Abstract
Acrobolbus mashpianus, a leafy liverwort of the family Acrobolbaceae, is described as new to science from the Chocó Biogeographical Region. It is characterized by strongly asymmetrical, deeply bilobed leaves with ciliate marginal teeth. The new species is illustrated and its relationships within the genus Acrobolbus are discussed. A key to the Ecuadorian species of Acrobolbus is provided.
References
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<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Briscoe, L., Engel, J.J., Söderström, L., Hagborg, A. & von Konrat, M. (2015) Notes on Early Land Plants Today. 66. Nomenclatural notes on Acrobolbaceae. <em>Phytotaxa</em> 202 (1): 58–62. https://doi.org/10.11646/phytotaxa.202.1.8</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Briscoe, L., Zerega, N., Lumbsch, T., Stech, M., Kraichak, E., von Konrat, M., Engel, J.J. & Wickett, N. (2017) Molecular, morphological, and biogeographic perspectives on the classification of Acrobolboideae (Acrobolbaceae, Marchantiophyta). <em>Phytotaxa</em> 319: 56–70. https://doi.org/10.11646/phytotaxa.319.1.2</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Burghardt, M. (2020) A first insight into the bryophyte flora of the Mashpi Ecological Reserve, Pichincha, Ecuador—Notes on the bryophytes of Ecuador V. <em>Nova Hedwigia</em> 111 (1–2): 59–76. https://doi.org/10.1127/nova_hedwigia/2020/0583</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Burghardt, M. & Gradstein, S.R. (2008) A revision of <em>Tylimanthus</em> (Acrobolbaceae, Marchantiophyta) in Tropical America, Africa, and Macaronesia. <em>Fieldiana: Botany</em> (n.ser.) 47: 199–210. https://doi.org/10.3158/0015-0746-47.1.199</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Engel, J.J. & Glenny, D.S. (2008) Austral hepaticae. 44. A revision of <em>Marsupidium</em> Mitt. for New Zealand. <em>Nova Hedwigia</em> 87 (3/4): 277–313. https://doi.org/10.1127/0029-5035/2008/0087-0277</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Engel, J.J. & Grolle, R. (1971) <em>Marsupidium</em> in South America. <em>Journal of the Hattori Botanical Laboratory</em> 34: 437–444.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Evans, A.W. (1891) A provisional list of the hepaticae of the Hawaiian Islands. <em>Transactions of the Connecticut Academy of Arts and Sciences</em> 8 (15): 253–261.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Evans, A.W. (1904) Hepaticae of Puerto Rico. IV. <em>Odontolejeunea</em>, <em>Cyclolejeunea</em> and <em>Prionolejeunea</em>. <em>Bulletin of the Torrey Botanical Club</em> 31 (4): 183–226. https://doi.org/10.2307/2478687</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gottsche, C.M. (1845) <em>Lejeunea aemula</em>.<em> In: </em>Gottsche, C.M., Lindenberg, J.B.W. & Nees, C.G. (Ed.) <em>Synopsis hepaticarum</em>, fasc. 2. Meissner, Hamburg, 145–304. https://doi.org/10.5962/bhl.title.15221</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gradstein, S.R., Churchill, S.P. & Salazar-Allen, N. (2001) Guide to the Bryophytes of Tropical America. <em>Memoirs of the New York Botanical Garden 86</em>: 1–577.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gradstein, S.R. (2021) The Liverworts and Hornworts of Colombia and Ecuador. <em>Memoirs of the New York Botanical Garden</em> 121: 1–723. https://doi.org/10.1007/978-3-030-49450-6</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Grolle, R. (1963) Ein neuer <em>Tylimanthus</em> aus Tasmanien. <em>Nova Hedwigia</em> 6 (3/4): 391–394.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Grolle, R. (1989) <em>Marsupidium</em> in Guyana. <em>Journal of the Hattori Botanical Laboratory</em> 66: 337–342.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Grolle, R. & Persson, H. (1966) Die Gattung <em>Tylimanthus</em> auf den Atlantischen Inseln. <em>Svensk Botanisk Tidskrift</em> 60 (1): 164–174. </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hässel, G.G. & Solari, S.S. (1972) Sinopsis de las especies andinopatagónicas del género <em>Tylimanthus</em> (Hepaticae). <em>Darwiniana</em> 17: 568–591.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hodgson, E.A. (1962) Hepatics from the subantarctic islands of New Zealand including ‘Cape Expedition’ collections from the Auckland and Campbell Islands. <em>Records of the Dominion Museum</em> 4 (11): 101–132.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Jack, J.B. & Stephani, F. (1892) Hepaticae Wallisianae. <em>Hedwigia</em> 31 (1): 11–27.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lindenberg, J.B.W. (1829) <em>Synopsis hepaticarum europaearum</em>. Eduardum Weberum, Bonnae, pp. 1–133.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lindenberg, J.B.W. (1840) <em>Species hepaticarum</em>, fasc. 2–4. Henry & Cohen, Bonn, pp. 37–120.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Mitten, W. (1867) <em>Tylimanthus</em>, <em>Marsupidium</em>.<em> In: </em>Hooker, J.D. (Ed.) <em>Handbook of the New Zealand Flora</em>, Part II, Reeve & Co., London, pp. 751.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Montagne, J.F.C. (1843) Quatrième centurie de plantes cellulaires exotiques nouvelles, décades I-VI. <em>Annales des Sciences Naturelles; Botanique (sér. 2)</em> 19: 238–266.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Montagne, J.F.C. (1856) <em>Sylloge generum specierumque cryptogamarum</em>. Sumptibus J.-B. Ballière, Paris, 498 pp.</span></span></span></p>
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