a long career as a botanist in the Herbarium of the Royal Botanic Gardens, Kew, United Kingdom, John Gilbert Baker (1834–1920) described one new genus, Kitchingia, in Crassulaceae subfam. Kalanchooideae, and eight species in this genus. He in addition published a new combination in Kitchingia for a species originally published in Kalanchoe. Otto Stapf (1857–1933), Austrian-born Kew botanist, described one new species of Kitchingia, and Alwin Berger (1871–1931), the German-born succulent plant expert, published two new combinations in Kitchingia. The genus Kitchingia was in the past variously upheld in Crassulaceae subfam. Kalanchooideae, but is today recognised in Kalanchoe at the rank of subgenus, as Kalanchoe subg. Kitchingia. The nomenclature and taxonomy of the names published in or transferred to Kitchingia by Baker, Stapf, and Berger are reviewed and discussed in a historical context. Where necessary, the names are typified.

References

  1. Adanson, M. (1763) Familles des plantes. Contenant une Préface Historike sur l’état ancien & actuel de la Botanike, & une Téorie de cette Science. II. Partie. Chez Vincent, Imprimeur-Libraire de Mgr le Comte de Provence, rue S, Severin, Paris, 640 pp. [https://www.biodiversitylibrary.org/item/6958#page/794/mode/1up]
  2. Andrews, H.[C.] (1798) The botanist’s repository, for new, and rare plants. Containing coloured figures of such plants, as have not hitherto appeared in any similar publication; with all their essential characters, botanically arranged, after the sexual system of the celebrated Linnaeus; in English and Latin. To each description is added, a short history of the plant, as to its time of flowering, culture, native place of growth, when introduced, and by whom. The whole executed by Henry Andrews, author of the coloured engravings of heaths, in folio. [Vereia crenata Andrews, basionym of Kalanchoe crenata (Andrews) Haw. treated in Vol. 1: t. 21 (1798).] Printed by T. Bensley, and published by the author, No 5, Knightsbridge. To be had of J. White, Fleet-street, and all the Booksellers, London, 42 plates, pages not numbered. [https://www.biodiversitylibrary.org/item/111072#page/94/mode/1up]
  3. Anonymous. (1933) Dr. Otto Stapf, F.R.S. Nature 132: 305. https://doi.org/10.1038/132305a0
  4. A.W.H. (1933) Otto Stapf—1857–1933. Obituary Notices of Fellows of the Royal Society 1: 115–118. https://doi.org/10.1098/rsbm.1933.0007
  5. Baillon, M.H. (1885) Liste des plantes de Madagascar (suite de la page 464). [Kalanchoe § Kitchingia]. Bulletin Mensuel de la Société Linnéenne de Paris 1 (59): 465–472. [https://www.biodiversitylibrary.org/item/41447#page/60/mode/1up]
  6. Baker, J.G. (1880) A synopsis of Aloineae and Yuccoideae. Journal of the Linnean Society Botany 18: 148–241. https://doi.org/10.1111/j.1095-8339.1880.tb00663.x
  7. Baker, J.G. (1881) Notes on a collection of flowering plants made by L. Kitching, Esq., in Madagascar in 1879 (Plates VII. & VIII.). Journal of the Linnean Society Botany 18: 264–281. [https://www.biodiversitylibrary.org/item/8373#page/271/mode/1up (text); https://www.biodiversitylibrary.org/item/8373#page/564/mode/1up (Plate VII, Figs 1–3)]
  8. Baker, J.G. (1882) Contributions to the flora of central Madagascar (to be continued). Journal of Botany, British and Foreign 20: 109–114. [https://www.biodiversitylibrary.org/item/35873#page/125/mode/1up]
  9. Baker, J.G. (1883) Contributions to the flora of Madagascar.—Part I. Polypetalæ [Plates XXII & XXIII]. Botanical Journal of the Linnean Society 20: 87–158. https://doi.org/10.1111/j.1095-8339.1883.tb00195.x
  10. Baker, J.G. (1884) Further contributions to the flora of central Madagascar. Botanical Journal of the Linnean Society 21: 317–353. https://doi.org/10.1111/j.1095-8339.1884.tb00548.x
  11. Baker, J.G. (1887) Further contributions to the flora of Madagascar. Botanical Journal of the Linnean Society 22: 441–537. https://doi.org/10.1111/j.1095-8339.1887.tb00472.x
  12. Baker, J.G. (1890) Further contributions to the Flora of Madagascar. (Plates L.–LIII.). Botanical Journal of the Linnean Society 25: 294–350. https://doi.org/10.1111/j.1095-8339.1890.tb02402.x
  13. Baker, J.G. (1896) Aloe. In: Thiselton-Dyer, W.T. (Ed.) Flora capensis [being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal]. Vol. 6. L. Reeve & Co., London, pp. 302–329. [https://www.biodiversitylibrary.org/item/15237#page/311/mode/1up]
  14. Berger, A. (1930) Crassulaceae. Unterfam. II. Kalanchoideae [sic, Kalanchooideae]. 6. Kalanchoe Adanson. 7. Kitchingia Baker. 8. Bryophyllum Salisb. In: Engler, A. & Prantl, K. (Eds.) Die natürlichen Pflanzenfamilien, edn 2, 18a. Verlag von Wilhelm Engelmann, Leipzig, pp. 378–386, 402–412.
  15. Berger, E. (2020) Mein Leben mit Alwin Berger. La Mortola—Stuttgart—Geneva. Im Auftrag des Alwin-Berger-Archivs herausgegeben und bearbeitet von Rainer Redies. Mit einer Bibliografie von Dr Detlev Menzing [Metzing] und einer Pflanzenliste von Dr Björn Schäfer. Eugen Ulmer, Stuttgart (Hohenheim), 256 pp.
  16. Boiteau, P. (1947) Les plantes grasses de Madagascar. [Overview of sections in Kalanchoe.] Cactus 1947 (12): 5–10, figures 1–4.
  17. Boiteau, P. & Allorge-Boiteau, L. (1995) Kalanchoe (Crassulacées) de Madagascar. Systématique, écophysiologie et phytochimie. ICSN, CNRS, 91198 Gif-sur-Yvette, Éditions Karthala, Paris, 252 pp.
  18. Boiteau, P. & Mannoni, O. (1948a) Les Kalanchoe (suite). Section I. Kitchingia. [§ 1. Sylvaticae and § 2. Campanulatae.] (à suivre). Cactus 1948 (13): 7–10, figures 1–6.
  19. Boiteau, P. & Mannoni, O. (1948b) Les Kalanchoe (suite). [§ 4. Epidendrae; Kalanchoe porphyrocalyx (Baker) Baill. and Kalanchoe uniflora Raym.-Hamet (in the key).] (à suivre). Cactus 1948 (17–18): 57–58, two unnumbered figures.
  20. Boiteau, P. & Mannoni, O. (1949) Les Kalanchoe (suite). [Kalanchoe uniflora Raym.-Hamet; § 5. Scandentes; § (unnumbered) Bulbilliferae.] (à suivre). Cactus 1949 (19): 9–14, seven unnumbered figures.
  21. Britten, J. (1920) John Gilbert Baker (1834–1920). The Journal of Botany, British and Foreign LVIII (694): 233–238. [https://www.biodiversitylibrary.org/item/33758#page/285/mode/1up]
  22. Byalt, V.V. (2008) New combinations in the genera Bryophyllum and Kalanchoë (Crassulaceae). Botanicheskii Zhurnal 93 (3): 461–465. [In Russian]
  23. Descoings, B. (2003) Kalanchoe. In: Eggli, U. (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer Verlag, Berlin, Heidelberg, New York, pp. 143–181; whole work 458 pp. + XLVIII plates.
  24. Descoings, B. (2006) Le genre Kalanchoe structure et définition. Journal de Botanique de la Société Botanique de France 33: 3–28. https://doi.org/10.3406/jobot.2006.2006
  25. Dorr, L.J. (1997) Plant collectors in Madagascar and the Comoro Islands. A biographical and bibliographical guide to individuals and groups who have collected herbarium material of algae, bryophytes, fungi, lichens, and vascular plants in Madagascar and the Comoro Islands. The Trustees, Royal Botanic Gardens, Kew, 524 pp.
  26. Eggli, U. (1992) Nomenclatural notes on two genera of Crassulaceae and a new combination. Bradleya 10: 83–84. https://doi.org/10.25223/brad.n10.1992.a6
  27. Eggli, U. (Ed.) (2003) List of abbreviations and symbols. In: Eggli, U. (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer Verlag, Berlin, Heidelberg, New York, pp. IX–X; whole work 458 pp. + XLVIII plates. https://doi.org/10.1007/978-3-642-55874-0
  28. Greuter, W., McNeill, J., Barrie, F.R., Burdet, H.-M., Demoulin, V., Filgueiras, T.S., Nicolson, D.H., Silva, P.C., Skog, J.E., Trehane, P., Turland, N.J. & Hawksworth, D.L. (2000) International Code of Botanical Nomenclature (Saint Louis Code) adopted by the Sixteenth International Botanical Congress St Louis, Missouri, U.S.A. July–August 1999. Koeltz Scientific Books, Königstein. [Regnum Vegetabile 138], 474 pp. [https://archive.bgbm.org/iapt/nomenclature/code/SaintLouis/0000St.Luistitle.htm]
  29. Hamet, R. (1907) Monographie du genre Kalanchoe. (A suivre. [To be continued]) Bulletin de l’Herbier Boissier 2me sér. 7 (11): 869–900. [https://www.biodiversitylibrary.org/item/105267#page/939/mode/1up]
  30. Hamet, R. (1908) Monographie du genre Kalanchoe (Suite et fin. [Continuation and end], with Errata on p. 17). Bulletin de l’Herbier Boissier 2me sér. 8 (1): 17–48. [https://www.biodiversitylibrary.org/item/104945#page/31/mode/1up]
  31. Hamet, R. (1910) Sur quelques Kalanchoe peu connus (Suite). Bulletin de la Société Botanique de France Quatrième series 57: 49–54. https://doi.org/10.1080/00378941.1910.10832165
  32. Haworth, A.H. (1812) Synopsis plantarum succulentarum, cum descriptionibus, synonymis, locis; observationibus anglicanis, culturaque. [Kalenchoe, sic, Kalanchoe, on pp. 109–110; Bryophyllum on p. 110.] Typis Richardi Taylor et Socii, Shoe-Lane, Londoni, 334 pp.; including ‘Appendix’. https://doi.org/10.5962/bhl.title.9462
  33. Hooker, W.J. & Baker, J.G. (1865) Synopsis filicum; or, A synopsis of all known ferns, including the Osmundaceae, Schizæsveae, Marattiaceae, and Ophioglossaceae (chiefly derived from the Kew herbarium). Robert Hardwicke, London, 523 pp.
  34. IPNI (2024+) The International Plant Names Index. Available from: https://www.ipni.org/ (accessed January 2024)
  35. Koorders, S.H. (1919) Beitrag zur Kenntniss der Flora von Java No. 18. Beschreibung von Kalanchoë Schumacheri vom Idjen-Plateau und Revision der Crassulaceae von Java. (Mit 2 lithographischen Tafeln.). Bulletin du Jardin Botanique de Buitenzorg sér. 3 1 (3): 169–180, Figures 14 and 15. [https://www.biodiversitylibrary.org/item/27173#page/205/mode/1up (text); https://www.biodiversitylibrary.org/item/27173#page/243/mode/1up (figures)]
  36. Lamarck, J. (1786) Encyclopédie méthodique. Botanique 2(1). Panckoucke, Paris & Plomteux, Liège, 774 pp. https://doi.org/10.5962/bhl.title.824
  37. Linnaeus, C. (1753) Species plantarum, exhibentes plantas rite cognitas, ad generarelatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundum systema sexuale digestas. Vol. 1. Salvius, Stockholm, 560 pp. [Aloe L. on pp. 319–323 and Cotyledon L., nom. cons., treated on pp. 429–430]. [https://www.biodiversitylibrary.org/item/13829#page/331/mode/1up [Aloe L.]; https://www.biodiversitylibrary.org/item/13829#page/441/mode/1up [Cotyledon L.]]
  38. Maire, R. (1977) Kalanchoe Adanson (1763) Rosales: Crassulaceae. In: Quézel, P.[A.] (Ed.) Flore de l’Afrique de Nord (Maroc, Algérie, Tunisie, Tripolitaine, Cyrénaïque et Sahara). Vol. XIV. Rhoeadales: Cruciferae fin Resedaceae. Sarraceniales: Droseraceae. Rosales: Crassulaceae. Éditions Paul Lechevalier éditeur S.A.R.L., 19, rue Augereau, 19, Paris, pp. 239–241, 251–252, 263. [https://archive.org/details/flore-afrique-du-nord/Vol%2014%20Flore%20de%20l%27Afrique%20Du%20Nord%20Maroc%20Algerie%20Tunisie%20Tripolitaine%20Cyrenaique%20Et%20Sahara%20R-Maire-1977/page/n119/mode/2up]
  39. Nash, G.V. (1919) Bryophyllum crenatum. Madagascar Sprouting-leaf. Native of central Madagascar. Family Crassulaceae. Orpine family. Addisonia: Colored illustrations and popular descriptions of plants 4 (4): 63, t. 152. [https://www.biodiversitylibrary.org/item/26178#page/138/mode/1up]
  40. Perrier de la Bâthie, [J.M.]H.[A.] (1923) Crassulacées malgache nouvelles. Bulletin du Muséum d’Histoire Naturelle 1923 [xxix] (6): 452–455. [https://www.biodiversitylibrary.org/item/213221#page/496/mode/1up]
  41. Perrier de la Bâthie, [J.M.]H.[A.] (1924) Les Crassulacées Malgaches. [Observations sur quelques espèces du genre Kalanchoe, pp. 21–24; Crassulacées Malgaches nouvelles, pp. 24–26; Stations et distribution géographique, pp. 26–28; Synopsis des Crassulacées Malgaches pp. 29–36.] Bulletin de l’Académie Malgache 6: 21–36. [https://www.biodiversitylibrary.org/item/50784#page/80/mode/1up]
  42. Perrier de la Bâthie, [J.M.]H.[A.] (1928) Observations nouvelles sur le genre Kalanchoe. Archives de Botanique II, Bulletin Mensuel, Caen 2 (2): 17–31.
  43. Persoon, C.H. (1805) Synopsis plantarum, seu enchiridium botanicum, complectens enumerationem systematicam specierum hucusque cognitorum. Pars prima. Apud Carol. Frid. Cramerum, Parisiis lutetiorum [Paris] et apud J.G. Cottam, Tubingæ [London], 546 pp. https://doi.org/10.5962/bhl.title.638
  44. Raymond-Hamet & Perrier de la Bâthie, [J.M.]H.[A.] (1915) Troisième contribution a l’étude des Crassulacées Malgaches. Annales du Musée Colonial de Marseille sér. 3 3: 63–117. [https://www.biodiversitylibrary.org/item/22992#page/85/mode/1up]
  45. Salisbury, R.A. (1805) The Paradisus Londinensis: or coloured figures of plants cultivated in the vicinity of the metropolis 1. Published by William Hooker, No. 6, Frith Street, city not stated, presumably London, 157 pp. & 117 plates. https://doi.org/10.5962/bhl.title.53520
  46. Schmalfuß, L. (2017) Alwin Berger: his life, his family and friends. Alwin Berger and others. The signs of German culture in the gardens and in the Riviera landscape before and after the Great War. Proceedings of a conference held on 28 October 2016 organised by Università degli Studi di Genova, Centro Servizi di Ateneo per i Giardini Botanici Hanbury, with the support of the Alwin Berger Archiv, Möschlitz, Germany. Giardini Botanici Hanbury, La Mortola, Ventimiglia, Italy.] Bollettino dei Musei e degli Istituti Biologici dell’Università di Genova 79: 9–16.
  47. Shtein, R. & Smith, G.F. (2021) A revision of the climbing kalanchoes (Crassulaceae subfam. Kalanchooideae) of Madagascar including the description of Kalanchoe sect. Invasores and K. ser. Vilana. Phytotaxa 482 (2): 93–120. https://doi.org/10.11646/phytotaxa.482.2.1
  48. Smith, G.F. (2021) Reinstatement of Kalanchoe [subg. Bryophyllum] sect. Alatae (Crassulaceae subfam. Kalanchooideae) for K. porphyrocalyx and K. uniflora, two Madagascan endemics. Bradleya 39: 177–181. https://doi.org/10.25223/brad.n39.2021.a17
  49. Smith, G.F. (2023a) The ‘woody clade’ in Kalanchoe: reinstatement and amendment of K. subg. Calophygia (Crassulaceae subfam. Kalanchooideae) published by Bernard Marie Descoings in 2006. Phytotaxa 616 (2): 113–127. https://doi.org/10.11646/phytotaxa.616.2.1
  50. Smith, G.F. (2023b) A reassessment of combinations previously proposed or published for Kalanchoe subg. Alatae (Crassulaceae subfam. Kalanchooideae) and its valid publication at the rank of subgenus. Phytotaxa 585 (1): 61–66. https://doi.org/10.11646/phytotaxa.585.1.7
  51. Smith, G.F. (2023c) A nomenclatural and taxonomic review of the names and new combination published between 1886 and 1931 in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by British botanist Nicholas Edward Brown (1849–1934). Phytotaxa 630 (4): 266–280. https://doi.org/10.11646/phytotaxa.630.4.2
  52. Smith, G.F. (2023d) A review of the infrageneric classifications proposed for Kalanchoe (Crassulaceae subfam. Kalanchooideae) from the earliest times up to 2000. Phytotaxa 619 (1): 1–38. https://doi.org/10.11646/phytotaxa.619.1.1
  53. Smith, G.F. (2024a) The unranked infrageneric names validly published in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by Alwin Berger in 1930 revisited, with publication of five names at the rank of section and one at the rank of subgenus. Phytotaxa 635 (4): 276–286. https://doi.org/10.11646/phytotaxa.635.4.2
  54. Smith, G.F. (2024b) Revisiting the nomenclature and taxonomy of the Malagasy Kalanchoe uniflora (K. subg. Alatae; Crassulaceae subfam. Kalanchooideae), which has Kitchingia uniflora as its basionym, and its homotypic and heterotypic synonyms. Phytotaxa 645 (1): 66–76. https://doi.org/10.11646/phytotaxa.645.1.6
  55. Smith, G.F. & Figueiredo, E. (2013) Succulent paradise. Twelve great gardens of the world. Struik Lifestyle, an imprint of Random House Struik (Pty) Ltd, Cape Town, 184 pp.
  56. Smith, G.F. & Figueiredo, E. (2019) Kalanchoe porphyrocalyx var. sambiranensis, a new variety in a Madagascan Kalanchoe (Crassulaceae subfam. Kalanchooideae). Phytotaxa 422 (2): 206–208. https://doi.org/10.11646/phytotaxa.422.2.8
  57. Smith, G.F. & Figueiredo, E. (2018) The infrageneric classification and nomenclature of Kalanchoe Adans. (Crassulaceae), with special reference to the southern African species. Bradleya 36: 162–172. https://doi.org/10.25223/brad.n36.2018.a10
  58. Smith, G.F. & Figueiredo, E. (2022) Clarifying the typification of the name Kalanchoe orgyalis (Crassulaceae subfam. Kalanchooideae). Phytotaxa 543 (1): 99–102. https://doi.org/10.11646/phytotaxa.543.1.11
  59. Smith, G.F. & Figueiredo, E. (2023a) Kalanchoe orgyalis. Crassulaceae subfam. Kalanchooideae. Flowering Plants of Africa 68: 92–105, t. 2389.
  60. Smith, G.F. & Figueiredo, E. (2023b) A review and analysis of the use of epithets in infrageneric, species, and infraspecific names in Kalanchoe (Crassulaceae subfam. Kalanchooideae): trends in the plant naming game. Phytotaxa 618 (3): 213–242. https://doi.org/10.11646/phytotaxa.618.3.1
  61. Smith, G.F. & Figueiredo, E. (2023c) Names validly published in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by Louis-René ‘Edmond’ Tulasne in 1857, with reference to names proposed in sched. by Hilsenberg and Bojer for two Malagasy species, and with notes on the type of the name K. tubiflora. Phytotaxa 584 (4): 275–284. https://doi.org/10.11646/phytotaxa.584.4.4
  62. Smith, G.F., Figueiredo, E. & Van Wyk, A.E. (2019) Kalanchoe (Crassulaceae) in southern Africa. Classification, biology, and cultivation. Academic Press, an imprint of Elsevier, London, San Diego, Cambridge (U.S.A.), and Oxford, 328 pp. https://doi.org/10.1016/C2017-0-00602-X
  63. Smith, G.F., Klein, D.-P. & Shtein, R. (2021b) A revision of the Malagasy Kalanchoe subg. Kitchingia (Crassulaceae subfam. Kalanchooideae): history, taxonomy, and nomenclature. Phytotaxa 507 (1): 67–80. https://doi.org/10.11646/phytotaxa.507.1.3
  64. Smith, G.F. & Monro, A.M. (2022a) The nomenclature and taxonomy of Crassulaceae subfam. Kalanchooideae resolved. Phytotaxa 555 (3): 241–251. https://doi.org/10.11646/phytotaxa.555.3.3
  65. Smith, G.F. & Monro, A.M. (2022b) (2908) Proposal to reject the name Crassulaceae subfam. Cotyledonoideae (Crassulaceae). Taxon 71 (4): 910–911. https://doi.org/10.1002/tax.12778
  66. Smith, G.F. & Monro, A.M. (2022c) Notes on the nomenclature and taxonomy of Chiastophyllum and Umbilicus oppositifolius (≡Cotyledon oppositifolia, ≡Chiastophyllum oppositifolium) (Crassulaceae subfam. Sempervivoideae). Phytotaxa 564 (2): 239–247. https://doi.org/10.11646/phytotaxa.564.2.7
  67. Smith, G.F. & Shtein, R. (2021) Nomenclature and taxonomic placement of the infraspecific taxa recognised in the Madagascan Kalanchoe miniata (Crassulaceae subfam. Kalanchooideae) by Perrier de la Bâthie and Boiteau and Mannoni in the 20th century. Phytotaxa 514 (2): 118–128. https://doi.org/10.11646/phytotaxa.514.2.3
  68. Smith, G.F., Shtein, R. & Klein, D.-P. (2021c) Kalanchoe subg. Alatae (Crassulaceae subfam. Kalanchooideae), a new rank for K. sect. Alatae. Phytotaxa 509 (2): 255–258. https://doi.org/10.11646/phytotaxa.509.2.9
  69. Smith, G.F., Shtein, R., Klein, D.-P., Parihar, B., Almeida, A., Rodewald, S. & Kadereit, G. (2021a) Special Review. Sexual and asexual reproduction in Kalanchoe (Crassulaceae subfam. Kalanchooideae): a review of known and newly recorded strategies. Haseltonia 28: 2–22. https://doi.org/10.2985/026.028.0101
  70. Stapf, O. (1908) Diagnoses africanae: XXIII. Bulletin of Miscellaneous Information 1908 (6): 257–262. https://doi.org/10.2307/4111575
  71. Stapf, O. (1909) Kitchingia uniflora. Madagascar. Crassulaceae. Curtis’s Botanical Magazine 135 4th ser. 5: Tab. 8286. [https://www.biodiversitylibrary.org/item/14261#page/227/mode/1up]
  72. Thiers, B. (2024-continuously updated) Index herbariorum: a global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (accessed January 2024)
  73. Tulasne, L.-R. [‘Edmond’] (1857) Fragmentum alterum (1). Floræ madagascariensis. II. Calanchoe [sic, Kalanchoe] pp. 149–150. Annales des Sciences Naturelles, Botanique sér. 4, 8: 44–163. [https://www.biodiversitylibrary.org/item/129817#page/153/mode/1up]
  74. Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Koeltz Botanical Books, Glashütten, 254 pp. https://doi.org/10.12705/Code.2018