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Type: Article
Published: 2023-10-11
Page range: 255-276
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New orchids in the flora of Vietnam VII (Orchidaceae: tribes Cypripedieae, Cranichideae, Orchideae, and Collabieae)

Komarov Botanical Institute of the Russian Academy of Sciences; 2; Prof. Popov Street; 197376 St. Petersburg; Russia
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LEONID V. AVERYANOV

 

 

Institute of Applied Technology; Thu Dau Mot University; No. 6; Tran Van On Street; Phu Hoa Ward; Thu Dau Mot City; Binh Duong Province; Vietnam
Institute of Tropical Biology; Vietnam Academy of Science and Technology; 85; Tran Quoc Toan; District 3; Ho Chi Minh City; Vietnam
Institute of Ecology and Biological Resources; Vietnam Academy of Science and Technology; 18; Hoang Quoc Viet Road; Cau Giay; Hanoi; Vietnam. Graduate University of Science and Technology; Vietnam Academy of Science and Technology; 18; Hoang Quoc Viet Road; Cau Giay; Hanoi; Vietnam
Department of Higher Plants; Biological Faculty; M.V. Lomonosov Moscow State University; 1; 12; Leninskie Gory; 119234 Moscow; Russia. Joint Russian-Vietnamese Tropical Scientific and Technological Center; 63; Nguyen Van Huyen Street; Cau Giay; 122000 Hanoi; Vietnam
Vietnam National University of Forestry; Xuan Mai; Chuong My; Hanoi; Vietnam
P.O. Box 8210; Cairns 4870; Queensland; Australia
Komarov Botanical Institute of the Russian Academy of Sciences; 2; Prof. Popov Street; 197376 St. Petersburg; Russia
Institute of Applied Technology, Thu Dau Mot University, No. 6, Tran Van On Street, Phu Hoa Ward, Thu Dau Mot City, Binh Duong Province, Vietnam
Department of Higher Plants, Biological Faculty, M.V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234 Moscow, Russia.
Tay Nguyen Institute for Scientific Research, Vietnam Academy of Science and Technology, Dalat, Vietnam.
endemism new nomenclatural combinations new records new species new synonyms plant diversity plant geography plant taxonomy monocots

Abstract

The seventh paper of the series on orchid diversity in Vietnam includes descriptions of four species new to science (Goodyera umbonata, Myrmechis brachyscapa, Rhomboda obcordata, and Vietorchis proboscidea) and five new national records (Calanthe speciosa, Habenaria coultousii, Myrmechis urceolata, Paphiopedilum barbigerum var. barbigerum, and Peristylus affinis). Two new nomenclatural combinations, Rhomboda aurea (≡ Odontochilus aureus) and Odontochilus tonkinensis (≡ Anoectochilus tonkinensis), are proposed. Odontochilus putaoensis described from Myanmar and reported also from China and Laos is synonymized with Odontochilus umbrosus described 22 years earlier from Vietnam. Data on relevant synonyms and etymology are provided, along with morphological descriptions for the newly described species. For the species new to science and for the taxa reported for Vietnam for the first time, the information is given about studied specimens, ecology, phenology, distribution, and conservation status assessed according to the IUCN Red List Categories and Criteria, accompanied by brief taxonomic notes and illustrations.

 

References

  1. Aung, Y.L., Mu, A.T. & Jin, X. (2018) Odontochilus putaoensis (Cranichideae, Orchidaceae), a new species from Myanmar. PhytoKeys 103: 19–26. https://doi.org/10.3897/phytokeys.103.25913
  2. Averyanov, L.V. (1990) Fam. Orchidaceae. In: Tzvelev, N.N. (Ed.) Vascular plants synopsis of Vietnamese flora 1. Nauka, Leningrad, pp. 20–167.
  3. Averyanov, L.V. (1996) New species of orchids (Orchidaceae) from Vietnam. Botanical Journal (St.-Petersburg) 81 (10): 73–83.
  4. Averyanov, L.V. (2006) Rare species of orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 9 (3): 48–89.
  5. Averyanov, L.V. (2007) New species of orchids from Vietnam. Taiwania 52 (4): 287–306. https://doi.org/10.6165/tai.2007.52(4).287
  6. Averyanov, L. (2008) The orchids of Vietnam. Illustrated survey. Part 1. Subfamilies Apostasioideae, Cypripedioideae and Spiranthoideae. Turczaninowia 11 (1): 5–168.
  7. Averyanov, L.V. (2011) The orchids of Vietnam. Illustrated survey. Part 3. Subfamily Epidendroideae (primitive tribes—Neottieae, Vanilleae, Gastrodieae, Nervilieae). Turczaninowia 14 (2): 15–100.
  8. Averyanov, L.V. & Averyanova, A.L. (2002) Rare species of orchids (Orchidaceae) in the flora of Vietnam. Turczaninowia 5 (4): 49–108.
  9. Averyanov, L.V. & Averyanova, A.L. (2003) Updated checklist of the orchids of Vietnam. Vietnam National University Publishing House, Hanoi, 102 pp.
  10. Averyanov, L., Cribb, P., Loc, P.K. & Nguyen, T.H. (2003a) Slipper orchids of Vietnam. With an introduction to the flora of Vietnam. Compass Press Limited, Kew, 308 pp.
  11. Averyanov, L.V., Loc, P.K., Hiep, N.T. & Harder, D.K. (2003b) Phytogeographic review of Vietnam and adjacent areas of Eastern Indochina. Komarovia 3: 1–83.
  12. Averyanov, L.V., Nuraliev, M.S., Kuznetsov, A.N. & Kuznetsova, S.P. (2013) Vietorchis furcata (Orchidaceae, Vietorchidinae)—a new species from Southern Vietnam. Taiwania 58 (4): 251–256. https://doi.org/10.6165/tai.2013.58.251
  13. Averyanov, L.V., Nguyen, K.S., Nguyen, T.T., Nguyen, P.T., Nong, V.D., Nguyen, V.C. & Chu, C.X. (2015) New orchids in the flora of Vietnam. Wulfenia 22: 137–188.
  14. Averyanov, L.V., Nguyen, K.S., Maisak, T.V., Nguyen, H.T., Orlov, N.L., Slastunov, D.D. & Sokolova, I.V. (2019a) New orchid species (Orchidaceae) in the flora of Laos. Novitates Systematicae Plantarum Vascularium 50: 25–40. https://doi.org/10.31111/novitates/2019.50.25
  15. Averyanov, L.V., Nguyen, V.C., Nguyen, K.S., Maisak, T.V. & Truong, B.V. (2019b) New orchids (Orchidaceae) in the flora of Vietnam I. Epidendroideae. Taiwania 64 (2): 176–188. https://doi.org/10.6165/tai.2019.64.176
  16. Averyanov, L.V., Truong, B.V., Nguyen, V.C., Nguyen, K.S. & Maisak, T.V. (2019c) New orchids (Orchidaceae) in the flora of Vietnam II. Vandeae. Taiwania 64 (3): 285–298. https://doi.org/10.6165/tai.2019.64.285
  17. Averyanov, L.V., Truong, B.V., Nguyen, V.C., Maisak, T.V., Dinh, Q.D., Nuraliev, M.S., Nguyen, K.S. & Chu, V.T. (2020) New orchids in the flora of Vietnam III (Collabieae, Malaxideae, Nervilieae and Orchideae). Taiwania 65 (4): 478‒492. https://doi.org/10.6165/tai.2020.65.478
  18. Averyanov, L.V., Thai, T.H., Truong, B.V., Nguyen, V.C., Nguyen, T.H., Maisak, T.V., Nguyen, K.S. & Nguyen, V.K. (2022a) New orchids in the flora of Vietnam IV (Orchidaceae, Aeridinae). Phytotaxa 555 (2): 113–135. https://doi.org/10.11646/phytotaxa.555.2.1
  19. Averyanov, L.V., Thai, T.H., Nguyen, V.C., Truong, B.V., Maisak, T.V., Doan, N.T. & Nguyen, K.S. (2022b) New orchids in the flora of Vietnam V (Orchidaceae, Aeridinae). Phytotaxa 572 (1): 43–60. https://doi.org/10.11646/phytotaxa.572.1.3
  20. Averyanov, L.V., Nguyen, V.C., Truong, B.V., Nguyen, K.S., Nguyen, C.H., Maisak, T.V., Doan, N.T., Nguyen, T.H., Pham, V.T., Thanh, P.T., Thai, T.H., Nguyen, V.K. & Trinh, N.B. (2023) New orchids in the flora of Vietnam VI (Orchidaceae, tribes Arethuseae, Cymbidieae, Diurideae, Epidendreae, Vandeae, and Vanilleae). Phytotaxa 597 (2): 87–110. https://doi.org/10.11646/phytotaxa.597.2.1
  21. Barretto, G.D’A. (1981) An addition to the orchid flora of Hong Kong. Orchadian 7: 10–13.
  22. Barretto, G., Cribb, P. & Gale, S. (2011) The wild orchids of Hong Kong. Natural History Publications (Borneo), Kota Kinabalu, 697 pp.
  23. Blume, C.L. (1825) Bijdragen tot de flora van Nederlandsch Indië. Ter Lands Drukkerij, Batavia, 1169 pp.
  24. Blume, C.L. (1858) Collection des orchideìes les plus remarquables de l’Archipel indien et du Japon. C.G. Sulpke, Amsterdam, 190 pp.
  25. Braem, G.J. (1988) Paphiopedilum. Brucke-Verlag Kurt Schmersow, Hildesheim, 249 pp.
  26. Braem, G.J., Baker, C.O. & Baker, M.L. (1999) The genus Paphiopedilum—natural history and cultivation. Vol. 2. Botanical Publishers Inc., Kissimmee, 185–363 pp.
  27. Chase, M.W., Cameron, K.M., Freudenstein, J.V., Pridgeon, A.M., Salazar, G., van den Berg, C. & Schuiteman, A. (2015) An updated classification of Orchidaceae. Botanical Journal of the Linnean Society 177 (2): 151–174. https://doi.org/10.1111/boj.12234
  28. Cheeseman, T.F. (1903) The flora of Rarotonga, the chief island of the Cook Group. Transactions of the Linnean Society of London 2, Botany, 6: 261–313. https://doi.org/10.1111/j.1095-8339.1903.tb00277.x
  29. Chen, S.C., Liu, Z.J., Zhu, G.H., Lang, K.Y., Ji, Z.H., Luo, Y.B., Jin, X.H., Cribb, P.J., Wood, J.J., Gale, S.W., Ormerod, P., Vermeulen, J.J., Wood, H.P., Clayton, D. & Bell, A. (2009) Orchidaceae. In: Wu, Z.Y., Raven, P.H. & Hong, D. (Eds.) Flora of China, vol. 25. Science Press, Beijing & Missouri Botanical Garden Press, St. Louis, pp. 211–235.
  30. Clayton, D. & Cribb, P. (2013) The genus Calanthe. Natural History Publications (Borneo), Kota Kinabalu, 411 pp.
  31. Comber, J.B. (1990) Orchids of Java. Bentham-Moxon Trust, Kew, 407 pp.
  32. Comber, J.B. (2001) Orchids of Sumatra. The Royal Botanic Gardens, Kew, 1026 pp.
  33. Cribb, P. (1998) The genus Paphiopedilum. Natural History Publications, Kota Kinabalu, 427 pp.
  34. Cribb, P.C. & Whistler, W.A. (2011) The orchids of Tonga, Niue, and The Cook Islands. Lankesteriana 11 (2): 99–177.
  35. Don, D. (1825) Prodromus florae Nepalensis. J. Gale, London, 256 pp.
  36. Downie, D.G. (1925) Contributions to flora of Siam. Additamentum XVII. Bulletin of Miscellaneous Information (Royal Botanic Gardens, Kew) 1925 (10): 404–423. https://doi.org/10.2307/4115102
  37. Fukuyama, N. (1934) Studia orchidacearum Japonicarum III. Orchidaceae Formosanae novae atque criticae. Botanical Magazine (Tokyo) 48 (571): 429–442. https://doi.org/10.15281/jplantres1887.48.429
  38. Fukuyama, N. (1936) Studia orchidacearum Japonicarum. VIII. Orchidaceae Formosae et Insulae Kôtô-syo novae atque criticae. Botanical Magazine (Tokyo) 50 (589): 16–24. https://doi.org/10.15281/jplantres1887.50.16
  39. Gagnepain, F. (1931a) Dix Orchidacées nouvelles d’Asie. Bulletin du Museìum National d’Histoire Naturelle 2, 3 (3): 322–328.
  40. Gagnepain, F. (1931b) Treize Orchidacées nouvelles d’Indo-Chine. Bulletin du Museìum National d’Histoire Naturelle 2, 3 (7): 679–687.
  41. Gagnepain, F. (1933) Espèces nouvelles de Marantacées, Orchidacées, Apostasiacées. Bulletin de la Société Botanique de France 80 (3): 348–352. https://doi.org/10.1080/00378941.1933.10833847
  42. Govaerts, R., Bernet, P., Kratochvil, K., Gerlach, G., Carr, G., Alrich, P., Pridgeon, A.M., Pfahl, J., Campacci, M.A., Holland Baptista, D., Tigges, H., Shaw, J., Cribb, P.J., George, A., Kreuz, K. & Wood, J. (2023) Orchidaceae. POWO: Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet. Available from: http://www.plantsoftheworldonline.org/ (accessed: 16 June 2023).
  43. Hooker, J.D. (1890) Orchidaceae. In: Hooker, J.D. (Ed.) The flora of British India 6. Reeve, London, pp. 1–198.
  44. Hu, S.Y. & Kong, S.L. (2011) 389. Orchidaceae. In: Hu, Q.M. & Wu, D.L. (Eds.) Flora of Hong Kong. Vol. 4. Agriculture, Fisheries and Conservation Department, Government of the Hong Kong Special Administrative Region, Hong Kong, pp. 294–352.
  45. IUCN Standards and Petitions Committee (2023) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from: https://www.iucnredlist.org/documents/RedListGuidelines.pdf (accessed on 16 June 2023).
  46. Jalal, J.S. & Jayanthi, J. (2015) An annotated checklist of the orchids of western Himalaya, India. Lankesteriana 15 (1): 7–50. https://doi.org/10.15517/lank.v15i1.18521
  47. King, G. & Pantling, R. (1896) A second series of new orchids from Sikkim. The Journal of the Asiatic Society of Bengal 65 (2): 118–134.
  48. King, G. & Pantling, R. (1898) The orchids of Sikkim Himalaya. Annals Royal Botanic Garden, Calcutta 8: 1–342.
  49. Kurzweil, (H. 2010a) A precursory study of the Calanthe group (Orchidaceae) in Thailand. Adansonia 3, 32 (1): 57–107. https://doi.org/10.5252/a2010n1a4
  50. Kurzweil, H. (2010b) Taxonomic studies in the genus Peristylus (Orchidaceae) in Thailand. Nordic Journal of Botany 28 (1): 21–46. https://doi.org/10.1111/j.1756-1051.2009.00373.x
  51. Kurzweil, H. (2011) 28. Peristylus. In: Santisuk, T. & Larsen, K. (Eds.) Flora of Thailand 12 (1). Orchidaceae 1. The Forest Herbarium, National Park, Wildlife and Plant Conservation Department, Bangkok, pp. 222–250.
  52. Kurzweil, H. (2014) 44. Calanthe. In: Santisuk, T. & Baslev, H. (Eds.) Flora of Thailand 12 (2). Orchidaceae 2. The Forest Herbarium, National Park, Wildlife and Plant Conservation Department, Bangkok, pp. 341–376.
  53. Lanorsavanh, S., Souvannakhoummane, K, Kumar, P., Gale, S.W., Lamxay, V., Bounphanmy, S. & Chanthavongsa, K. (2019) Two new distribution records of Odontochilus Blume (Orchidaceae) from Laos. Thai Journal Botany 11 (2): 85–93.
  54. Lang, K.-Y. (1996) Three new species and two new combinations of Orchidaceae from China. Acta Phytotaxonomica Sinica 34 (6): 635–640.
  55. Lindley, J. (1821–1826) Collectanea botanica, or, Figures and botanical illustrations of rare and curious exotic plants. Richard & Arthur Taylor, Shoe-Lane, London, no pagination. https://doi.org/10.5962/bhl.title.6215
  56. Lindley, J. (1830–1840) The genera and species of orchidaceous plants. James Ridgway and Sons, London, 554 pp. https://doi.org/10.5962/bhl.title.499
  57. Lindley, J. (1857) Contributions to the orchidology of India, 1. Journal of the Proceedings of the Linnean Society. Botany 1 (4): 170–190. https://doi.org/10.1111/j.1095-8339.1857.tb02444.x
  58. Liu, Z., Chen, S., Chen, L. & Lei, S. (2009) The genus Paphiopedilum in China. Science Press, Beijing, 371 pp.
  59. Nguyen, T.T. (2001) The Orchids of Vietnam 1. Nha Xuat Ban Nong Nghiep, Ho Chi Minh, 424 pp.
  60. Odyuo, N., Roy, D., Bhattacharjee, A. & Ormerod, P. (2019) Rhomboda monensis (Orchidaceae), a new species of terrestrial orchid from India. Phytotaxa 405 (1): 61–64. https://doi.org/10.11646/phytotaxa.405.1.6
  61. Olędrzyńska, N. & Szlachetko, D.L. (2021) Contribution to the taxonomic revision of Brachycorythis-complex (Orchidaceae, Orchidoideae). Biodiversity Research and Conservation 62 (1): 5–117. https://doi.org/10.2478/biorc-2021-0004
  62. Olędrzyńska, N., Szlachetko, D.L. & Mieszkowska, A. (2016) Notes on the genera Silvorchis and Vietorchis (Orchidaceae-Orchidoideae). Phyton 56 (2): 193–200. http://dx.doi.org/10.12905/0380.phyton56(2)2016-0193
  63. Ormerod, P. (1995) A reinstatement of Rhomboda Lindl. (Orchidaceae subtribe Goodyerinae). Orchadian 11: 323–339.
  64. Ormerod, P. (1998) Miscellaneous Goodyerinae studies. Australian Orchid Review 63 (4): 10–11.
  65. Ormerod, P. (2002) Taxonomic changes in Goodyerinae (Orchidaceae: Orchidoideae). Lindleyana 17 (4): 189–238.
  66. Ormerod, P. & Bhattacharjee, A. (2014) The generic identity of Zeuxine pulchra King & Pantl. (Orchidaceae: Goodyerinae). Phytotaxa 191 (1): 177–180. https://doi.org/10.11646/phytotaxa.191.1.13
  67. Ormerod, P., Kurzweil, H. & Watthana, S. (2021) Annotated list of Orchidaceae for Myanmar. Phytotaxa 481 (1): 1–262. https://doi.org/10.11646/phytotaxa.481.1.1
  68. Pearce, N.R. & Cribb, P.J. (2002) The orchids of Bhutan. In: Flora of Bhutan. Vol. 3 (3). Royal Botanic Garden Edinburgh, UK & Royal Government of Bhutan, Huddersfield, 643 pp.
  69. Pfitzer, E. (1888) Untersuchungen über Bau und Entwicklung der Orchideenblüthe. I. Theil: Cypripedilinae, Ophrydinae, Neottiinae. Jahrbücher für Wissenschaftliche Botanik 19: 155–177.
  70. Poyarkov, N.A., Nguyen, T.V., Popov, E.S., Geissler, P., Pawangkhanant, P., Neang, T., Suwannapoom, C. & Orlov, N.L. (2021) Recent progress in taxonomic studies, biogeographic analysis and revised checklist of amphibians in Indochina. Russian Journal of Herpetology 28 (3A): 1–110. https://doi.org/10.30906/1026-2296-2021-28-3A-1-110
  71. Reichenbach, H.G. (1874) Enumeration of the orchids collected by the rev. E.C. Parish in the neighborhood of Moulmein, with descriptions of the new species. Transactions of the Linnean Society of London 30 (1): 133–155. https://doi.org/10.1111/j.1096-3642.1874.tb00004.x
  72. Reichenbach, H.G. (1878) IV. Orchideae Wilkesianae indescriptae. Otia Botanica Hamburgensia 1: 50–56.
  73. Rokaya, M.B., Raskoti, B.B., Timsina, B. & Münzbergova, Z. (2013) An annotated checklist of the orchids of Nepal. Nordic Journal of Botany 31: 511–550. https://doi.org/10.1111/j.1756-1051.2013.01230.x
  74. Schuiteman, A., Bonnet, P., Svengsuksa, B. & Barthélémy, D. (2008) An annotated checklist of the Orchidaceae of Laos. Nordic Journal of Botany 26 (5–6): 257–316. https://doi.org/10.1111/j.1756-1051.2008.00265.x
  75. Seidenfaden, G. (1969) Contributions to the orchid flora of Thailand. Botanisk Tidsskrift 65 (1–2): 100–162.
  76. Seidenfaden, G. (1977) Orchid genera in Thailand 5. Orchidoideae. Dansk Botanisk Arkiv 31 (2): 1–149.
  77. Seidenfaden, G. (1992) The orchids of Indochina. Opera Botanica 114: 385–395.
  78. Seidenfaden, G. & Wood, J.J. (1992) The orchids of Peninsular Malaysia and Singapore. Olsen & Olsen, Fredensborg, 779 pp.
  79. Smith, J.J. (1907) Die orchideen von Java. Bulletin du Deìpartement de l’Agriculture aux Indes Neìerlandaises 13: 1–78.
  80. Sun, M., Jin, Y., Huang, J., Kumar, P. & Tian, H.Z. (2021) Odontochilus putaoensis (Orchidaceae), a new record and key to the genus for China. Rheedea 31 (3): 200–205. https://doi.org/10.22244/rheedea.2021.31.03.11
  81. Szlachetko, D.L., Kras, M. & Mytnik, J. (2006) Matériaux pour la révision taxinomique du complexe Brachycorytis (Orchidaceae, Orchidoideae). Richardiana 6 (2): 72‒90.
  82. Tang, Е. & Wang, F.T. (1940) Contributions to the knowledge of eastern Asiatic Orchidaceae I. Bulletin of the Fan Memorial Institute of Biology, Botany 10: 23‒46.
  83. Tang, T. & Wang, F.T. (1951) Contributions to the knowledge of Eastern Asiatic Orchidaceae 2. Acta Phytotaxonomica Sinica 1 (1): 23–102.
  84. Thiers, B. (2023) [continuously updated] Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 16 June 2023).
  85. Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashütten. https://doi.org/10.12705/Code.2018
  86. Vietnam Administrative Atlas (2015) Vietnam Publishing House of Natural Resources, Environment and Cartography, Hanoi, 131 pp.
  87. Ya, J.-D., Zhang, T., Pandey, T.R., Liu, C., Han, Z.-D., Ye, D.-P., He, D.-M., Liu, Q., Yang, L., Huang, L., Zhang, R.-Z., Jiang, H. & Cai, J. (2021) New contributions to Goodyerinae and Dendrobiinae (Orchidaceae) in the flora of China. Plant Diversity 43 (5): 362–378. https://doi.org/10.1016/j.pld.2021.05.006
  88. Yokota, M. & Inoue, K. (2016) Calanthe R.Br. In: Iwatsuki, K., Boufford, D.E. & Ohba, H. (Eds.) Flora of Japan. IVb. Kodansha LTD, Tokyo, pp. 268–276.
  89. Yukawa, T. (2016) Taxonomic notes on the Orchidaceae of Japan and adjacent regions. Bulletin of the National Museum of Nature and Science B 42 (3): 103–111.
  90. Yukawa, T. & Cribb, P. (2014) Nomenclatural changes in the genus Calanthe (Orchidaceae). Bulletin of the National Museum of Nature and Science B, 40 (4): 145–152.
  91. Zhou, X., Cheng, Z., Liu, Q., Zhang, J., Hu, A., Huang, M., Hu, C. & Tian, H. (2016) An updated checklist of Orchidaceae for China, with two new national records. Phytotaxa 276 (1): 1–148. https://doi.org/10.11646/phytotaxa.276.1.1