Abstract
The heterotypic synonyms Verbascum limnense and Celsia tomentosa are neotypified and lectotypified, respectively, and their nomenclature is clarified. The former binomial represents the correct name at species rank for a range-restricted, paleoendemic plant confined to Central Greece. Furthermore, the new nothospecies Verbascum × sipiadense, representing the hybrid formula V. limnense × V. sinuatum, is described from the island of Evvia (Euboea).
References
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<p>Boissier, P.E. (1856) <em>Diagnoses plantarum Orientalium novarum ser. 2</em>, 3. B. Herrmann, Leipzig, 177 pp.</p>
<p>Boissier, P.E. (1879) <em>Flora Orientalis </em>4(2). H. Georg, Genevae [Genève], Basileae [Basel] & Lugduni [Lyon], pp. 281–1276.</p>
<p>Dimopoulos, P., Raus, Th., Bergmeier, E., Constantinidis, Th., Iatrou, G., Kokkini, S., Strid, A. & Tzanoudakis, D. (2013) <em>Vascular plants of Greece: An annotated checklist</em>. Botanischer Garten und Botanisches Museum Berlin-Dahlem, Berlin; Hellenic Botanical Society, Athens, 372 pp.</p>
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<p>Ferguson, I.K. (1972) <em>Verbascum </em>L. <em>In</em>: Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (Eds.) <em>Flora Europaea</em> 3. Cambridge University Press, Cambridge, pp. 205–216.</p>
<p>Fischer, E. (1997) Notulae ad Floram Germanicam II. Typifications of Linnean names of Scrophulariaceae occurring in Germany. <em>Feddes Repertorium</em> 108: 111–117. https://doi.org/10.1002/fedr.19971080110</p>
<p>Fraas, C.N. (1845) <em>Synopsis plantarum florae classicae</em>. E.A. Fleischmann, München, 320 pp.</p>
<p>Hertel, H. & Schreiber, A. (1988) Die Botanische Staatssammlung München 1813–1988 (Eine Übersicht über die Sammlungsbestände). <em>Mitteilungen der Botanischen Staatssammlung München</em> 26: 81–512. Available from: http://indexcoll.botanischestaatssammlung.de/ (accessed 6 April 2021)</p>
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<p>Halácsy, E. von (1902) <em>Conspectus florae Graecae</em> 2. W. Engelmann, Lipsiae [Leipzig], 612 pp.</p>
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<p>Huber-Morath, A. & Rechinger, K.H. (1960) Zur Kenntnis der Gattungen <em>Verbascum</em> und <em>Celsia</em> in Griechenland. <em>Mitteilungen des Thüringischen Botanischen Vereins</em> 2: 42–55.</p>
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<p>Lamarck, J.B.P.A.M. (1779) [“1778”] <em>Flore françoise</em> 2. L’Imprimerie Royale, Paris, 684 pp.</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum</em>. Impensis Laurentii Salvii, Holmiae [Stockholm], 1200 pp.</p>
<p>Murbeck, S.S. (1925) <em>Monographie der Gattung Celsia</em>. C.W.K. Gleerup, Lund, 238 pp.</p>
<p>Murbeck, S.S. (1933) <em>Monographie der Gattung Verbascum</em>. H. Ohlsson, Lund, 630 pp.</p>
<p>Murbeck, S.S. (1939) <em>Weitere Studien über die Gattungen Verbascum und Celsia</em>. C.W.K. Gleerup, Lund, 70 pp.</p>
<p>Nyman, C.F. (1881) <em>Conspectus Florae Europaeae. </em>3. Typis Officinae Bohlinianae, Örebro Sueciae, pp. 493–677.</p>
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<p>Rechinger, K.H. (1944) [“1943”] Flora Aegaea. Flora der Inseln und Halbinseln des Ägäischen Meeres. <em>Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, Denkschriften</em> 105 (1): 1–924 + tab. I–XXV.</p>
<p>Rechinger, K.H. (1961) Die Flora von Euboea. I. Teil. <em>Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie </em>80: 294–382.</p>
<p>Strid, A. (2016a) Atlas of the Aegean flora, Part 1: Texts and Plates. <em>Englera </em>33 (1): 1–700.</p>
<p>Strid, A. (2016b) Atlas of the Aegean flora, Part 2: Maps. <em>Englera </em>33 (2): 1–878.</p>
<p>Trigas, P. (2003) <em>Contribution to the study of the endemism of the flora of the island of Evvia.</em> PhD Thesis, University of Patras, Greece, 341 pp. [in Greek with an English summary].</p>
<p>Turland, N.J. (2019) <em>The Code Decoded</em>—<em>A user’s guide to the International Code of Nomenclature for algae, fungi, and plants, Second edition</em>. Pensoft, Sofia, 196 pp. https://doi.org/10.3897/ab.e38075</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (Eds.) (2018) <em>International Code of Nomenclature for</em> <em>algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. Regnum Vegetabile 159. Koeltz Botanical Books, Glashütten, 254 pp. https://doi.org/10.12705/Code.2018</p>
<p>Zografidis, A. (2017) New floristic records in Greece. <em>Parnassiana Archives</em> 5: 61–66.</p>
<p>Zuccarini, J.G. (1837) Plantarum novarum vel minus cognitarum, quae in horto botanico herbarioque regio Monacensi servantur, fasc. II. <em>Abhandlungen der Mathematisch-Physikalischen Klasse der Königlich Bayerischen Akademie der Wissenschaften </em>2: 309–380.</p>
<p>Boissier, P.E. (1856) <em>Diagnoses plantarum Orientalium novarum ser. 2</em>, 3. B. Herrmann, Leipzig, 177 pp.</p>
<p>Boissier, P.E. (1879) <em>Flora Orientalis </em>4(2). H. Georg, Genevae [Genève], Basileae [Basel] & Lugduni [Lyon], pp. 281–1276.</p>
<p>Dimopoulos, P., Raus, Th., Bergmeier, E., Constantinidis, Th., Iatrou, G., Kokkini, S., Strid, A. & Tzanoudakis, D. (2013) <em>Vascular plants of Greece: An annotated checklist</em>. Botanischer Garten und Botanisches Museum Berlin-Dahlem, Berlin; Hellenic Botanical Society, Athens, 372 pp.</p>
<p>Ferguson, I.K. (1971) Notes on the genus <em>Verbascum</em> (Scrophulariaceae). Flora Europaea Notulae Systematicae ad Floram Europaeam spectantes No. 9. <em>Botanical Journal of the Linnean Society </em>64: 229–233. https://doi.org/10.1111/j.1095-8339.1971.tb02145.x</p>
<p>Ferguson, I.K. (1972) <em>Verbascum </em>L. <em>In</em>: Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M. & Webb, D.A. (Eds.) <em>Flora Europaea</em> 3. Cambridge University Press, Cambridge, pp. 205–216.</p>
<p>Fischer, E. (1997) Notulae ad Floram Germanicam II. Typifications of Linnean names of Scrophulariaceae occurring in Germany. <em>Feddes Repertorium</em> 108: 111–117. https://doi.org/10.1002/fedr.19971080110</p>
<p>Fraas, C.N. (1845) <em>Synopsis plantarum florae classicae</em>. E.A. Fleischmann, München, 320 pp.</p>
<p>Hertel, H. & Schreiber, A. (1988) Die Botanische Staatssammlung München 1813–1988 (Eine Übersicht über die Sammlungsbestände). <em>Mitteilungen der Botanischen Staatssammlung München</em> 26: 81–512. Available from: http://indexcoll.botanischestaatssammlung.de/ (accessed 6 April 2021)</p>
<p>Ghahremaninejad, F., Riahi, M., Babaei, M., Attar, F., Beh¸cet, L. & Sonboli, A. (2014) Monophyly of <em>Verbascum</em> (Scrophularieae: Scrophulariaceae): evidence from nuclear and plastid phylogenetic analyses. <em>Australian Journal of Botany</em> 62: 638–646. https://doi.org/10.1071/BT14159</p>
<p>Goerdten, U. (2021) Carl Fraas. <em>In</em>: <em>Wikipedia, Die freie Enzyklopädie</em>. Version: 13 May 2021. Available from: https://de.wikipedia.org/w/index.php?title=Carl_Fraas&oldid=211922493 (accessed 28 May 2021)</p>
<p>Halácsy, E. von (1898) Die bisher bekannten <em>Verbascum</em>-Arten Griechenlands. <em>Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien</em> 48: 119–150.</p>
<p>Halácsy, E. von (1902) <em>Conspectus florae Graecae</em> 2. W. Engelmann, Lipsiae [Leipzig], 612 pp.</p>
<p>Hayek, A. von (1929) Prodromus Florae Peninsulae Balcanicae 2(2). <em>Repertorium Specierum Novarum Regni Vegetabilis Beihefte </em>30: 97–240.</p>
<p>Huber-Morath, A. (1979) Novitiae Florae Anatolicae XIII. <em>Bauhinia</em> 6: 369–374.</p>
<p>Huber-Morath, A. & Rechinger, K.H. (1960) Zur Kenntnis der Gattungen <em>Verbascum</em> und <em>Celsia</em> in Griechenland. <em>Mitteilungen des Thüringischen Botanischen Vereins</em> 2: 42–55.</p>
<p>Kalheber, H. (2006) Bavarian plant collectors in Greece. 1. Franz Xaver Berger, Franz Zuccarini and Carl Nikolaus Fraas. <em>Willdenowia</em> 36: 565–578. https://doi.org/10.3372/wi.36.36153</p>
<p>Kuntze, O. (1891) <em>Revisio generum plantarum</em> 2. A. Felix, Leipzig [etc.], pp. 375–1011.</p>
<p>Lamarck, J.B.P.A.M. (1779) [“1778”] <em>Flore françoise</em> 2. L’Imprimerie Royale, Paris, 684 pp.</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum</em>. Impensis Laurentii Salvii, Holmiae [Stockholm], 1200 pp.</p>
<p>Murbeck, S.S. (1925) <em>Monographie der Gattung Celsia</em>. C.W.K. Gleerup, Lund, 238 pp.</p>
<p>Murbeck, S.S. (1933) <em>Monographie der Gattung Verbascum</em>. H. Ohlsson, Lund, 630 pp.</p>
<p>Murbeck, S.S. (1939) <em>Weitere Studien über die Gattungen Verbascum und Celsia</em>. C.W.K. Gleerup, Lund, 70 pp.</p>
<p>Nyman, C.F. (1881) <em>Conspectus Florae Europaeae. </em>3. Typis Officinae Bohlinianae, Örebro Sueciae, pp. 493–677.</p>
<p>POWO (2019, continuously updated) <em>Plants of the World Online</em>. The Royal Botanic Gardens, Kew. Available from: http://www.plantsoftheworldonline.org/ (accessed 5 April 2021)</p>
<p>Rechinger, K.H. (1944) [“1943”] Flora Aegaea. Flora der Inseln und Halbinseln des Ägäischen Meeres. <em>Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, Denkschriften</em> 105 (1): 1–924 + tab. I–XXV.</p>
<p>Rechinger, K.H. (1961) Die Flora von Euboea. I. Teil. <em>Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie </em>80: 294–382.</p>
<p>Strid, A. (2016a) Atlas of the Aegean flora, Part 1: Texts and Plates. <em>Englera </em>33 (1): 1–700.</p>
<p>Strid, A. (2016b) Atlas of the Aegean flora, Part 2: Maps. <em>Englera </em>33 (2): 1–878.</p>
<p>Trigas, P. (2003) <em>Contribution to the study of the endemism of the flora of the island of Evvia.</em> PhD Thesis, University of Patras, Greece, 341 pp. [in Greek with an English summary].</p>
<p>Turland, N.J. (2019) <em>The Code Decoded</em>—<em>A user’s guide to the International Code of Nomenclature for algae, fungi, and plants, Second edition</em>. Pensoft, Sofia, 196 pp. https://doi.org/10.3897/ab.e38075</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (Eds.) (2018) <em>International Code of Nomenclature for</em> <em>algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. Regnum Vegetabile 159. Koeltz Botanical Books, Glashütten, 254 pp. https://doi.org/10.12705/Code.2018</p>
<p>Zografidis, A. (2017) New floristic records in Greece. <em>Parnassiana Archives</em> 5: 61–66.</p>
<p>Zuccarini, J.G. (1837) Plantarum novarum vel minus cognitarum, quae in horto botanico herbarioque regio Monacensi servantur, fasc. II. <em>Abhandlungen der Mathematisch-Physikalischen Klasse der Königlich Bayerischen Akademie der Wissenschaften </em>2: 309–380.</p>