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Issue: Vol. 626 No. 4: 28 Nov. 2023
Type: Article
Published: 2023-11-28
DOI: 10.11646/phytotaxa.626.4.1
Page range: 223-246
Abstract views: 110
PDF downloaded: 15

A critical inventory of the family Amaranthaceae s.str. in Saudi Arabia

Botany and Microbiology Department; Faculty of Science; Beni-Suef University; Beni-Suef; Egypt
Department of Biology; College of Science; Princess Nourah bint Abdulrahman University; Riyadh; Saudi Arabia
Department of Environmental Biology; Univeristy of Rome Sapienza; 00185 Rome; Italy
Aliens Asia flora morphology new record typification Eudicots

Abstract

The aims of the present study are to assess and update the floristic diversity of Amaranthaceae s.str. in Saudi Arabia. A total number of 33 taxa were indentified including 28 species, 4 subspecies, and 7 varieties. The richest genus is Amaranthus with 16 taxa (12 species); Aerva and Alternanthera have 3 species each; Achyranthes and Celosia 2 species each; 6 genera (Digera, Gomphrena, Nothosaerva, Psilotrichum, Pupalia, Saltia) include just one species each. Most of the taxa are native (18, 54.55%), 14 (42.42%) are aliens (6 casual taxa, 5 naturalized, and 1 invasive), one (Alternanthera sessilis) is here considered as doubtfully native, two (Amaranthus tricolor, Gomphrena globosa) are found only in cultivation. Two species (Amaranthus tortuosum, Pupalia lappacea var. lappacea) are excluded from the national flora. Diagnostic keys at genus, species, and infraspecific ranks are provided. The following names are lectotypified: Achyranthes argentea var. obovata (G; isolectotype at HBG), Achyranthes fruticosa var. pubescens (G), Achyranthes papposa (C; isolectotypes at C and BM), Achyranthes polystachya (BM), Aerva javanica var. bovei (FI), Achyranthes lanata (illustration by Burman), Digera arvensis (C; isolectotype at C), Psilostachys gnaphalobryum (TUB; isolectotypes at GOET, HBG, HEID, JE, MEL, MPU, P, and TUB), Telanthera bettzickiana (LE).

 

References

  1. Abbas, A.M., Al-Kahtani, M.A., Alfaifi, M.Y., Elbehairi, S.E.I. & Badry, M.O. (2020) Floristic Diversity and Phytogeography of JABAL Fayfa: A Subtropical Dry Zone, South-West Saudi Arabia. Diversity 12 (9): 345. https://doi.org/10.3390/d12090345
  2. Abdullah, M.A., Al.Yasee, H., Al-Sudani, Y. & Moneruzaman, M. (2017) Developing weeds of agricultural crops at different levels of heights, in taif area of Saudia Arabia. Bulgarian Journal of Agricultural Science 23 (5): 762–769.
  3. Al-Eisawi, D.M. & Al-Ruzayza, S. (2015) The flora of holy Mecca district, Saudi Arabia. International Journal of Biodiversity Conservation 7 (3): 173–189. https://doi.org/10.5897/IJBC2014.0773
  4. Alfarhan, A.H., Thomas, J. & Rajakrishan, R. (2021) Impact Assessment and Management of Invasive Species in Plant Diversity Centers and Agriculture Fields of Saudi Arabia In: Pullaiah, T. & Ielmini, M.R. (Eds.) Invasive Alien Species: Observations and Issues from Around the World, vol. 2. Wiley-Blackwell, pp. 207–225. https://doi.org/10.1002/9781119607045.ch22
  5. Al-Namazi, Ali, A., Algarni, S.M., Justin, S.H. Wan, Al Mosallam, M.S. & Alotaibi, F. (2022) Floristic composition of Jandaf Mountain as biodiversity hotspot area in southwestern Saudi Arabia. Saudi Journal of Biological Science 29 (5): 3654–3660. https://doi.org/10.1016/j.sjbs.2022.03.003
  6. Al-Turki, T.A., Filfilan, S.A. & Mehmood, S.F. (2000) A cytological study of flowering plants from Saudi Arabia. Willdenowia 30: 339–358. https://doi.org/10.3372/wi.30.30211
  7. Alwadie, H.M. (2005) Morphology and Distribution of Three Genera of Amaranthaceae in the South Western Area of Saudi Arabia. Journal of King Saud University 18 (1): 51–62.
  8. Aljieddani, G.S., Al-Harbi, N.A., Al-Qathani, S.M., Al-Absy, K.M., Abdullatif, B.M. & Dahan, T.N. (2021) Inventory of some introduced and invasive plant species in some governorates of the Kingdom of Saudi Arabia. Applied Ecology and Environmental Research 19 (6): 4373–4388. https://doi.org/10.15666/aeer/1906_43734388
  9. APG IV [Angiosperm Phylogeny Group IV] (2016) An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV. Botanical Journal of Linnean Society 181 (1): 1–20. https://doi.org/10.1111/boj.12385
  10. Atiqur Rahman, M., Al-Said, M.S., Mossa, J.S., Al-Yahya, M.A. & Fahad Al-Hemaid, M.A. (2002) A Check List of Angiosperm Flora of Farasan Islands, Kingdom of Saudi Arabia. Pakistan Journal of Biological Science 5 (11): 1162–1166. https://doi.org/10.3923/pjbs.2002.1162.1166
  11. Bao, B., Clemants, S.E. & Borsch, T. (2003) Psilotrichum Blume. In: Wu, Z.Y., Raven, P.H. & Hong, D.Y. (Eds.) Flora of China, vol. 5. Science Press, Beijing and Missouri Botanical Garden Press, St. Louis, pp. 423.
  12. Barrie, F.R. (2006) Report of the General Committee: 9. Taxon 55 (3): 795–800. https://doi.org/10.2307/25065657
  13. Bauhin, J. & Cherler, J.H. (1651) Historia Plantarum Universalis, vol. 2. Fr. Lud. Graffenried, Ebroduni, 1074 pp.
  14. Blume, C.L. von. (1826) Bijdragen tot de Flora van Nederlandsch Indie, parts 10–13. Ter Lands Drukkerij, Batavia, 245 pp. https://doi.org/10.5962/bhl.title.115427
  15. Borsch, T., Berendsohn, W., Dalcin, E., Delmas, M., Demissew, S., Elliott, A., Fritsch, P., Fuchs, A., Geltman, D., Güner, A., Haevermans, T., Knapp, S., Le Roux, M. M., Loizeau, P.-A., Miller, C., Miller, J., Miller, J. T., Palese, R., Paton, A., Parnell, J., Pendry, C., Qin, H.-N., Sosa, V., Sosef, M., Von Raab-Straube, E., Ranwashe, F., Raz, L., Salimov, R., Smets, E., Thiers, B., Thomas, W., Tulig, M., Ulate, W., Ung, V., Watson, M., Wyse Jackson, P. & Zamora, N. (2020) World Flora Online: Placing taxonomists at the heart of a definitive and comprehensive global resource on the world’s plants. Taxon 69 (4): 1311–1341. https://doi.org/10.1002/tax.12373
  16. Burman, J. (1737) Thesaurus zeylanicus. Amsterdam: apud Janssonio-Waesbergios & Salomonem Schouten. https://doi.org/10.5962/bhl.title.734
  17. Burman, N.L. (1768) Flora Indica. Lugduni Batavorum [Leiden]: apud Cornelium Haek; Amstelaedami [Amsterdam]: apud Johannem Schreuderum.
  18. Chaudhary, S.A. (1998) Flora of the Kingdom of the Saudi Arabia, vol. 1. Ministry of Agriculture & Water, Riyadh, 692 pp.
  19. Chaudhary, S.A., Parker, C. & Kasasian, L. (1981) Weeds of central, southern and eastern Arabian Peninsula. Tropical Pest Management 27 (2): 181–190. https://doi.org/10.1080/09670878109413649
  20. Clemants, S.E. (2003) Alternanthera Forssk. In: Flora of North America Editorial Committee (Eds.) Flora of North America North Mexico (Magnoliophyta: Caryophyllydae, part 1), 4. Oxford University Press, Oxford, pp. 447–451.
  21. Collenette, S. (1999) Wildflowers of Saudi Arabia. National commission for wildlife conservation and development (NCWCD), 799 pp.
  22. Costea, M., Sanders, A. & Waines, G. (2001b) Notes on some little known Amaranthus taxa (Amaranthaceae) in the United States. Sida 19 (4): 975–992.
  23. Das, S. (2013) Infraspecific variability of Amaranthus tricolor (Amaranthaceae) in India with a new variety. Phytotaxa 88 (2): 25–30. https://doi.org/10.11646/phytotaxa.88.2.2
  24. Das, S. (2016) Amaranthus: A Promising Crop of Future. Springer, Singapore, 208 pp. https://doi.org/10.1007/978-981-10-1469-7
  25. Dodoens, R. (1616) Stirpium Historiae pemptades sex, sive libri XXX. Ex Officina Plantiniana, Antuerpiae, 872 pp.
  26. El-Ghazali, G.E.B. & Al-Soqeer, A.R.A. (2013) A Checklist of the Weed Flora of Qassim Region, Saudi Arabia. Australian Journal of Basic and Applied Science 7 (2): 900–905.
  27. El Hadidi, N. & El Hadidy, A.M.H. (1981) Flora of Egypt. Amaranthaceae Juss. Taeckholmia, Additional Series 1 (2): 1–8.
  28. Fawcett, W. & Rendle, A.B. (1914) Flora of Jamaica containing descriptions of the flowering plants known from the island, vol. 3. British Museum, London, 208 pp.
  29. Fernald, M.L. (1945) Botanical specialities of the Seward Forest and adjacent Areas of Southeastern Virginia. Rhodora 47: 93–142.
  30. Filias, F., Gaulliez, R. & Guedes, M. (1980) Amaranthus blitum vs. A. lividus (Amaranthaceae). Taxon 29: 149–150. https://doi.org/10.2307/1219612
  31. Forsskål, P. (1775) Flora Aegyptiaco-Arabica: Sive Descriptiones Plantarum, quas per Aegyptum Inferiorem et Arabiam felicem detexit, iIllustravit Petrus Forskål. Möller, Kjǿbenhavn, 220 pp. https://doi.org/10.5962/bhl.title.6625
  32. Ghazanfar, S.A. (2023 onwards) Plants of the Middle East. Flora of Saudi Arabia. Checklist of the Flowering plants of Saudi Arabia (excluding Poaceae). Available from: https://www.plantsofthemiddleeast.com/checklists/checklist-saudiarabia.htm (accessed: 12 September 2023).
  33. Ghazanfar, S.A. & Fisher, M. (2013) Vegetation of the Arabian Peninsula. Kluwer Academic Publisher, Dordrecht/Boston/London, 363 pp.
  34. Green, M.L. (1929) Amaranthus caudatus L. In: Hitchcock, A.S. (Ed.) Nomenclature. Proposals by British Botanists. Wyman & Sons, London, p. 188.
  35. Govaerts, R, Lughadha, E.N., Black, N., Turner, N. & Paton, A. (2021) The World Checklist of Vascular Plants, a continuously updated resource for exploring global plant diversity. Scientific Data 8: 215. https://doi.org/10.1038/s41597-021-00997-6
  36. Hall, M., Llwellyn, O.A., Miller, A.G., Al-Abbasi, T.M., Al-Wetaid, A.H., Al-Harbi, L.J. & Al-Shammari, K.F. (2022) Important plant areas in the Arabian Peninsula: 2. Farasan Archipelago. Edinburgh Journal of Botany 67 (2): 189–208. https://doi.org/10.1017/S0960428610000089
  37. Hassan, A., Al-Shaye, N., Alghamdo, S., Shereen, Korany, M. & Iamonico, D. (2022) Taxonomic revision of the genus Amaranthus (Amaranthaceae) in Saudi Arabia. Phytotaxa 576 (2): 135–157. https://doi.org/10.11646/phytotaxa.576.2.1
  38. Hepper, F.N. & Friis, I. (1994) The Plants of Pehr Forsskål’s Flora Aegyptiaco-Arabica. Collected on the Royal Danish Expedition to Egypt and Yemen 1761-63. Royal Botanic Gardens, Kew & Copenhagen, 400 pp.
  39. Hernández-Ledesma, P., Berendsohn, W.G., Borsch, T., Von Mering, S., Akhani, H., Arias, S., Castañeda-Noa, I., Eggli, U., Eriksson, R., Flores-Olvera, H., Fuentes-Bazán, S., Kadereit, G., Klak, C., Korotkova, N., Nyffeler, R., Ocampo, G., Ochoterena, H., Oxelman, B., Rabeler, R.K., Sanchez, A., Schlumpberger, B.O. & Uotila, P. (2015) A taxonomic backbone for the global synthesis of species diversity in the angiosperm order Caryophyllales. Willdenowia 45 (3): 281–383. https://doi.org/10.3372/wi.45.45301
  40. Hitchcock, A.S. (1929) Nomenclature. Proposals by British Botanists. Wyman & Sons, London, pp. 188.
  41. Hochstetter, C.F. (1844) Nova genera plantarum Africae, proponit et describit. Continuatio. Flora 27: 1–8.
  42. Hornemann, J.W. (1819) Hortus regius botanicus Hafniensis in usum tyronum et botanophilorum. Supplementum. E. A. H. Mölleri, Hafinae, 172 pp.
  43. HUH Index of Botanists (2013a-onwards) Index of botanists, Harward University Herbaria & Libraries. Regel, Eduard August von. Available from: https://kiki.huh.harvard.edu/databases/botanist_search.php?mode=details&id=333 (accessed: 12 September 2023).
  44. HUH Index of Botanists (2013b-onwards) Index of botanists, Harward University Herbaria & Libraries. Forsskål, Pehr. Available from: https://kiki.huh.harvard.edu/databases/botanist_search.php?mode=details&id=1609 (accessed: 12 September 2023).
  45. HUH Index of Botanists (2013c-onwards) Index of botanists, Harward University Herbaria & Libraries. Wallich, Nathaniel. Available from: https://kiki.huh.harvard.edu/databases/botanist_search.php?mode=details&id=776 (accessed: 12 September 2023).
  46. Iamonico, D. (2011) On the presence of Amaranthus polygonoides L. (Amaranthaceae) in Europe. Phyton 50 (2): 205–219.
  47. Iamonico, D. (2014a) Lectotypification of Linnaean names in the genus Achyranthes L. (Amaranthaceae). Taxon 63 (2): 405–407. https://doi.org/10.12705/632.2
  48. Iamonico, D. (2014b) Lectotypification of Linnaean names in the genus Amaranthus L. (Amaranthaceae). Taxon 63: 146–150. https://doi.org/10.12705/631.34
  49. Iamonico, D. (2015a) Taxonomic revision of the genus Amaranthus (Amaranthaceae) in Italy. Phytotaxa 199 (1): 1–84. https://doi.org/10.11646/phytotaxa.199.1.1
  50. Iamonico, D. (2015b) Amaranthaceae Juss. In: Euro+Med Plantbase – the information resource for Euro-Mediterranean plant diversity. Available from https://www.europlusmed.org/cdm_dataportal/taxon/3bff5e2c-a0ab-458d-804b-5ee334c024d5 (accessed: 12 September 2023).
  51. Iamonico, D. (2016a) Nomenclature survey of the genus Amaranthus (Amaranthaceae). 3. Names linked to the Italian flora. Plant Biosystems 150 (3): 519–531. https://doi.org/10.1080/11263504.2014.987188
  52. Iamonico, D. (2016b) Nomenclature survey of the genus Amaranthus (Amaranthaceae). 4. Detailed questions arising around the name Amaranthus gracilis. Botanica Serbica 40 (1): 61–68.
  53. Iamonico, D. (2016c) Nomenclature survey of the genus Amaranthus (Amaranthaceae). 5. Moquin-Tandon’s names. Phytotaxa 273 (2): 81–114. https://doi.org/10.11646/phytotaxa.273.2.1
  54. Iamonico, D. (2017) Achyranthes L. In: Pignatti, S. (Ed.) Flora d’Italia, vol. 2. Ed agricole, Milano, pp. 233–234.
  55. Iamonico, D. (2020) A nomenclature survey of the genus Amaranthus (Amaranthaceae). 7. Wildenow’s names. Willdenowia 50 (1): 147–155. https://doi.org/10.3372/wi.50.50114
  56. Iamonico, D. (2023) Nomenclature Survey of the Genus Amaranthus (Amaranthaceae): 12. Questions about Amaranthus caudatus. Plants 12: 1566. https://doi.org/10.3390/plants12071566
  57. Iamonico, D. & Das, S. (2014) Amaranthus bengalense (Amaranthaceae) a new species from India, with taxonomical notes on A. blitum aggregate. Phytotaxa 181 (5): 293–300. https://doi.org/10.11646/phytotaxa.181.5.4
  58. Iamonico, D. & El Mokni, R. (2017) Amaranthus palmeri (Amaranthaceae) in Tunisia, a second record for the continental African flora and nomenclatural notes on A. sonoriensis nom. nov. pro A. palmeri var. glomeratus. Bothalia 47 (1): a2100. https://doi.org/10.4102/abc.v47i1.2100
  59. Iamonico, D. & El Mokni, R. (2019) A new addition to the alien flora of Tunisia, Amaranthus spinosus L. (Amaranthaceae s.l.), with notes on A. diacanthus Raf.. Acta Botanica Croatica 78 (1): 91–94. https://doi.org/10.2478/botcro-2018-0009
  60. Iamonico, D. & Friis, I. (2017) The lectotypification of two names referred by Linnaeus to the genus Illecebrum (Caryophyllales): Illecebrum benghalense and Iresine javanica, with comments on Aerva tomentosa Forssk. Taxon 66 (5): 1208–1212. https://doi.org/10.12705/665.14
  61. Iamonico, D. & Palmer, J. (2020) Nomenclature survey of the genus Amaranthus (Amaranthaceae). 6. Names linked to the Australian flora. Australian Systematic Botany 33 (2): 169–173. https://doi.org/10.1071/SB18062
  62. Iamonico, D. & Sánchez Del Pino, I. (2016) Taxonomic revision of the genus Alternanthera (Amaranthaceae) in Italy. Plant Biosystems 150 (2): 333–342. https://doi.org/10.1080/11263504.2015.1019588
  63. Jarvis, C.E. (1992) Seventy-Two Proposals for the Conservation of Types of Selected Linnaean Generic Names, the Report of Subcommittee 3C on the Lectotypification of Linnaean Generic Names. Taxon 41 (3): 552–583. https://doi.org/10.2307/1222833
  64. Jussieu, A.L. de (1803) Observations sur la fimille des plantes amarantacées. Annales du Museum National d’Histoire Naturelle. Paris 2: 131–135.
  65. IPNI (2023) The International Plant Names Index. Psilotrichum cordatum Moq. Available from https://www.ipni.org/n/61151-1 (accessed: 12 September 2023).
  66. Kadereit, G., Borsch, T., Weising, K. & Freitag, H. (2003) Phylogeny of Amaranthaceae and Chenopodiaceae and the evolution of C4 photosynthesis. International Journal of Plant Sciences 164 (6): 959–986. https://doi.org/10.1086/378649
  67. Lambdon, P.W., Pyšek, P., Basnou, C., Hejda, M., Arianoutsou, M., Essl, F., Jarošík, V., Pergl, J., Winter, M., Anastasiu, P., Andriopoulos, P., Bazos, I., Brundu, G., Celesti-Grapow, L., Chassot, P., Delipetrou, P., Josefsson, M., Kark, S., Klotz, S., Kokkoris, Y., Kuehn, I., Marchante, H., Perglová, I., Pino, J., Vila, M., Zikos, A., Roy, D. & Hulme, P.E. (2008) Alien flora of Europe: species diversity, temporal trends, geographical patterns and research needs. Preslia 80: 101–149.
  68. Linnaeus, C. (1753) Species Plantarum, vol. 1. Laurentius Salvius, Stockholm, 590 pp.
  69. Lorite, J (2016) An updated checklist of the vascular flora of Sierra Nevada (SE Spain). Phytotaxa 261 (1): 1–57. https://doi.org/10.11646/17094
  70. Mears, A.J. (1980) The Linnaean species of Gomphrena L. (Amaranthaceae). Taxon 29 (1): 85–95. https://doi.org/10.2307/1219601
  71. Melville, R. (1958) Notes on Alternanthera. Kew Bulletin 13 (1): 171–175. https://doi.org/10.2307/4117643
  72. Miller, A.G. & Cope, T.A. (1996) Flora of the Arabian peninsula and Socotra, vol. 1. Edinburgh university press, Edinburgh, 586 pp.
  73. Moquin-Tandon, C.H.B.A. (1849) Amaranthaceae Juss. In: Candolle, A.P. De (Ed.) Prodromus systematics naturalis regni vegetabilis, vol. 13, part. 2. Treuttel & Wurtz, Paris, pp. 231–424.
  74. Mosyakin, S.L. & Robertson, K.R. (2003) Amaranthus L. In: Flora of North America Editorial Committee (Eds.) Flora of North America North of Mexico (Magnoliophyta: Caryophyllidae, part 1) 4. Oxford University Press, Oxford, pp. 410–435.
  75. Nobis, M., Klichowska, E., Wolanin, M., Nobis, A. & Nowak, A. (2022) Typification of five plant names described based on specimens collected by Józef Warszewicz in Central and South America. PhytoKeys 192: 45–61. https://doi.org/10.3897/phytokeys.192.78409
  76. Osman, A.K.E. & El-Ameid Abedin, M.A. (2019) Floristic diversity of Wadi Ar'ar, Saudi Arabia. Journal of Taibah University for Science 13 (1): 772–789. https://doi.org/10.1080/16583655.2019.1634177
  77. Patzelt, A. & al. (2020) Studies in the flora of Arabia: XXXIV. Sixty new records from the Sultanate of Oman. Edinburgh Journal of Botany 77: 413–437. https://doi.org/10.1017/S0960428620000086
  78. Peruzzi, L., Domina, G., Bartolucci, F., Galasso, G., Peccenini, S., Raimondo, F.M., Albano, A., Alessandrini, A., Banfi, E., Barberis, G., Bernardo, L., Bovio,M., Brullo,S., Brundu,G., Brunu, A., Camarda, I., Carta, L., Conti, F., Croce, A., Iamonico,D., Iberite, M., Iriti, G., Longo, D., Marsili, S., Medagli, P., Pistarino, A., Salmeri, C., Santangelo, A., Scassellati, E., Selvi, F., Soldano, A., Stinca, A., Villani, M., Wagensommer, R.P. & Passalacqua, N.G. (2015) An inventory of the names of vascular plants endemic to Italy, their loci classici and types. Phytotaxa 196 (1): 1–217. https://doi.org/10.11646/phytotaxa.196.1.1
  79. POWO (2023) Plants of the World Online. Available from: https://powo.science.kew.org/ (accessed: 12 September 2023).
  80. Pyšek, P., Richardson, D.M., Rejmánek, M., Webster, G.L., Williamson, M. & Kirschner, J. (2002) Alien plants in checklists and floras: towards better communication between taxonomists and ecologists. Taxon 53: 131–143. https://doi.org/10.2307/4135498
  81. Raus, Th. (1997) Amaranthus L. In: Strid, A. & Tan, K. (Eds.) Flora Hellenica, vol. 1. Koeltz Scientific Books, Königstein, pp. 138–146.
  82. Raus, Th. (2022) Taxonomic, nomenclatural and floristic review of Amaranthaceae of Greece and neighbouring countries. Willdenowia 52: 335–357. https://doi.org/10.3372/wi.52.52304
  83. Regel, E.A. von (1862a) Index Seminum, quae Hortus Botanicus Imperialis Petropolitanus pro Mutua Commutatione Offert. Accedunt Animadversiones Botanicae Nonnullae. St. Petersburg.
  84. Regel, E.A. von (1862b) Gartenflora, vol. 11. Ferdinand Enke, Erlangen, 448 pp.
  85. Reveal, J.L. & Jarvis, C.E. (2009) Typification of names of temperate North American plants proposed by Linnaeus. Taxon 58: 977–984. https://doi.org/10.1002/tax.583027
  86. Richardson, D.M. & Pyšek, P. (2006) Plant invasions: merging the concepts of species invasiveness and community invasibility. Progress in Physical Geography 30 (3): 409–431. https://doi.org/10.1191/0309133306pp490pr
  87. Robertson, K.R. (2003) Achyranthes L. In: Flora of North America Editorial Committee (eds.) Flora of North America north of Mexico 4. Oxford University Press, New York & Oxford, pp. 435–437.
  88. Robertson, K.R. & Clemants., S.E. (2003) Amaranthaceae Juss. In: North American Editorial Committee (Eds.) Flora of North America: Magnoliophyta: Caryophyllidae, vol. 1 (4). Oxford University Press, Oxford, pp. 405–456.
  89. Sánchez-del Pino, I. & Iamonico, D. (2016) Jamesbondia, a new Subgenus of Alternanthera (Gomphrenoideae, Amaranthaceae) from Central America and the Caribbean Islands. Plant Biosystems 150 (2): 190–200. https://doi.org/10.1080/11263504.2014.941034
  90. Sindhu, A., Venugopalan Nair Saradamma, A.K., Walsan Kalarikkal, V. & Iamonico, D. (2020) Amaranthus rajasekharii (Amaranthaceae), a new species from Kerala (SW-India). Phytotaxa 433 (2): 153–160. https://doi.org/10.11646/phytotaxa.433.2.6
  91. Sindhu, A., Iamonico, D., Sánchez-Del Pino, I. & Venugopalan Nair Saradamma, A.K. (2021a) Alternanthera indica (Amaranthaceae), a new species from Kerala (India). Phytotaxa 482 (2): 191–196. https://doi.org/10.11646/phytotaxa.482.2.7
  92. Sindhu, A., Iamonico, D. & Venugopalan Nair Saradamma, A.K. (2021b) Amaranthus powellii (Amaranthaceae), a new addition for the flora of India and a preliminary list of the indian Amaranthus species. Hacquetia 20 (2): 257–262. https://doi.org/10.2478/hacq-2021-0005
  93. Thiers, B. (2023 [continuously updated]) Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden's Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (accessed: 12 September 2023).
  94. Thiv, M., Thulin, M., Kilian, N. & Linder, P. (2006) Eritreo-Arabian Affinities of the Socotran Flora as Revealed from the Molecular Phylogeny of Aerva (Amaranthaceae). Systematic Botany 31 (3): 560–570. https://doi.org/10.1600/036364406778388683
  95. Thomas, H. (2011+) Flora of Saudi Arabia – Checklist. Available from: http://www.plantdiversityofsaudiarabia.info/Biodiversity-Saudi-Arabia/Flora/Checklist/Cheklist.htm (accessed: 12 September 2023).
  96. Townsend, C.C. (1973) Notes on Amaranthaceae—I. Kew Bulletin 28 (1): 141–146. https://doi.org/10.2307/4117075
  97. Townsend, C.C. (1974) Amaranthaceae. In: Nasir, E. & Ali, S.I. (Eds.) Flora of West Pakistan, vol. 71. Ferozsons Press, Rawalpindi, pp. 1–49.
  98. Townsend, C.C. (1979) A Survey of Pupalia Juss. Notes on Amaranthaceae: V. Kew Bulletin 34 (1): 131–142. https://doi.org/10.2307/4117978
  99. Townsend, C.C. (1980) A New Psilotrichum from Somalia with a Note on Centemopsis: Notes on Amaranthaceae: XI. Kew Bulletin 35 (2): 377–378. https://doi.org/10.2307/4114586
  100. Townsend, C.C. (1985) Amaranthaceae Juss. In: Polhill, R.M. (Ed.) Flora of Tropical East Africa. A.A. Balkema, Rotterdam, pp. 1–136.
  101. Townsend, C.C. (1993) Amaranthaceae Juss. In: Kubitzki, K., Rohwer, J.G. & Bittrich, V. (Eds.) The families and genera of vascular plants, vol. 2. Springer, Berlin, Heidelberg & New York, pp. 70–91.
  102. Townsend, C.C. (1994) Amaranthacées. In: Bosser, J., Cadet, T., Guého, J. & Marais, W. (Eds.) Flore des Mascareignes: La Réunion, Maurice, Rodrigues, vol. 142. Royal Botanic Gardens, Kew, pp. 1–32.
  103. Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress, Shenzhen, China, July 2017. Regnum Vegetabile 159: i–xxxviii + 1–254. https://doi.org/10.12705/Code.2018
  104. Tutin, T.G. (1993) Achyranthes L. In: Tutin, T.G., Burges, N.A., Chater, A.O, Edmondson, J.R., Heywood, V.H., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (Eds.) Flora Europaea (second edition), vol. 1. Cambridge University Press, Cambridge, pp. 132.
  105. Wagensommer, R.P. (2023) Floristic studies in the light of biodiversity knowledge and conservation. Plants 12 (16): 2973.
  106. https://doi.org/10.3390/plants12162973
  107. Wagensommer, R.P., Perrino, E.V., Albano, A., Medagli, P. & Passalacqua, N.G. (2016) Lectotypification of four Lacaita’s names in the genus Centaurea (Asteraceae). Phytotaxa 269 (1): 54–58.
  108. https://doi.org/10.11646/phytotaxa.269.1.7
  109. Wallich, N. (1829) Numerical list of dried specimens of plants in the Museum of the Holl. East India Company /which have been supplied by Dr. Wallich, superintendent of the botanic garden at Calcutta. London, 306 pp. https://doi.org/10.5962/bhl.title.1917
  110. Webb, P.B. (1849) Niger flora; or, An enumeration of the plants of western tropical Africa. H. Bailliere, London, 588 pp. + 50 illustrations.
  111. Weinmann, F., Zika, P.F., Giblin, D.E. & Legler, B. (2002 onwards) Checklist of the Vascular Plants of Washington State. University of Washington Herbarium. Available from: http://biology.burke.washington.edu/herbarium/waflora/checklist.php (accessed: 18 May 2023).
  112. Wigth, R. (1853) Icones plantarum Indiae Orientalis :or figures of Indian plants, vol. 6. Printed for the author, Madras, 44 pp. + plates 1776–2101. https://doi.org/10.5962/bhl.title.104737
  113. Zhang, Z., Idrees, M., Wang, F., Zhang, N., Chen, W., Ahmad, M., Sultana, S., Jiao, Y., Zheng, X., Li, M., Prodhan, Z.H. & Arfan, M. (2023) Typification and Nomenclature Notes on Twenty-Nine Names in Asparagus (Asparagaceae). Plants 12: 2537. https://doi.org/10.3390/plants12132537