Abstract
This paper presents new data on the orchid diversity of Vietnam obtained in 2015–2021. It contains descriptions of three new species (Cleisostoma furcatum, Malleola micrantha, Schoenorchis hungii), one new variety (Chiloschista parishii var. minutiflora) and one new form (Cleisostoma furcatum forma albiflora). Additionally six species, Ascocentropsis malipoensis, Chiloschista sweelimii, Pomatocalpa undulatum, Trachoma rhopalorrhachis, Trichoglottis maculata, and Uncifera acuminata are newly recorded for Vietnam. The paper also provides an amended description of a very rare and poorly known species, Cleisostoma equestre. All studied taxa belong to subtribe Aeridinae (tribe Vandeae, subfamily Epidendroideae). The accepted plant name, main synonyms, data on type specimens, ecology, phenology and distribution, estimated IUCN Red List status, studied specimens, brief taxonomic notes, and illustrations are provided for each species listed.
References
<p>Averyanov, L.V. (2012a) New orchids (Orchidaceae) in the flora of Vietnam. <em>Turczaninowia </em>15 (1): 11–18.</p>
<p>Averyanov, L.V. (2012b) New orchid taxa and records in the flora of Vietnam. <em>Taiwania</em> 57 (2): 127–152.</p>
<p>Averyanov, L.V. (2013) The orchids of Vietnam. Illustrated survey. Part 4. Subfamily Epidendroideae (tribes – Arethuseae and Malaxideae). <em>Turczaninowia</em> 16 (1): 5–163.</p>
<p> https://doi.org/10.14258/turczaninowia.16.4.7</p>
<p>Averyanov, L.V. & Vuong, T.B. (2015) Review of the genus <em>Miguelia</em> (Orchidaceae) with a new species, <em>M. cruenta</em>, from southern Vietnam. <em>Taiwania</em> 60 (1): 33–38.</p>
<p>Averyanov, L.V. & Averyanova, A.L. (2003) <em>Updated checklist of the orchids of Vietnam</em>. Vietnam National University Publishing House, Hanoi, 102 pp.</p>
<p>Averyanov, L.V., Phan, K.L., Nguyen, T.H. & Harder, D.K. (2003) Phytogeographic review of Vietnam and adjacent areas of Eastern Indochina. <em>Komarovia</em> 3: 1–83.</p>
<p>Averyanov, L.V., Duy, N.V. & Loc, P.K. (2012) <em>Hymenorchis phitamii</em> (Orchidaceae) – new genus and species in the flora of Vietnam. <em>Taiwania</em> 57 (4): 372–376.</p>
<p>Averyanov, L.V., Nuraliev, M.S., Kuznetsov, A.N. & Kuznetsova, S.P. (2013) <em>Vietorchis furcata</em> – a new orchid species (Orchidaceae) from southern Vietnam. <em>Taiwania</em> 58 (4): 251–256.</p>
<p>Averyanov, L.V., Loc, P.K. & Canh, C.X. (2014) <em>Aerides phongii</em> (Orchidaceae), a new species from southern Vietnam. <em>Turczaninowia</em> 17 (1): 6–9.</p>
<p> https://doi.org/10.14258/turczaninowia.17.1.2</p>
<p>Averyanov, L.V., Duy, N.V., Vinh, T.T., Hoi, Q.V. & Cong, V.K. (2015a) Four new species of orchids (Orchidaceae) in eastern Vietnam. <em>Phytotaxa</em> 238 (2): 136–148.</p>
<p> https://doi.org/10.11646/phytotaxa.238.2.2</p>
<p>Averyanov, L.V., Tich, N.T. & Canh, N.V. (2015b) New species of the genus <em>Cleisostoma </em>in the flora of Vietnam. <em>Taiwania</em> 60 (3): 107–116.</p>
<p>Averyanov, L.V., Nguyen, K.S., Nguyen, T.T., Nguyen, P.T., Nong, V.D., Nguyen, V.C. & Xuan, C.C. (2015c) New orchids in the flora of Vietnam. <em>Wulfenia</em> 22: 137–188.</p>
<p>Averyanov, L.V., Ponert, J., Nguyen, P.T., Nong, V.D., Nguyen, K.S. & Nguyen, V.C. (2016a) The survey of <em>Dendrobium </em>sect. <em>Formosae</em> in Cambodia, Laos and Vietnam. <em>Adansonia</em> 38 (2): 55–73.</p>
<p> https://doi.org/10.5252/a2016n2a5</p>
<p>Averyanov, L.V., Ormerod, P.A., Nong, V.D., Tien, T.V., Chen, T. & Zhang, D.-X. (2016b) <em>Bidoupia phongii</em>, new orchid genus and species (Orchidaceae, Orchidoideae, Goodyerinae) from southern Vietnam. <em>Phytotaxa</em> 266 (4): 289–294.</p>
<p> https://doi.org/10.11646/phytotaxa.266.4.6</p>
<p>Averyanov, L.V., Nong, V.D., Nguyen, K.S., Maisak, T.V., Nguyen, V.C., Phan, Q.T., Nguyen, P.T., Nguyen, T.T. & Truong, B.V. (2016c) New species of orchids (Orchidaceae) in the flora of Vietnam. <em>Taiwania</em> 61 (4): 319–354.</p>
<p>Averyanov, L.V., Vuong, T.B. & Tam, T.Q. (2016d) The genus <em>Liparis</em> (Orchidaceae) in Hon Ba nature reserve, Vietnam, Khanh Hoa province. <em>Turtzaninowia</em> 19 (2): 34–49.</p>
<p> https://doi.org/10.14258/turczaninowia.19.2.4</p>
<p>Averyanov, L.V., Nguyen, K.S., Maisak, T.V., Konstantinov, E.L., Nguyen, T.H. & Bounphanmy, S. (2016e) New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos. <em>Turczaninowia</em> 19 (3): 5–58.</p>
<p> https://doi.org/10.14258/turczaninowia.19.3.1</p>
<p>Averyanov, L.V., Nguyen, K.S., Nong, V.D., Nguyen, V.C., Truong, B.V. & Maisak, T.V. (2017a) <em>Bulbophyllum</em> sect. <em>Hirtula</em> in eastern Indochina. <em>Taiwania</em> 62 (1): 1–23.</p>
<p>Averyanov, L.V., Canh, C.X., Tuan, N.H., Phu, V.A., Nguyen, K.S., Nguyen, T.H. & Maisak, T.V. (2017b) The genus <em>Cypripedium</em> (Orchidaceae) in the flora of Vietnam. <em>Turczaninowia</em> 20 (1): 118–124.</p>
<p> https://doi.org/10.14258/turczaninowia.20.1.10</p>
<p>Averyanov, L.V., Nguyen, V.D., Nguyen, K.S., Dinh, Q.D. & Maisak, T.M. (2018a) <em>Silvorchis vietnamica</em> (Orchidaceae, Orchidoideae, Vietorchidinae), a new miniature mycotrophic species from southern Vietnam. <em>Nordic Journal of Botany </em>36 (7): e01883. [7 pp.]</p>
<p> https://doi.org/10.1111/njb.01883</p>
<p>Averyanov, L.V., Duy, N.V., Tuan, N.H., Nuraliev, M.S., Maisak, T.M. & Nguyen, C.A. (2018b) New species of <em>Bulbophyllum</em> (Orchidaceae) in the flora of Vietnam. <em>Phytotaxa</em> 369 (1): 1–14.</p>
<p> https://doi.org/10.11646/phytotaxa.369.1.1</p>
<p>Averyanov, L.V., Nuraliev, M.S., Kuznetsov, A.N. & Kuznetsova, S.P. (2018c) <em>Biermannia longicheila</em> (Orchidaceae, Aeridinae), a new species from southern Vietnam. <em>Phytotaxa</em> 343 (2): 194–198.</p>
<p> https://doi.org/10.11646/phytotaxa.343.2.11</p>
<p>Averyanov, L.V., Nguyen, V.C., Truong, B.V., Maisak, T.V., Luu, H.T., Nguyen, K.S., Dinh, Q.D., Nguyen, H.T., Chu, X.C., Tran, G., Nguyen, V.K. & Le, H.S. (2018d) New orchids (Orchidaceae: Cymbidieae and Vandeae) in the flora of Vietnam. <em>Taiwania</em> 63 (2): 119–138.</p>
<p>Averyanov, L.V., Nguyen, V.C., Nguyen, H.T., Truong, B.V., Nguyen, P.T., Nguyen, S.K., Maisak, T.V., Nguyen, H.T., Bui, D.N. & Chu, X.C. (2018e) New orchids (Orchidaceae: Epidendroideae and Vandoideae) in the flora of Vietnam. <em>Taiwania</em> 63 (3): 195–219.</p>
<p>Averyanov, L.V., Nguyen, K.S., Truong, B.V., Nguyen, V.C., Maisak, T.V., Thai, T.H., Dat, P.T.T. & Tu, B.N. (2019a) New species of <em>Bulbophyllum</em> (Orchidaceae) in the flora of Vietnam Ⅱ. <em>Phytotaxa</em> 404 (6): 231–244.</p>
<p> https://doi.org/10.11646/phytotaxa.404.6.2</p>
<p>Averyanov, L.V., Nuraliev, M.S., Maisak, T.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2019b) <em>Didymoplexis holochelia</em> (Orchidaceae, Gastrodiinae), a new species from northern Vietnam. <em>Phytotaxa</em> 405 (1): 54–60.</p>
<p> https://doi.org/10.11646/phytotaxa.405.1.5</p>
<p>Averyanov, L.V., Truong, B.V. & Maisak, T.V. (2019c). <em>Bulbophyllum</em> (Orchidaceae) in the flora of Vietnam IⅡ. The revision of <em>B.</em> sect. <em>Lemniscata</em>. <em>Phytotaxa</em> 416 (1): 51–58.</p>
<p> https://doi.org/10.11646/phytotaxa.416.1.6</p>
<p>Averyanov, L.V., Nguyen, V.C., Nguyen, K.S., Maisak, T.V. & Truong, B.V. (2019d) New orchids (Orchidaceae) in the flora of Vietnam I. Epidendroideae. <em>Taiwania</em> 64 (2): 176–188.</p>
<p>Averyanov, L.V., Truong, B.V., Nguyen, V.C., Nguyen, K.S. & Maisak, T.V. (2019e) New orchids (Orchidaceae) in the flora of Vietnam II. Vandeae. <em>Taiwania</em> 64 (3): 285–298.</p>
<p>Averyanov, L.V., Eskov, А.K., Truong, B.V., Nguyen, V.C., Prilepsky, N.G., Nuraliev, M.S., Maisak, T.V. & Manh, V. (2020a) The genus <em>Octarrhena</em> (Orchidaceae, Thelasiinae) in Vietnam with description of two new species. <em>Phytotaxa</em> 459 (4): 265–275.</p>
<p> https://doi.org/10.11646/phytotaxa.459.4.2</p>
<p>Averyanov, L.V., Truong, B.V., Nguyen, V.C., Maisak, T.V., Dinh, Q.D., Nuraliev, M.S., Nguyen, K.S. & Chu, V.T. (2020b) New orchids in the flora of Vietnam Ⅲ (Collabieae, Malaxideae, Nervilieae and Orchideae). <em>Taiwania</em> 65 (4): 478‒492.</p>
<p>Averyanov, L., Dat, P.T.T., Truong, B.V., Orlov, N, Maisak, T., Nguyen, T.H. & Dinhh, Q.D. (2021) Studies of <em>Bulbophyllum </em>(Orchidaceae) in eastern Indochina Ⅳ. New species in the flora of Laos and Vietnam. <em>Phytotaxa</em> 514 (3): 187–204.</p>
<p> https://doi.org/10.11646/phytotaxa.514.3.1</p>
<p>Blume, C.L. (1825) <em>Bijdragen tot de flora van Nederlandsch Indië</em>. Batavia, Ter Lands Drukkerij, 1169 pp.</p>
<p> https://doi.org/10.5962/bhl.title.115427</p>
<p>Breda, J.G.S. (1827–1829) <em>Genera et Species Orchidearum et Asclepiadearum</em>. Batavia, 55 pp.</p>
<p>Chase, M.W, Cameron, K.M., Freudenstein, J.V., Pridgeon, A.M., Salazar, G., Van den Berg, C. & Schuiteman, A. (2015) An updated classification of Orchidaceae. <em>Botanical Journal of the Linnean Society</em> 177 (2): 151–174.</p>
<p> https://doi.org/10.1111/boj.12234</p>
<p>Choudhary, R.K., Tran, B., Hai, D.V., Quang, B.H., Nong, L.V., Kumar, P., Park, S.-H. & Lee, J. (2013) <em>Cordiglottis longipedicellata</em> (Orchidaceae), a new species from Vietnam. <em>Annales Botanici Fennici</em> 50 (1–2): 95–98.</p>
<p> https://doi.org/10.5735/085.050.0118</p>
<p>Dang, M.Q., Averyanov, L.V., Dang, V.S., Maisak, T., Bui, V.H., Tu, B.N. & Truong, B.V. (2022) <em>Bulbophyllum</em> section <em>Rhytionanthos</em> (Orchidaceae) in Vietnam with description of new taxa and new national record. <em>Phytotaxa</em> 530 (3): 257–270.</p>
<p> https://doi.org/10.11646/phytotaxa.530.3.1</p>
<p>Dang, M.Q., Averyanov, L.V., Dang, V.S., Bui, V.H., Amsler, R., Maisak, T. & Truong, B.V. (2021) A new miniature species, <em>Bulbophyllum nghiasonii</em>, sect. <em>Brachyantha</em> (Orchidaceae) from northern Vietnam. <em>Phytotaxa</em> 505 (2): 235–239.</p>
<p> https://doi.org/10.11646/phytotaxa.505.2.10</p>
<p>Don, D. (1825) P<em>rodromus florae Nepalensis</em>. Londini, J. Gale, 256 pp.</p>
<p>Duong, M.T.T.N., Dang, V.S., Averyanov, L.V., Truong, Q.T., Amsler, R. & Truong, B.V. (2021) Two new species of <em>Bulbophyllum</em> sect. <em>Racemosae</em> (Orchidaceae) from Vietnam. <em>Nordic Journal of Botany</em> 2021: e03062: 1–6.</p>
<p>Freycinet, H.L.C.S. (1826–1829) <em>Voyage autour du monde, entrepris par ordre du roi. Exeìcuteì sur les corvettes de S.M. l’Uranie</em>, 4. Paris, Chez Pillet, 522 pp.</p>
<p>Gale, S.W., Schuiteman, A., Watthana, S., Sando, T., Souvannakhoummane, K., Averyanov, L. & Suddee, S. (2016) Studies in Asian <em>Nervilia</em> (Nervilieae, Epidendroideae, Orchidaceae) VI: <em>N. mekongensis</em>, a new species from Thailand, Cambodia, Laos and Vietnam. <em>Phytotaxa</em> 247 (4): 267–273.</p>
<p> https://doi.org/10.11646/phytotaxa.247.4.4</p>
<p>Garay, L.A. (1972) On the systematics of the monopodial orchids I. B<em>otanical Museum leaflets, Harvard University</em> 23 (4): 149–212.</p>
<p> https://doi.org/10.5962/p.168559</p>
<p>Guillaumin, A. (1964) Plantes nouvelles, rares ou critiques des serres du museìum (Notules sur quelques orchideìes d’Indochine. XXXIV). <em>Bulletin du Museìum national d’histoire naturelle</em> 2, 36 (2): 268–269.</p>
<p>Holttum, R.E. (1966) A new <em>Chiloschista</em> from Malaya. <em>Orchid Review</em> 174: 146–147.</p>
<p>Hsu, T.-C., Chen, C.-W., Hung, H.-C., Tram, N.K.T., Truong, Q.C., Luu, H.T. & Li, C.-W. (2020) New and noteworthy orchids (Orchidaceae) discovered in Langbiang Plateau, southern Vietnam 1. <em>Taiwania</em> 65 (2): 237–248.</p>
<p>IUCN Standards and Petitions Subcommittee (2019) <em>Guidelines for using the IUCN Red List categories and criteria</em> Ver. 14. Available from: https://cmsdocs.s3.amazonaws.com/RedListGuidelines.pdf (accessed 1 May 2022)</p>
<p>Kocyan, A. & Schuiteman, A. (2014) New combinations in Aeridinae. <em>Phytotaxa</em> 161 (1): 61–85.</p>
<p> https://doi.org/10.11646/phytotaxa.161.1.3</p>
<p>Kumar, P., Gale, S.W., Kocyan, A., Fischer, G.A., Averyanov, L., Borosova, R., Bhattacharjee, A., Li, J.-H. & Pang, K.S. (2014) <em>Gastrochilus kadooriei</em> (Orchidaceae), a new species from Hong Kong, with notes on allied taxa in section Microphyllae found in the region. <em>Phytotaxa</em> 164 (2): 91–103.</p>
<p> https://doi.org/10.11646/phytotaxa.164.2.3</p>
<p>Lin, T.-P., Lui, H.-Y., Hsieh, C.-F. & Wang, K.-H. (2016) Complete list of the native orchids of Taiwan and their type information. <em>Taiwania</em> 61 (2): 78‒126.</p>
<p>Lindley, J. (1830–1840) <em>The genera and species of orchidaceous plants</em>. London, Ridgways, 554 pp.</p>
<p> https://doi.org/10.5962/bhl.title.499</p>
<p>Lindley, J. (1832) <em>Angraecum eburneum</em>. <em>Edwards’s botanical register</em> 18, tab. 1522.</p>
<p>Lindley, J. (1838) 88. <em>Saccolabium gemmatum</em>. <em>Edwards’s botanical register</em> 24: 50.</p>
<p>Lindley, J. (1858) Contributions to the Orchidology of India. 2. <em>Journal of the proceedings of the Linnean Society. Botany</em> 3: 1–63.</p>
<p> https://doi.org/10.1111/j.1095-8339.1858.tb02055.x</p>
<p>Liu, Z.-J. & Chen, L.-J. (2009) <em>Singchia</em> and <em>Gunnaria</em>, two new genera of Orchidaceae. <em>Journal of Systematics and Evolution</em> 47 (6): 599–604.</p>
<p> https://doi.org/10.1111/j.1759-6831.2009.00047.x</p>
<p>Minh, Q.D., Averyanov, L.V., Maisak, T.M., Bui, V.H., Dang, V.S., Truong, Q.T. & Truong, B.V. (2021) New taxa of <em>Bulbophyllum</em> (Orchidaceae) in the Flora of Vietnam. <em>Taiwania </em>66 (2): 258–266.</p>
<p>Nguyen, D.H., Averyanov, L.V., Dang, V.S., Bui, V.H., Maisak, T. & Truong, B.V. (2021a) <em>Gastrochilus dresslerii</em> (Orchidaceae), a new species from northern Vietnam. <em>Phytotaxa</em> 498 (2): 145–149.</p>
<p> https://doi.org/10.11646/phytotaxa.498.2.8</p>
<p>Nguyen, D.H., Averyanov, L.V., Dang, V.S., Pham, P.D., Maisak, T., Bu, V.H., Dang, M.Q. & Truong, B.V. (2021b) <em>Bulbophyllum xuansonii</em> (Orchidaceae), a new miniature orchid from northern Vietnam and new national record of <em>B. gracilipes</em>. <em>Phytotaxa</em> 522 (2): 150–156.</p>
<p> https://doi.org/10.11646/phytotaxa.522.2.7</p>
<p>Nguyen, H.T. & Averyanov, L.V. (2017) Two endangered ornamental orchid species, <em>Bulbophyllum coweniorum</em> and <em>Esmeralda bella</em> (Orchidaceae), new in the flora of Vietnam. <em>Turczaninowia</em> 20 (1): 68–74.</p>
<p> https://doi.org/10.14258/turczaninowia.20.1.5</p>
<p>Nguyen, M.T., Averyanov, L.V., Bui, V.H., Hoang, N.S., Dang, V.S. & Truong, B.V. (2020) <em>Vanilla tiendatii</em>, a new climbing orchid from Vietnam. <em>Taiwania</em> 65 (4): 438‒442.</p>
<p>Nguyen, V.C., Averyanov, L., Hoang, N.S., Dang, V.S. & Truong, B.V. (2020a) <em>Sunipia quangdangii</em>, new orchid species (Orchidaceae) from southern Vietnam. <em>Phytotaxa</em> 452 (1): 92–98</p>
<p>Nguyen, V.C., Averyanov, L.V., Nguyen, V.K., Pham, T.T.D., Maisak, T.V. & Truong, B.V. (2020b) <em>Cylindrolobus chienii</em> (Orchidaceae), a new species from Central Highlands of Vietnam. <em>Taiwania </em>65 (3): 272‒276.</p>
<p>Nguyen, V.C., Averyanov, L.V., Maisak, T.V., Nguyen, T.L.T., Nguyen, V.K. & Truong, B.V. (2021) <em>Dendrobium parahendersonii</em>, a new orchid species (Orchidaceae) from southern Vietnam. <em>Taiwania</em> 66 (3): 360‒363.</p>
<p>Nguyen, V.C., Averyanov, L.V., Maisak, T.V., Nguyen, T.L.T., Nguyen, V.K. & Truong, B.V. (2022) <em>Gastrochilus pankajkumarii</em>, (Aeridinae, Epidendroideae, Orchidaceae) a new lithophytic orchid from southern Vietnam. <em>Taiwania</em> 67 (1): 35‒39.</p>
<p>Nguyen, V.T., Averyanov, L.V., Dang, V.S., Maisak, T.V., Bui, V.H., Dang, M.Q., Boo, S.M. & Truong, B.V. (2022a) A new species of <em>Bulbophyllum</em> from northern of Vietnam. <em>Phytotaxa</em> 542 (1): 95–99.</p>
<p> https://doi.org/10.11646/phytotaxa.542.1.9</p>
<p>Nguyen, V.T., Averyanov, L.V., Dang, V.S., Maisak, T.V., Bui, V.H., Nguyen, V.C. & Truong, B.V. (2022b) A new species, <em>Bulbophyllum mamillatum</em> and a new national record, <em>B. retusum</em> (Orchidaceae) in the flora of Vietnam. <em>Taiwania</em> 67 (2): 181‒185.</p>
<p>Nong, V.D. & Averyanov, L.V. (2015) <em>Bulbophyllum bidoupense</em> and <em>Schoenorchis hangianae –</em>new species of orchids (Orchidaceae) from southern Vietnam. <em>Phytotaxa</em> 213 (2): 113–121.</p>
<p> https://doi.org/10.11646/phytotaxa.213.2.4</p>
<p>Nuraliev, M.S., Efimov, P.G., Averyanov, L.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2014) <em>Cephalanthera exigua</em> (Orchidaceae), a new species and genus for the flora of Vietnam. <em>Wulfienia</em> 21: 95–102.</p>
<p>Nuraliev, M.S., Averyanov, L.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2015) Review of the genus <em>Plocoglottis</em> (Orchidaceae) in Cambodia, Laos and Vietnam. <em>Wulfenia </em>22: 189–199.</p>
<p>O’Byrne, P. (2017) A clarification of the <em>Malleola dentifera - Robiquetia insectifera - R. vietnamensis</em> complex. <em>Malesian Orchid Journal</em> 20: 91–121.</p>
<p>Pearce, N.R. & Cribb, P.J. (2002) <em>The orchids of Bhutan.</em> The Charlesworth Group, Huddersfield, 643 pp.</p>
<p>Pham, P.D., Averyanov, L.V., Dang, V.S., Nguyen, D.H., Maisak, T., Truong, Q.T., Dang, M.Q. & Truong, B.V. (2021) A new species and new record of <em>Bulbophyllum</em> (Orchidaceae) from Vietnam. <em>Phytotaxa</em> 522 (1): 68–72.</p>
<p> https://doi.org/10.11646/phytotaxa.522.1.8</p>
<p>Reichenbach, H.G. (1857) Orchideae Zollingerianae itineris primi racensentur. <em>Bonplandia</em> 5: 34–43.</p>
<p>Rice, R. (2018) <em>Photo Intro to: Vandoid Orchid Genera in Asia Paperback</em>. Nature & Travel Books, 202 pp.</p>
<p>Schuiteman, A., Averyanov, L. & Rybkova, R. (2013) <em>Vanilla atropogon</em>, a new species from Vietnam. <em>Orchid Journal</em> 1 (1): 10–16.</p>
<p>Schuiteman, A. & Averyanov, L.V. (2017) Systematics of <em>Yoania prainii</em> (Orchidaceae: Calypsoinae). <em>Phytotaxa</em> 309 (3): 288–290.</p>
<p> https://doi.org/10.11646/phytotaxa.309.3.12</p>
<p>Seidenfaden, G. (1988) Orchid genera in Thailand XIV. Fifty-nine Vandoid genera. <em>Opera Botanica</em> 95: 1–399.</p>
<p>Seidenfaden, G. & Wood, J.J. (1992) <em>The orchids of Peninsular Malaysia and Singapore</em>. Fredensborg, Olsen & Olsen, 779 pp.</p>
<p>Smith, J.J. (1905) <em>Die Orchideen von Java</em>. Leiden, E. J. Brill, 672 pp.</p>
<p> https://doi.org/10.5962/bhl.title.86938</p>
<p>Smith, J.J. (1912) <em>Sarcanthus </em>Lindl. und die nächstverwandten Gattungen. <em>Natuurkundig tijdschrift voor Nederlandsch Indië</em> 72: 79–115.</p>
<p>Smith, J.J. (1918) Die Orchideen von Java. <em>Bulletin du Jardin botanique de Buitenzorg</em> Ser. 2, 26: 1–135.</p>
<p>Smith, J.J. (1927) <em>Malleola dentifera</em> J.J.Sm. <em>Bulletin du Jardin botanique de Buitenzorg.</em> Ser. 3, 9: 191.</p>
<p>Thiers, B. (2020) [continuously updated] <em>Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium</em>. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 1 May 2022)</p>
<p>Truong, B.V., Averyanov, L.V., Amsler, R., Nguyen, V.C., Tran, H., Maisak, T.V. & Dang, V.S. (2020a) New species, <em>Bulbophyllum trongquyetii</em> (Orchidaceae) from Vietnam. <em>Phytotaxa </em>464 (4): 293–298.</p>
<p>Truong, B.V., Dang, V.S., Tran, H., Maisak, T.V. & Averyanov, L.V. (2020b) <em>Bulbophyllum sridithii</em> (Orchidaceae), new species from Vietnam. <em>Taiwania</em> 65 (4): 473‒477.</p>
<p>Truong, B.V., Hsu, T.C., Bui, V.H., Tu, B.N., Dang, V.S., Luu, H.T. & Sutsgu, K. (2020c) The genus <em>Lecanorchis</em> Blume (Orchidaceae) in the flora of Vietnam. <em>Taiwania</em> 65 (1): 86‒94.</p>
<p><em>Vietnam Administrative Atlas</em> (2015) Vietnam Publishing House of Natural Resources, Environment and Cartography, Hanoi, 131 pp.</p>
<p>Zhou, Z., Cheng, Z., Liu, Q., Zhang, J., Hu, A., Huang, M., Hu, C. & Tian, H. (2016) An updated checklist of Orchidaceae for China, with two new national records. <em>Phytotaxa</em> 276 (1): 1–148.</p>
<p> https://doi.org/10.11646/phytotaxa.276.1.1</p>
<p>Averyanov, L.V. (2012b) New orchid taxa and records in the flora of Vietnam. <em>Taiwania</em> 57 (2): 127–152.</p>
<p>Averyanov, L.V. (2013) The orchids of Vietnam. Illustrated survey. Part 4. Subfamily Epidendroideae (tribes – Arethuseae and Malaxideae). <em>Turczaninowia</em> 16 (1): 5–163.</p>
<p> https://doi.org/10.14258/turczaninowia.16.4.7</p>
<p>Averyanov, L.V. & Vuong, T.B. (2015) Review of the genus <em>Miguelia</em> (Orchidaceae) with a new species, <em>M. cruenta</em>, from southern Vietnam. <em>Taiwania</em> 60 (1): 33–38.</p>
<p>Averyanov, L.V. & Averyanova, A.L. (2003) <em>Updated checklist of the orchids of Vietnam</em>. Vietnam National University Publishing House, Hanoi, 102 pp.</p>
<p>Averyanov, L.V., Phan, K.L., Nguyen, T.H. & Harder, D.K. (2003) Phytogeographic review of Vietnam and adjacent areas of Eastern Indochina. <em>Komarovia</em> 3: 1–83.</p>
<p>Averyanov, L.V., Duy, N.V. & Loc, P.K. (2012) <em>Hymenorchis phitamii</em> (Orchidaceae) – new genus and species in the flora of Vietnam. <em>Taiwania</em> 57 (4): 372–376.</p>
<p>Averyanov, L.V., Nuraliev, M.S., Kuznetsov, A.N. & Kuznetsova, S.P. (2013) <em>Vietorchis furcata</em> – a new orchid species (Orchidaceae) from southern Vietnam. <em>Taiwania</em> 58 (4): 251–256.</p>
<p>Averyanov, L.V., Loc, P.K. & Canh, C.X. (2014) <em>Aerides phongii</em> (Orchidaceae), a new species from southern Vietnam. <em>Turczaninowia</em> 17 (1): 6–9.</p>
<p> https://doi.org/10.14258/turczaninowia.17.1.2</p>
<p>Averyanov, L.V., Duy, N.V., Vinh, T.T., Hoi, Q.V. & Cong, V.K. (2015a) Four new species of orchids (Orchidaceae) in eastern Vietnam. <em>Phytotaxa</em> 238 (2): 136–148.</p>
<p> https://doi.org/10.11646/phytotaxa.238.2.2</p>
<p>Averyanov, L.V., Tich, N.T. & Canh, N.V. (2015b) New species of the genus <em>Cleisostoma </em>in the flora of Vietnam. <em>Taiwania</em> 60 (3): 107–116.</p>
<p>Averyanov, L.V., Nguyen, K.S., Nguyen, T.T., Nguyen, P.T., Nong, V.D., Nguyen, V.C. & Xuan, C.C. (2015c) New orchids in the flora of Vietnam. <em>Wulfenia</em> 22: 137–188.</p>
<p>Averyanov, L.V., Ponert, J., Nguyen, P.T., Nong, V.D., Nguyen, K.S. & Nguyen, V.C. (2016a) The survey of <em>Dendrobium </em>sect. <em>Formosae</em> in Cambodia, Laos and Vietnam. <em>Adansonia</em> 38 (2): 55–73.</p>
<p> https://doi.org/10.5252/a2016n2a5</p>
<p>Averyanov, L.V., Ormerod, P.A., Nong, V.D., Tien, T.V., Chen, T. & Zhang, D.-X. (2016b) <em>Bidoupia phongii</em>, new orchid genus and species (Orchidaceae, Orchidoideae, Goodyerinae) from southern Vietnam. <em>Phytotaxa</em> 266 (4): 289–294.</p>
<p> https://doi.org/10.11646/phytotaxa.266.4.6</p>
<p>Averyanov, L.V., Nong, V.D., Nguyen, K.S., Maisak, T.V., Nguyen, V.C., Phan, Q.T., Nguyen, P.T., Nguyen, T.T. & Truong, B.V. (2016c) New species of orchids (Orchidaceae) in the flora of Vietnam. <em>Taiwania</em> 61 (4): 319–354.</p>
<p>Averyanov, L.V., Vuong, T.B. & Tam, T.Q. (2016d) The genus <em>Liparis</em> (Orchidaceae) in Hon Ba nature reserve, Vietnam, Khanh Hoa province. <em>Turtzaninowia</em> 19 (2): 34–49.</p>
<p> https://doi.org/10.14258/turczaninowia.19.2.4</p>
<p>Averyanov, L.V., Nguyen, K.S., Maisak, T.V., Konstantinov, E.L., Nguyen, T.H. & Bounphanmy, S. (2016e) New and rare orchids (Orchidaceae) in the flora of Cambodia and Laos. <em>Turczaninowia</em> 19 (3): 5–58.</p>
<p> https://doi.org/10.14258/turczaninowia.19.3.1</p>
<p>Averyanov, L.V., Nguyen, K.S., Nong, V.D., Nguyen, V.C., Truong, B.V. & Maisak, T.V. (2017a) <em>Bulbophyllum</em> sect. <em>Hirtula</em> in eastern Indochina. <em>Taiwania</em> 62 (1): 1–23.</p>
<p>Averyanov, L.V., Canh, C.X., Tuan, N.H., Phu, V.A., Nguyen, K.S., Nguyen, T.H. & Maisak, T.V. (2017b) The genus <em>Cypripedium</em> (Orchidaceae) in the flora of Vietnam. <em>Turczaninowia</em> 20 (1): 118–124.</p>
<p> https://doi.org/10.14258/turczaninowia.20.1.10</p>
<p>Averyanov, L.V., Nguyen, V.D., Nguyen, K.S., Dinh, Q.D. & Maisak, T.M. (2018a) <em>Silvorchis vietnamica</em> (Orchidaceae, Orchidoideae, Vietorchidinae), a new miniature mycotrophic species from southern Vietnam. <em>Nordic Journal of Botany </em>36 (7): e01883. [7 pp.]</p>
<p> https://doi.org/10.1111/njb.01883</p>
<p>Averyanov, L.V., Duy, N.V., Tuan, N.H., Nuraliev, M.S., Maisak, T.M. & Nguyen, C.A. (2018b) New species of <em>Bulbophyllum</em> (Orchidaceae) in the flora of Vietnam. <em>Phytotaxa</em> 369 (1): 1–14.</p>
<p> https://doi.org/10.11646/phytotaxa.369.1.1</p>
<p>Averyanov, L.V., Nuraliev, M.S., Kuznetsov, A.N. & Kuznetsova, S.P. (2018c) <em>Biermannia longicheila</em> (Orchidaceae, Aeridinae), a new species from southern Vietnam. <em>Phytotaxa</em> 343 (2): 194–198.</p>
<p> https://doi.org/10.11646/phytotaxa.343.2.11</p>
<p>Averyanov, L.V., Nguyen, V.C., Truong, B.V., Maisak, T.V., Luu, H.T., Nguyen, K.S., Dinh, Q.D., Nguyen, H.T., Chu, X.C., Tran, G., Nguyen, V.K. & Le, H.S. (2018d) New orchids (Orchidaceae: Cymbidieae and Vandeae) in the flora of Vietnam. <em>Taiwania</em> 63 (2): 119–138.</p>
<p>Averyanov, L.V., Nguyen, V.C., Nguyen, H.T., Truong, B.V., Nguyen, P.T., Nguyen, S.K., Maisak, T.V., Nguyen, H.T., Bui, D.N. & Chu, X.C. (2018e) New orchids (Orchidaceae: Epidendroideae and Vandoideae) in the flora of Vietnam. <em>Taiwania</em> 63 (3): 195–219.</p>
<p>Averyanov, L.V., Nguyen, K.S., Truong, B.V., Nguyen, V.C., Maisak, T.V., Thai, T.H., Dat, P.T.T. & Tu, B.N. (2019a) New species of <em>Bulbophyllum</em> (Orchidaceae) in the flora of Vietnam Ⅱ. <em>Phytotaxa</em> 404 (6): 231–244.</p>
<p> https://doi.org/10.11646/phytotaxa.404.6.2</p>
<p>Averyanov, L.V., Nuraliev, M.S., Maisak, T.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2019b) <em>Didymoplexis holochelia</em> (Orchidaceae, Gastrodiinae), a new species from northern Vietnam. <em>Phytotaxa</em> 405 (1): 54–60.</p>
<p> https://doi.org/10.11646/phytotaxa.405.1.5</p>
<p>Averyanov, L.V., Truong, B.V. & Maisak, T.V. (2019c). <em>Bulbophyllum</em> (Orchidaceae) in the flora of Vietnam IⅡ. The revision of <em>B.</em> sect. <em>Lemniscata</em>. <em>Phytotaxa</em> 416 (1): 51–58.</p>
<p> https://doi.org/10.11646/phytotaxa.416.1.6</p>
<p>Averyanov, L.V., Nguyen, V.C., Nguyen, K.S., Maisak, T.V. & Truong, B.V. (2019d) New orchids (Orchidaceae) in the flora of Vietnam I. Epidendroideae. <em>Taiwania</em> 64 (2): 176–188.</p>
<p>Averyanov, L.V., Truong, B.V., Nguyen, V.C., Nguyen, K.S. & Maisak, T.V. (2019e) New orchids (Orchidaceae) in the flora of Vietnam II. Vandeae. <em>Taiwania</em> 64 (3): 285–298.</p>
<p>Averyanov, L.V., Eskov, А.K., Truong, B.V., Nguyen, V.C., Prilepsky, N.G., Nuraliev, M.S., Maisak, T.V. & Manh, V. (2020a) The genus <em>Octarrhena</em> (Orchidaceae, Thelasiinae) in Vietnam with description of two new species. <em>Phytotaxa</em> 459 (4): 265–275.</p>
<p> https://doi.org/10.11646/phytotaxa.459.4.2</p>
<p>Averyanov, L.V., Truong, B.V., Nguyen, V.C., Maisak, T.V., Dinh, Q.D., Nuraliev, M.S., Nguyen, K.S. & Chu, V.T. (2020b) New orchids in the flora of Vietnam Ⅲ (Collabieae, Malaxideae, Nervilieae and Orchideae). <em>Taiwania</em> 65 (4): 478‒492.</p>
<p>Averyanov, L., Dat, P.T.T., Truong, B.V., Orlov, N, Maisak, T., Nguyen, T.H. & Dinhh, Q.D. (2021) Studies of <em>Bulbophyllum </em>(Orchidaceae) in eastern Indochina Ⅳ. New species in the flora of Laos and Vietnam. <em>Phytotaxa</em> 514 (3): 187–204.</p>
<p> https://doi.org/10.11646/phytotaxa.514.3.1</p>
<p>Blume, C.L. (1825) <em>Bijdragen tot de flora van Nederlandsch Indië</em>. Batavia, Ter Lands Drukkerij, 1169 pp.</p>
<p> https://doi.org/10.5962/bhl.title.115427</p>
<p>Breda, J.G.S. (1827–1829) <em>Genera et Species Orchidearum et Asclepiadearum</em>. Batavia, 55 pp.</p>
<p>Chase, M.W, Cameron, K.M., Freudenstein, J.V., Pridgeon, A.M., Salazar, G., Van den Berg, C. & Schuiteman, A. (2015) An updated classification of Orchidaceae. <em>Botanical Journal of the Linnean Society</em> 177 (2): 151–174.</p>
<p> https://doi.org/10.1111/boj.12234</p>
<p>Choudhary, R.K., Tran, B., Hai, D.V., Quang, B.H., Nong, L.V., Kumar, P., Park, S.-H. & Lee, J. (2013) <em>Cordiglottis longipedicellata</em> (Orchidaceae), a new species from Vietnam. <em>Annales Botanici Fennici</em> 50 (1–2): 95–98.</p>
<p> https://doi.org/10.5735/085.050.0118</p>
<p>Dang, M.Q., Averyanov, L.V., Dang, V.S., Maisak, T., Bui, V.H., Tu, B.N. & Truong, B.V. (2022) <em>Bulbophyllum</em> section <em>Rhytionanthos</em> (Orchidaceae) in Vietnam with description of new taxa and new national record. <em>Phytotaxa</em> 530 (3): 257–270.</p>
<p> https://doi.org/10.11646/phytotaxa.530.3.1</p>
<p>Dang, M.Q., Averyanov, L.V., Dang, V.S., Bui, V.H., Amsler, R., Maisak, T. & Truong, B.V. (2021) A new miniature species, <em>Bulbophyllum nghiasonii</em>, sect. <em>Brachyantha</em> (Orchidaceae) from northern Vietnam. <em>Phytotaxa</em> 505 (2): 235–239.</p>
<p> https://doi.org/10.11646/phytotaxa.505.2.10</p>
<p>Don, D. (1825) P<em>rodromus florae Nepalensis</em>. Londini, J. Gale, 256 pp.</p>
<p>Duong, M.T.T.N., Dang, V.S., Averyanov, L.V., Truong, Q.T., Amsler, R. & Truong, B.V. (2021) Two new species of <em>Bulbophyllum</em> sect. <em>Racemosae</em> (Orchidaceae) from Vietnam. <em>Nordic Journal of Botany</em> 2021: e03062: 1–6.</p>
<p>Freycinet, H.L.C.S. (1826–1829) <em>Voyage autour du monde, entrepris par ordre du roi. Exeìcuteì sur les corvettes de S.M. l’Uranie</em>, 4. Paris, Chez Pillet, 522 pp.</p>
<p>Gale, S.W., Schuiteman, A., Watthana, S., Sando, T., Souvannakhoummane, K., Averyanov, L. & Suddee, S. (2016) Studies in Asian <em>Nervilia</em> (Nervilieae, Epidendroideae, Orchidaceae) VI: <em>N. mekongensis</em>, a new species from Thailand, Cambodia, Laos and Vietnam. <em>Phytotaxa</em> 247 (4): 267–273.</p>
<p> https://doi.org/10.11646/phytotaxa.247.4.4</p>
<p>Garay, L.A. (1972) On the systematics of the monopodial orchids I. B<em>otanical Museum leaflets, Harvard University</em> 23 (4): 149–212.</p>
<p> https://doi.org/10.5962/p.168559</p>
<p>Guillaumin, A. (1964) Plantes nouvelles, rares ou critiques des serres du museìum (Notules sur quelques orchideìes d’Indochine. XXXIV). <em>Bulletin du Museìum national d’histoire naturelle</em> 2, 36 (2): 268–269.</p>
<p>Holttum, R.E. (1966) A new <em>Chiloschista</em> from Malaya. <em>Orchid Review</em> 174: 146–147.</p>
<p>Hsu, T.-C., Chen, C.-W., Hung, H.-C., Tram, N.K.T., Truong, Q.C., Luu, H.T. & Li, C.-W. (2020) New and noteworthy orchids (Orchidaceae) discovered in Langbiang Plateau, southern Vietnam 1. <em>Taiwania</em> 65 (2): 237–248.</p>
<p>IUCN Standards and Petitions Subcommittee (2019) <em>Guidelines for using the IUCN Red List categories and criteria</em> Ver. 14. Available from: https://cmsdocs.s3.amazonaws.com/RedListGuidelines.pdf (accessed 1 May 2022)</p>
<p>Kocyan, A. & Schuiteman, A. (2014) New combinations in Aeridinae. <em>Phytotaxa</em> 161 (1): 61–85.</p>
<p> https://doi.org/10.11646/phytotaxa.161.1.3</p>
<p>Kumar, P., Gale, S.W., Kocyan, A., Fischer, G.A., Averyanov, L., Borosova, R., Bhattacharjee, A., Li, J.-H. & Pang, K.S. (2014) <em>Gastrochilus kadooriei</em> (Orchidaceae), a new species from Hong Kong, with notes on allied taxa in section Microphyllae found in the region. <em>Phytotaxa</em> 164 (2): 91–103.</p>
<p> https://doi.org/10.11646/phytotaxa.164.2.3</p>
<p>Lin, T.-P., Lui, H.-Y., Hsieh, C.-F. & Wang, K.-H. (2016) Complete list of the native orchids of Taiwan and their type information. <em>Taiwania</em> 61 (2): 78‒126.</p>
<p>Lindley, J. (1830–1840) <em>The genera and species of orchidaceous plants</em>. London, Ridgways, 554 pp.</p>
<p> https://doi.org/10.5962/bhl.title.499</p>
<p>Lindley, J. (1832) <em>Angraecum eburneum</em>. <em>Edwards’s botanical register</em> 18, tab. 1522.</p>
<p>Lindley, J. (1838) 88. <em>Saccolabium gemmatum</em>. <em>Edwards’s botanical register</em> 24: 50.</p>
<p>Lindley, J. (1858) Contributions to the Orchidology of India. 2. <em>Journal of the proceedings of the Linnean Society. Botany</em> 3: 1–63.</p>
<p> https://doi.org/10.1111/j.1095-8339.1858.tb02055.x</p>
<p>Liu, Z.-J. & Chen, L.-J. (2009) <em>Singchia</em> and <em>Gunnaria</em>, two new genera of Orchidaceae. <em>Journal of Systematics and Evolution</em> 47 (6): 599–604.</p>
<p> https://doi.org/10.1111/j.1759-6831.2009.00047.x</p>
<p>Minh, Q.D., Averyanov, L.V., Maisak, T.M., Bui, V.H., Dang, V.S., Truong, Q.T. & Truong, B.V. (2021) New taxa of <em>Bulbophyllum</em> (Orchidaceae) in the Flora of Vietnam. <em>Taiwania </em>66 (2): 258–266.</p>
<p>Nguyen, D.H., Averyanov, L.V., Dang, V.S., Bui, V.H., Maisak, T. & Truong, B.V. (2021a) <em>Gastrochilus dresslerii</em> (Orchidaceae), a new species from northern Vietnam. <em>Phytotaxa</em> 498 (2): 145–149.</p>
<p> https://doi.org/10.11646/phytotaxa.498.2.8</p>
<p>Nguyen, D.H., Averyanov, L.V., Dang, V.S., Pham, P.D., Maisak, T., Bu, V.H., Dang, M.Q. & Truong, B.V. (2021b) <em>Bulbophyllum xuansonii</em> (Orchidaceae), a new miniature orchid from northern Vietnam and new national record of <em>B. gracilipes</em>. <em>Phytotaxa</em> 522 (2): 150–156.</p>
<p> https://doi.org/10.11646/phytotaxa.522.2.7</p>
<p>Nguyen, H.T. & Averyanov, L.V. (2017) Two endangered ornamental orchid species, <em>Bulbophyllum coweniorum</em> and <em>Esmeralda bella</em> (Orchidaceae), new in the flora of Vietnam. <em>Turczaninowia</em> 20 (1): 68–74.</p>
<p> https://doi.org/10.14258/turczaninowia.20.1.5</p>
<p>Nguyen, M.T., Averyanov, L.V., Bui, V.H., Hoang, N.S., Dang, V.S. & Truong, B.V. (2020) <em>Vanilla tiendatii</em>, a new climbing orchid from Vietnam. <em>Taiwania</em> 65 (4): 438‒442.</p>
<p>Nguyen, V.C., Averyanov, L., Hoang, N.S., Dang, V.S. & Truong, B.V. (2020a) <em>Sunipia quangdangii</em>, new orchid species (Orchidaceae) from southern Vietnam. <em>Phytotaxa</em> 452 (1): 92–98</p>
<p>Nguyen, V.C., Averyanov, L.V., Nguyen, V.K., Pham, T.T.D., Maisak, T.V. & Truong, B.V. (2020b) <em>Cylindrolobus chienii</em> (Orchidaceae), a new species from Central Highlands of Vietnam. <em>Taiwania </em>65 (3): 272‒276.</p>
<p>Nguyen, V.C., Averyanov, L.V., Maisak, T.V., Nguyen, T.L.T., Nguyen, V.K. & Truong, B.V. (2021) <em>Dendrobium parahendersonii</em>, a new orchid species (Orchidaceae) from southern Vietnam. <em>Taiwania</em> 66 (3): 360‒363.</p>
<p>Nguyen, V.C., Averyanov, L.V., Maisak, T.V., Nguyen, T.L.T., Nguyen, V.K. & Truong, B.V. (2022) <em>Gastrochilus pankajkumarii</em>, (Aeridinae, Epidendroideae, Orchidaceae) a new lithophytic orchid from southern Vietnam. <em>Taiwania</em> 67 (1): 35‒39.</p>
<p>Nguyen, V.T., Averyanov, L.V., Dang, V.S., Maisak, T.V., Bui, V.H., Dang, M.Q., Boo, S.M. & Truong, B.V. (2022a) A new species of <em>Bulbophyllum</em> from northern of Vietnam. <em>Phytotaxa</em> 542 (1): 95–99.</p>
<p> https://doi.org/10.11646/phytotaxa.542.1.9</p>
<p>Nguyen, V.T., Averyanov, L.V., Dang, V.S., Maisak, T.V., Bui, V.H., Nguyen, V.C. & Truong, B.V. (2022b) A new species, <em>Bulbophyllum mamillatum</em> and a new national record, <em>B. retusum</em> (Orchidaceae) in the flora of Vietnam. <em>Taiwania</em> 67 (2): 181‒185.</p>
<p>Nong, V.D. & Averyanov, L.V. (2015) <em>Bulbophyllum bidoupense</em> and <em>Schoenorchis hangianae –</em>new species of orchids (Orchidaceae) from southern Vietnam. <em>Phytotaxa</em> 213 (2): 113–121.</p>
<p> https://doi.org/10.11646/phytotaxa.213.2.4</p>
<p>Nuraliev, M.S., Efimov, P.G., Averyanov, L.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2014) <em>Cephalanthera exigua</em> (Orchidaceae), a new species and genus for the flora of Vietnam. <em>Wulfienia</em> 21: 95–102.</p>
<p>Nuraliev, M.S., Averyanov, L.V., Kuznetsov, A.N. & Kuznetsova, S.P. (2015) Review of the genus <em>Plocoglottis</em> (Orchidaceae) in Cambodia, Laos and Vietnam. <em>Wulfenia </em>22: 189–199.</p>
<p>O’Byrne, P. (2017) A clarification of the <em>Malleola dentifera - Robiquetia insectifera - R. vietnamensis</em> complex. <em>Malesian Orchid Journal</em> 20: 91–121.</p>
<p>Pearce, N.R. & Cribb, P.J. (2002) <em>The orchids of Bhutan.</em> The Charlesworth Group, Huddersfield, 643 pp.</p>
<p>Pham, P.D., Averyanov, L.V., Dang, V.S., Nguyen, D.H., Maisak, T., Truong, Q.T., Dang, M.Q. & Truong, B.V. (2021) A new species and new record of <em>Bulbophyllum</em> (Orchidaceae) from Vietnam. <em>Phytotaxa</em> 522 (1): 68–72.</p>
<p> https://doi.org/10.11646/phytotaxa.522.1.8</p>
<p>Reichenbach, H.G. (1857) Orchideae Zollingerianae itineris primi racensentur. <em>Bonplandia</em> 5: 34–43.</p>
<p>Rice, R. (2018) <em>Photo Intro to: Vandoid Orchid Genera in Asia Paperback</em>. Nature & Travel Books, 202 pp.</p>
<p>Schuiteman, A., Averyanov, L. & Rybkova, R. (2013) <em>Vanilla atropogon</em>, a new species from Vietnam. <em>Orchid Journal</em> 1 (1): 10–16.</p>
<p>Schuiteman, A. & Averyanov, L.V. (2017) Systematics of <em>Yoania prainii</em> (Orchidaceae: Calypsoinae). <em>Phytotaxa</em> 309 (3): 288–290.</p>
<p> https://doi.org/10.11646/phytotaxa.309.3.12</p>
<p>Seidenfaden, G. (1988) Orchid genera in Thailand XIV. Fifty-nine Vandoid genera. <em>Opera Botanica</em> 95: 1–399.</p>
<p>Seidenfaden, G. & Wood, J.J. (1992) <em>The orchids of Peninsular Malaysia and Singapore</em>. Fredensborg, Olsen & Olsen, 779 pp.</p>
<p>Smith, J.J. (1905) <em>Die Orchideen von Java</em>. Leiden, E. J. Brill, 672 pp.</p>
<p> https://doi.org/10.5962/bhl.title.86938</p>
<p>Smith, J.J. (1912) <em>Sarcanthus </em>Lindl. und die nächstverwandten Gattungen. <em>Natuurkundig tijdschrift voor Nederlandsch Indië</em> 72: 79–115.</p>
<p>Smith, J.J. (1918) Die Orchideen von Java. <em>Bulletin du Jardin botanique de Buitenzorg</em> Ser. 2, 26: 1–135.</p>
<p>Smith, J.J. (1927) <em>Malleola dentifera</em> J.J.Sm. <em>Bulletin du Jardin botanique de Buitenzorg.</em> Ser. 3, 9: 191.</p>
<p>Thiers, B. (2020) [continuously updated] <em>Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium</em>. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 1 May 2022)</p>
<p>Truong, B.V., Averyanov, L.V., Amsler, R., Nguyen, V.C., Tran, H., Maisak, T.V. & Dang, V.S. (2020a) New species, <em>Bulbophyllum trongquyetii</em> (Orchidaceae) from Vietnam. <em>Phytotaxa </em>464 (4): 293–298.</p>
<p>Truong, B.V., Dang, V.S., Tran, H., Maisak, T.V. & Averyanov, L.V. (2020b) <em>Bulbophyllum sridithii</em> (Orchidaceae), new species from Vietnam. <em>Taiwania</em> 65 (4): 473‒477.</p>
<p>Truong, B.V., Hsu, T.C., Bui, V.H., Tu, B.N., Dang, V.S., Luu, H.T. & Sutsgu, K. (2020c) The genus <em>Lecanorchis</em> Blume (Orchidaceae) in the flora of Vietnam. <em>Taiwania</em> 65 (1): 86‒94.</p>
<p><em>Vietnam Administrative Atlas</em> (2015) Vietnam Publishing House of Natural Resources, Environment and Cartography, Hanoi, 131 pp.</p>
<p>Zhou, Z., Cheng, Z., Liu, Q., Zhang, J., Hu, A., Huang, M., Hu, C. & Tian, H. (2016) An updated checklist of Orchidaceae for China, with two new national records. <em>Phytotaxa</em> 276 (1): 1–148.</p>
<p> https://doi.org/10.11646/phytotaxa.276.1.1</p>