Abstract
The nomenclature of the three names that Emmanuel Drake del Castillo published in Kalanchoe (Crassulaceae subfam. Kalanchooideae), K. antanosiana, K. beharensis, and K. linearifolia, in 1903 is reviewed and discussed. Kalanchoe beharensis and K. linearifolia are today accepted species, but K. antanosiana is widely treated as a synonym of K. orgyalis. Corrections are made to the typification of the names K. beharensis and K. vantieghemii to maintain the current application of these names, with the latter being a homotypic synonym of K. beharensis.
References
<p>Adanson, M. (1763) <em>Familles des Plantes</em>.<em> Seconde Partie. Etat actuel de la Botanike</em>. Chez Vincent, Imprimeur-Libraire de M<sup>gt</sup> le Comte de Provence, Paris, 640 pp. [https://www.biodiversitylibrary.org/item/6958#page/794/mode/1up]</p>
<p>Baena, S., Beentje, H., Cameron, A., Lowry, P., Moat, J., Rabehevitra, D., Rahantamalala, J., Rakotoarisoa, S., Randriarimalala, Z.L., Rasolohery, A., Schatz, G., Smith, P. & Tabor, K. (2007) Atlas of the vegetation of Madagascar. / Atlas de la vegetation de Madagascar. <em>In</em>: J. Moat & P. Smith (Eds.) <em>Atlas of the vegetation of Madagascar. / Atlas de la vegetation de Madagascar</em>. Kew Publishing, Royal Botanic Gardens, Kew, Richmond, Surrey, 124 pp.</p>
<p>Baillon, M.H. (1887) Liste des plantes de Madagascar (suite de la page 624). <em>Bulletin Mensuel de la Société Linnéenne de Paris</em> 1 (No. 84): 671–672. [https://www.biodiversitylibrary.org/page/11162078#page/24/mode/1up]</p>
<p>Baillon, M.H. (1888) <em>Kalanchoe grandidieri</em>, Plate 57 [of Plates 51–88]. <em>In</em>: A. Grandidier, <em>Histoire physique, naturelle et politique Madagascar publiée par Alfred Grandidier. Vol. XXX. Histoire naturelle des plantes par M. E. Drake del Castillo</em>. Tome II – Texte – 1, partie. 50 Fascicule. Imprimé par Autorisation de M. Le Garde des Sceaux. À l’Imprimerie Nationale, Paris, 130 plates. [https://www.biodiversitylibrary.org/item/41106#page/71/mode/1up]</p>
<p>Baker, J.G. (1882) Contributions to the flora of central Madagascar (continued from p. 70 and to be continued). <em>Journal of Botany, British and Foreign. London</em> 20: 109–114. [https://www.biodiversitylibrary.org/item/35873#page/125/mode/1up]</p>
<p>Boiteau, P. & Allorge-Boiteau, L. (1995) Kalanchoe <em>(Crassulacées) de Madagascar. Systématique, écophysiologie et phytochimie</em>. ICSN, CNRS, 91198 Gif-sur-Yvette, Éditions Karthala, Paris, 252 pp.</p>
<p>Berger, A. (1930) Crassulaceae. Unterfam. II. Kalanchoideae [sic Kalanchooideae]. 6. <em>Kalanchoe</em>. <em>In</em>: A. Engler & K. Prantl (Eds.) <em>Die natürlichen Pflanzenfamilien</em>. ed. 2, 18a. Verlag von Wilhelm Engelmann, Leipzig, pp. 402–412.</p>
<p>Descoings, B. (2003) <em>Kalanchoe</em>. <em>In</em>: U. Eggli (Ed.) <em>Illustrated handbook of succulent plants: Crassulaceae</em>. Springer Verlag, Berlin, pp. 143–181.</p>
<p>Dorr, L.J. (1997) <em>Plant collectors in Madagascar and the Comoro Islands. A biographical and bibliographical guide to individuals and groups who have collected herbarium material of algae, bryophytes, fungi, lichens, and vascular plants in Madagascar and the Comoro Islands</em>. The Trustees, Royal Botanic Gardens, Kew, 524 pp.</p>
<p>Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). <em>Bulletin du Muséum d’Histoire Naturelle</em> 9: 35–46. [https://www.biodiversitylibrary.org/page/42893852#page/45/mode/1up]</p>
<p>Figueiredo, E. & Smith, G.F. (2017) (56) Request for a binding decision on the descriptive statement associated with <em>Kalanchoe delagoensis</em> (Crassulaceae). <em>Taxon</em> 66: 717. https://doi.org/10.12705/663.37</p>
<p>Hamet, R. (1906 [Janvier–Mars]) Note sur une nouvelle espèce de <em>Kalanchoe</em> [<em>Kalanchoe vantieghemii</em>, as ‘<em>Van Tieghemi</em>’]. <em>Journal de Botanique</em> <em>[Morot]</em> XX [20<sup>e</sup> [année] (N<sup>o</sup> 1–3)]: 109–111. [https://www.biodiversitylibrary.org/item/22106#page/113/mode/1up]</p>
<p>Hamet, R. (1907a) Monographie du genre <em>Kalanchoe</em>. <em>Bulletin de l’Herbier Boissier</em>, sér. 2, 7: 869–900. [https://www.biodiversitylibrary.org/item/105267#page/939/mode/1up]</p>
<p>Hamet, R. (1907b [Mars]) Note sur deux <em>Kalanchoe</em> malgaches. [<em>Kalanchoe grandidieri</em> Baill. and <em>K. bonnieri</em> Raym.-Hamet]. <em>Bulletin de la Société Botanique de France</em> [Tome Cinquante-Quatrième], Quatrième serie—Tome VII, 54: 138–139. [https://www.biodiversitylibrary.org/item/8676#page/178/mode/1up]</p>
<p>Hamet, R. (1908a) Monographie du genre <em>Kalanchoe</em>. Suite et fin. <em>Bulletin de l’Herbier Boissier</em>, sér. 2, 8: 17–48. [https://www.biodiversitylibrary.org/item/104945#page/31/mode/1up]</p>
<p>Hamet, R. (1908b) <em>Kalanchoe luciae</em> sp. nov. <em>Bulletin de l’Herbier Boissier</em>, sér. 2, 8: 254–257. [https://www.biodiversitylibrary.org/item/104945#page/290/mode/1up]</p>
<p>Hamet, R. (1910a [Janvier]) Sur quelques <em>Kalanchoe</em> peu connus. <em>Bulletin de la Société Botanique de France</em> [Tome Cinquante-Septième], Quatrième serie—Tome X, Quatrième series 57: 18–24. [https://www.biodiversitylibrary.org/item/8679#page/56/mode/1up]</p>
<p>Hamet, R. (1910b [Janvier]) Sur quelques <em>Kalanchoe</em> peu connus (Suite). <em>Bulletin de la Société Botanique de France</em> [Tome Cinquante-Septième], Quatrième serie—Tome X, Quatrième series 57: 49–54. [https://www.biodiversitylibrary.org/item/8679#page/87/mode/1up]</p>
<p>Hamet, R. (1910c) <em>Kalanchoe aliciae </em>sp. nova et <em>K. beharensis</em> Drake del Castillo. <em>Bulletin de la Société Botanique de France</em> 57: 191–194. [https://www.biodiversitylibrary.org/item/8679#page/241/mode/1up]</p>
<p>Hamet, R. (1912 [12 April]) Observations sur le <em>Kalanchoe tubiflora</em> nom. nov. <em>Beihefte zum Botanischen Centralblatt </em>29 (Heft 1): 41–44. [https://www.biodiversitylibrary.org/item/27107#page/523/mode/1up]</p>
<p>Harvey, W.H. (1862) Order LIII. Crassulaceae, D.C. VIII. <em>Kalanchoe</em>, Adans. <em>In</em>: W.H. Harvey & O.W. Sonder (Eds.) <em>Flora capensis</em> [<em>being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal</em>]. Vol. 2. L. Reeve & Co., Ltd, Kent, pp. 378–380. [https://www.biodiversitylibrary.org/item/15233#page/386/mode/1up]</p>
<p>Raymond-Hamet. (1915) Sur quelques <em>Kalanchoe</em> de la flore Malgache. I. <em>Bryophyllum crenatum</em> Baker, <em>Kalanchoe laxiflora</em> Baker, <em>Kalanchoe Tieghemi</em> Raymond-Hamet et <em>Kalanchoe crenata</em> Raymond-Hamet. II. <em>Kalanchoe orgyalis</em> Baker et <em>Kalanchoe antanosiana</em> Drake del Castillo. III. <em>Kitchingia porphyrocalyx</em> Baker, <em>Kalanchoe porphyrocalyx</em> Baillon et <em>Kalanchoe sulphurea</em> Baker. IV. <em>Kitchingia campanulata</em> Baker, <em>Kitchingia parviflora</em> Baker, <em>Kitchingia panduriformis</em> Baker, <em>Kitchingia amplexicaulis</em> Baker, <em>Kalanchoe campanulata</em> Baillon. <em>Kalanchoe parviflora</em> Baillon, <em>Kalanchoe panduriformis</em> Baillon et <em>Kalanchoe amplexicaulis</em> Baillon. <em>Annales du Musée Colonial de Marseille</em>, sér. 3, 3: 123–164. [https://www.biodiversitylibrary.org/item/22992#page/145/mode/1up]</p>
<p>Raymond-Hamet & Marnier-Lapostolle, J. (1964) Le genre <em>Kalanchoe</em> au Jardin Botanique “Les Cèdres”. <em>Archives du Muséum National d’Histoire Naturelle</em> Septième Série, Tome VIII: 1–110; Plates I–XXXVII. [https://www.biodiversitylibrary.org/item/280857#page/21/mode/1up]</p>
<p>Scott Elliot, G.F. (1891 [22 August]) New and little known Madagascar plants, collected and enumerated by G.F. Scott Elliot, M.A., B.Sc. F.L.S. <em>The Journal of the Linnean Society, Botany</em> XXIX(197): 1–67, with Plates I–XII. [https://www.biodiversitylibrary.org/item/8386#page/9/mode/1up]</p>
<p>Smith, G.F. (2020) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). <em>Bradleya</em> 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15</p>
<p>Smith, G.F. & Figueiredo, E. (2022) Clarifying the typification of the name <em>Kalanchoe orgyalis</em> (Crassulaceae subfam. Kalanchooideae). <em>Phytotaxa</em> 543 (1): 99–102. https://doi.org/10.11646/phytotaxa.543.1.11</p>
<p>Smith, G.F., Figueiredo, E. & Walker, C.C. (2021) <em>Kalanchoe beharensis</em>. Crassulaceae subfam. Kalanchooideae. Madagascar. <em>Flowering Plants of Africa</em> 67: 50–59, plate 2365.</p>
<p>Tulasne, L.-R. [‘Edmond’]. (1857) Fragmentum alterum (1). Floræ madagascariensis. II. <em>Calanchoe </em>[sic, <em>Kalanchoe</em>] pp. 149–150. <em>Annales des Sciences Naturelles; Botanique </em>sér. 4, 8: 44–163. [https://www.biodiversitylibrary.org/item/129817#page/153/mode/1up]</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) <em>International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. Koeltz Botanical Books, Glashütten. [Regnum Vegetabile 159], 254 pp. https://doi.org/10.12705/Code.2018</p>
<p>Baena, S., Beentje, H., Cameron, A., Lowry, P., Moat, J., Rabehevitra, D., Rahantamalala, J., Rakotoarisoa, S., Randriarimalala, Z.L., Rasolohery, A., Schatz, G., Smith, P. & Tabor, K. (2007) Atlas of the vegetation of Madagascar. / Atlas de la vegetation de Madagascar. <em>In</em>: J. Moat & P. Smith (Eds.) <em>Atlas of the vegetation of Madagascar. / Atlas de la vegetation de Madagascar</em>. Kew Publishing, Royal Botanic Gardens, Kew, Richmond, Surrey, 124 pp.</p>
<p>Baillon, M.H. (1887) Liste des plantes de Madagascar (suite de la page 624). <em>Bulletin Mensuel de la Société Linnéenne de Paris</em> 1 (No. 84): 671–672. [https://www.biodiversitylibrary.org/page/11162078#page/24/mode/1up]</p>
<p>Baillon, M.H. (1888) <em>Kalanchoe grandidieri</em>, Plate 57 [of Plates 51–88]. <em>In</em>: A. Grandidier, <em>Histoire physique, naturelle et politique Madagascar publiée par Alfred Grandidier. Vol. XXX. Histoire naturelle des plantes par M. E. Drake del Castillo</em>. Tome II – Texte – 1, partie. 50 Fascicule. Imprimé par Autorisation de M. Le Garde des Sceaux. À l’Imprimerie Nationale, Paris, 130 plates. [https://www.biodiversitylibrary.org/item/41106#page/71/mode/1up]</p>
<p>Baker, J.G. (1882) Contributions to the flora of central Madagascar (continued from p. 70 and to be continued). <em>Journal of Botany, British and Foreign. London</em> 20: 109–114. [https://www.biodiversitylibrary.org/item/35873#page/125/mode/1up]</p>
<p>Boiteau, P. & Allorge-Boiteau, L. (1995) Kalanchoe <em>(Crassulacées) de Madagascar. Systématique, écophysiologie et phytochimie</em>. ICSN, CNRS, 91198 Gif-sur-Yvette, Éditions Karthala, Paris, 252 pp.</p>
<p>Berger, A. (1930) Crassulaceae. Unterfam. II. Kalanchoideae [sic Kalanchooideae]. 6. <em>Kalanchoe</em>. <em>In</em>: A. Engler & K. Prantl (Eds.) <em>Die natürlichen Pflanzenfamilien</em>. ed. 2, 18a. Verlag von Wilhelm Engelmann, Leipzig, pp. 402–412.</p>
<p>Descoings, B. (2003) <em>Kalanchoe</em>. <em>In</em>: U. Eggli (Ed.) <em>Illustrated handbook of succulent plants: Crassulaceae</em>. Springer Verlag, Berlin, pp. 143–181.</p>
<p>Dorr, L.J. (1997) <em>Plant collectors in Madagascar and the Comoro Islands. A biographical and bibliographical guide to individuals and groups who have collected herbarium material of algae, bryophytes, fungi, lichens, and vascular plants in Madagascar and the Comoro Islands</em>. The Trustees, Royal Botanic Gardens, Kew, 524 pp.</p>
<p>Drake del Castillo, E. (1903) Note sur les plantes recueillies par M. Guillaume Grandidier dans le sud de Madagascar, en 1898 et 1901. (A suivre). <em>Bulletin du Muséum d’Histoire Naturelle</em> 9: 35–46. [https://www.biodiversitylibrary.org/page/42893852#page/45/mode/1up]</p>
<p>Figueiredo, E. & Smith, G.F. (2017) (56) Request for a binding decision on the descriptive statement associated with <em>Kalanchoe delagoensis</em> (Crassulaceae). <em>Taxon</em> 66: 717. https://doi.org/10.12705/663.37</p>
<p>Hamet, R. (1906 [Janvier–Mars]) Note sur une nouvelle espèce de <em>Kalanchoe</em> [<em>Kalanchoe vantieghemii</em>, as ‘<em>Van Tieghemi</em>’]. <em>Journal de Botanique</em> <em>[Morot]</em> XX [20<sup>e</sup> [année] (N<sup>o</sup> 1–3)]: 109–111. [https://www.biodiversitylibrary.org/item/22106#page/113/mode/1up]</p>
<p>Hamet, R. (1907a) Monographie du genre <em>Kalanchoe</em>. <em>Bulletin de l’Herbier Boissier</em>, sér. 2, 7: 869–900. [https://www.biodiversitylibrary.org/item/105267#page/939/mode/1up]</p>
<p>Hamet, R. (1907b [Mars]) Note sur deux <em>Kalanchoe</em> malgaches. [<em>Kalanchoe grandidieri</em> Baill. and <em>K. bonnieri</em> Raym.-Hamet]. <em>Bulletin de la Société Botanique de France</em> [Tome Cinquante-Quatrième], Quatrième serie—Tome VII, 54: 138–139. [https://www.biodiversitylibrary.org/item/8676#page/178/mode/1up]</p>
<p>Hamet, R. (1908a) Monographie du genre <em>Kalanchoe</em>. Suite et fin. <em>Bulletin de l’Herbier Boissier</em>, sér. 2, 8: 17–48. [https://www.biodiversitylibrary.org/item/104945#page/31/mode/1up]</p>
<p>Hamet, R. (1908b) <em>Kalanchoe luciae</em> sp. nov. <em>Bulletin de l’Herbier Boissier</em>, sér. 2, 8: 254–257. [https://www.biodiversitylibrary.org/item/104945#page/290/mode/1up]</p>
<p>Hamet, R. (1910a [Janvier]) Sur quelques <em>Kalanchoe</em> peu connus. <em>Bulletin de la Société Botanique de France</em> [Tome Cinquante-Septième], Quatrième serie—Tome X, Quatrième series 57: 18–24. [https://www.biodiversitylibrary.org/item/8679#page/56/mode/1up]</p>
<p>Hamet, R. (1910b [Janvier]) Sur quelques <em>Kalanchoe</em> peu connus (Suite). <em>Bulletin de la Société Botanique de France</em> [Tome Cinquante-Septième], Quatrième serie—Tome X, Quatrième series 57: 49–54. [https://www.biodiversitylibrary.org/item/8679#page/87/mode/1up]</p>
<p>Hamet, R. (1910c) <em>Kalanchoe aliciae </em>sp. nova et <em>K. beharensis</em> Drake del Castillo. <em>Bulletin de la Société Botanique de France</em> 57: 191–194. [https://www.biodiversitylibrary.org/item/8679#page/241/mode/1up]</p>
<p>Hamet, R. (1912 [12 April]) Observations sur le <em>Kalanchoe tubiflora</em> nom. nov. <em>Beihefte zum Botanischen Centralblatt </em>29 (Heft 1): 41–44. [https://www.biodiversitylibrary.org/item/27107#page/523/mode/1up]</p>
<p>Harvey, W.H. (1862) Order LIII. Crassulaceae, D.C. VIII. <em>Kalanchoe</em>, Adans. <em>In</em>: W.H. Harvey & O.W. Sonder (Eds.) <em>Flora capensis</em> [<em>being a systematic description of the plants of the Cape Colony, Caffraria, & Port Natal</em>]. Vol. 2. L. Reeve & Co., Ltd, Kent, pp. 378–380. [https://www.biodiversitylibrary.org/item/15233#page/386/mode/1up]</p>
<p>Raymond-Hamet. (1915) Sur quelques <em>Kalanchoe</em> de la flore Malgache. I. <em>Bryophyllum crenatum</em> Baker, <em>Kalanchoe laxiflora</em> Baker, <em>Kalanchoe Tieghemi</em> Raymond-Hamet et <em>Kalanchoe crenata</em> Raymond-Hamet. II. <em>Kalanchoe orgyalis</em> Baker et <em>Kalanchoe antanosiana</em> Drake del Castillo. III. <em>Kitchingia porphyrocalyx</em> Baker, <em>Kalanchoe porphyrocalyx</em> Baillon et <em>Kalanchoe sulphurea</em> Baker. IV. <em>Kitchingia campanulata</em> Baker, <em>Kitchingia parviflora</em> Baker, <em>Kitchingia panduriformis</em> Baker, <em>Kitchingia amplexicaulis</em> Baker, <em>Kalanchoe campanulata</em> Baillon. <em>Kalanchoe parviflora</em> Baillon, <em>Kalanchoe panduriformis</em> Baillon et <em>Kalanchoe amplexicaulis</em> Baillon. <em>Annales du Musée Colonial de Marseille</em>, sér. 3, 3: 123–164. [https://www.biodiversitylibrary.org/item/22992#page/145/mode/1up]</p>
<p>Raymond-Hamet & Marnier-Lapostolle, J. (1964) Le genre <em>Kalanchoe</em> au Jardin Botanique “Les Cèdres”. <em>Archives du Muséum National d’Histoire Naturelle</em> Septième Série, Tome VIII: 1–110; Plates I–XXXVII. [https://www.biodiversitylibrary.org/item/280857#page/21/mode/1up]</p>
<p>Scott Elliot, G.F. (1891 [22 August]) New and little known Madagascar plants, collected and enumerated by G.F. Scott Elliot, M.A., B.Sc. F.L.S. <em>The Journal of the Linnean Society, Botany</em> XXIX(197): 1–67, with Plates I–XII. [https://www.biodiversitylibrary.org/item/8386#page/9/mode/1up]</p>
<p>Smith, G.F. (2020) The kalanchoes (Crassulaceae subfam. Kalanchooideae) of Raymond-Hamet (1890–1972). <em>Bradleya</em> 38: 120–140. https://doi.org/10.25223/brad.n38.2020.a15</p>
<p>Smith, G.F. & Figueiredo, E. (2022) Clarifying the typification of the name <em>Kalanchoe orgyalis</em> (Crassulaceae subfam. Kalanchooideae). <em>Phytotaxa</em> 543 (1): 99–102. https://doi.org/10.11646/phytotaxa.543.1.11</p>
<p>Smith, G.F., Figueiredo, E. & Walker, C.C. (2021) <em>Kalanchoe beharensis</em>. Crassulaceae subfam. Kalanchooideae. Madagascar. <em>Flowering Plants of Africa</em> 67: 50–59, plate 2365.</p>
<p>Tulasne, L.-R. [‘Edmond’]. (1857) Fragmentum alterum (1). Floræ madagascariensis. II. <em>Calanchoe </em>[sic, <em>Kalanchoe</em>] pp. 149–150. <em>Annales des Sciences Naturelles; Botanique </em>sér. 4, 8: 44–163. [https://www.biodiversitylibrary.org/item/129817#page/153/mode/1up]</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) <em>International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. Koeltz Botanical Books, Glashütten. [Regnum Vegetabile 159], 254 pp. https://doi.org/10.12705/Code.2018</p>