A new variety of fern from Borneo, Sphaerostephanos unitus var. dimorphophylla (Thelypteridaceae)

TAKEO HAYASHI; MONICA SULEIMAN; HIROSHI OKADA; HIROKAZU TSUKAYA

doi:10.11646/phytotaxa.346.3.8

Issue: Vol. 346 No. 3: 6 April 2018

Type: Article

Published: 2018-04-06

DOI: 10.11646/phytotaxa.346.3.8

Page range: 287–292

Abstract views: 20

PDF downloaded: 1

**A new variety of fern from Borneo, Sphaerostephanos unitus var. dimorphophylla (Thelypteridaceae)**

TAKEO HAYASHI⁺⁻
MONICA SULEIMAN⁺⁻
HIROSHI OKADA⁺⁻
HIROKAZU TSUKAYA⁺⁻

Department of Biological Sciences, Graduate School of Science, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan

Institute for Tropical Biology and Conservation Universiti Malaysia Sabah, Jalan UMS, 88400 Kota Kinabalu, Sabah, Malaysia

Graduate School of Science, Osaka City University, Sugimoto, Sumiyoshi-ku, Osaka 558-8585, Japan Institute of Natural Environmental Sciences, Hyogo Prefectural University, Sanda, Hyogo 669-1546, Japan

Department of Biological Sciences, Graduate School of Science, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan Bio-Next Project, Okazaki Institute for Integractive Bioscience, National Institutes of Natural Sciences, Yamate Build. #3, 5-1, Higashiyama, Myodaiji, Okazaki, Aichi 444-8787, Japan

Borneo fern Imbak Canyon Malaysia rheophyte Sabah Sphaerostephanos taxonomy Thelypteridaceae Pteridophyters

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Abstract

The genus Sphaerostephanos is a sub-group of Thelypteridaceae. We collected an undescribed taxon of this genus in Imbak Canyon, Sabah, Borneo. We describe this specimen as a new variety with a drawing and key of this genus.