Abstract
The taxonomic limits of two Moroccan Arabis species are revised based on a critical evaluation of morphology in light of phylogenetic and cytogenetic studies that also included the monospecific European Fourraea. As a result, the new tribe Fourraeeae, the new genus Hurkaea and new combinations H. josiae, H. conringioides, and H. conringioides subsp. humbertii are proposed. Keys to distinguish genera, species and subspecies are provided.
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<p>Ball, J. (1873) Descriptions of some new species, subspecies, and varieties of plants collected in Morocco by J. D. Hooker, G. Maw, and J. Ball. <em>Journal of Botany</em> 11: 296–307.</p>
<p>Burdet, H.M. (1967) Contribution à l’étude caryologique des genres <em>Cardaminopsis</em>, <em>Turritis</em> et <em>Arabis</em> en Europe. <em>Candollea</em> 22 (1): 107–156.</p>
<p>Candolle, A.P. de. (1821) Mémoire sur la famille des Crucifères. <em>Mémoires du Muséum d’Histoire Naturelle</em> 7: 169–252.</p>
<p>Couvreur, T.L.P., Franzke, A., Al-Shehbaz, I.A., Bakker, F.T., Koch, M.A. & Mummenhoff, K. (2010) Molecular phylogenetics, temporal diversification and principles of evolution in the mustard family (Brassicaceae). <em>Molecular Biology and Evolution</em> 27 (1): 55–71. https://doi.org/10.1093/molbev/msp202</p>
<p>Darwin, C. (1859) <em>On the origin of species by means of natural selection</em>. John Murray, London, 502 pp.</p>
<p>Dersch, G. (1968) Über einige Chromosomenzählungen an mitteleuropäischen Blütenpflanzen. <em>Abhandlungen und Berichte des Vereins für Naturkunde Kassel</em> 62 (4): 9–12.</p>
<p>Dobignard, A. & Jordan, D. (1987) Nouvelles observations sur la flore du Maroc. 1. Contribution à l’étude de la flore du Haut-Atlas. <em>Saussurea</em> 18: 67–104.</p>
<p>Dobignard, A., Chatelain, C., Fischer, M., Orso, J. & Jeanmonod, D. (2011) <em>Index synonymique de la flore d’Afrique du Nord</em> 3, Dicotyledoneae: Balsaminaceae-Euphorbiaceae. Conservatoire et Jardin botaniques de la Ville de Genève, Geneve, 449 pp.</p>
<p>Font Quer, P. (1928) <em>Iter Maroccanum 1927. Exsiccata schedula</em>. Natural Science Museum, Barcelona, 739 num.</p>
<p>Font Quer, P. (1929) <em>Iter Maroccanum 1928. Exsiccata schedula</em>. Natural Science Museum, Barcelona, 438 num.</p>
<p>Fourreau, J. (1868) Catalogue des plantes qui croissent spontanément le long du Rhône. <em>Annales de la Société Linnéenne de Lyon</em> sér. 2, 16: 301–404. https://doi.org/10.5962/bhl.title.12050</p>
<p>Galland, N. (1984) [Report LXXXV/III]. Chromosome number reports LXXXV. <em>Taxon</em> 33(4): 756–760. https://doi.org/10.1002/j.1996-8175.1984.tb04374.x</p>
<p>Galland, N. (1988) Recherche sur l’origine de la flore orophile du Maroc: étude caryologique et cytogéographique. <em>Travaux de l’Institut Scientifique, Université Mohammed V, Sér. Botanique</em> 35: 1–168.</p>
<p>Galland, N. & Favarger, C. (1985) Notes de taxonomie et de nomenclature sur la flore orophile du Maroc. <em>Candollea</em> 40 (1): 231–235. https://doi.org/10.5169/seals-879783</p>
<p>Garcke, A. (1858) <em>Flora von Nord- und Mittel-Deutschland</em>. Ed. 4. Gustav Bosselmann, Berlin, 428 pp.</p>
<p>German, D.A., Friesen, N., Neuffer, B., Al-Shehbaz I.A. & Hurka, H. (2009) Contribution to ITS phylogeny of the Brassicaceae, with special reference to some Asian taxa. <em>Plant Systematics and Evolution</em> 283 (1): 33–56. https://doi.org/10.1007/s00606-009-0213-5</p>
<p>Greuter, W. & Raus, T. (Eds.) (1984) Med-Checklist notulae, 8. <em>Willdenowia</em> 13 (2): 283–288.</p>
<p>Grimm, J.F.C. (1767) Synopsis methodica stirpium agri isenacensis consignata. <em>Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum</em> 3 App.: 249–364.</p>
<p>Hall, J.C., Tisdale, T.E., Donohue, K., Wheeler, A., Al-Yahya, M.A. & Kramer, E.M. (2011) Convergent evolution of a complex fruit structure in the tribe Brassiceae (Brassicaceae). <em>American Journal of Botany</em> 98 (12): 1989–2003. https://doi.org/10.3732/ajb.1100203</p>
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