Abstract
Four new species of Aspidistra (Asparagaceae), namely A. shiwandashanensis, A. ovatianthera, A. yunbiaoi and A. xiaoyunii, all local China endemics, are described and illustrated. The first three species are from Guangxi Zhuang Autonomous Region, and the last one is from Guangdong Province. The morphology, ecology, phenology, distribution, pollen morphology and the photo are provided for all the mentioned taxa.
References
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Averyanov, L.V., Le, T.A., Nguyen, K.S., Tillich, H.J., Nguyen, D.D., Tuan, A.H.L., Tran, H.D., Dat, P.T.T. & Maisak, T.V. (2019) Aspidistra erosa, A. sarcantha, and A. verruculosa (Asparagaceae), three new species from Vietnam. Phytotaxa 404: 102–110. https://doi.org/10.11646/phytotaxa.404.3.2</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Averyanov, L.V., Nguyen, K.S., Son, H.T., Tillich, H.J., Wynn-Jones, B. & Maisak, T.V. (2020) New taxa and new records in Aspidistra (Convallariaceae s.s.) of Laos and Vietnam. Nordic Journal of Botany 38: e02877. https://doi.org/10.1111/njb.02877</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Baillon, H. (1894) Etude d` un nouvel Aspidistra. Bulletin Mensuel de la Société Linnéenne de Paris 2: 1129–1132.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Cai, L., Peng, S., Tian, J., Dao, Z.L., Wei, N., Hu, G.W. & Wang, Q.F. (2018) Aspidistra austroyunnanensis (Asparagaceae), a new species from southern Yunnan, China. Phytotaxa 356: 233–237. https://doi.org/10.11646/phytotaxa.356.3.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ding, H.B., Xiong, K.C., Yang,B., Yin,J.T., Bouamanivong, S. & Tan, Y.H. (2021) New species and taxonomic notes of Aspidistra for the flora of China and Laos. Taiwania 66 (4): 439–449. https://doi.org/10.6165/tai.2021.66.439</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Erdtman, G. (1978) Handbook of Palynology. Science press. Beijing, 429 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Huang, J.L. & Hong, D.Y. (1996) Pollen morphology in the subtribe of Aspidistrinae (Liliaceae, s.l.). Journal of Systematis and Evolution 35 (2): 117–124.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Huang, X.Y., Peng, Y.D., Nong, D.X., Huang, B.Y. & Yu, L.Y. (2022) Aspidistra saccata (Asparagaceae), a new species with erect stem from limestone areas in Guangxi, China. Phytotaxa 541 (3): 297–300. https://doi.org/10.11646/phytotaxa.541.3.10</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hsu, T.C., Luu, H.T. & Li, C.W. (2022) Aspidistra heterotepala (Asparagaceae), a new species from southern Vietnam. Taiwania 67 (1): 21?24. https://doi.org/ 10.6165/tai.2022.67.21</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ker Gawler, J.B. (1822) Aspidistra lurida. The Botanical Register 8: 628. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Li, G.Z. (2004) The genus Aspidistra. Guangxi Science & Technology Publishing House, Nanning, 229 pp. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Li, G.Z. & Lin, C.R. (2016) Aspidistra. In: Li, S.G. (Ed.) Flora of Guangxi 5. Guangxi Science & Technology Publising House, Nanning, pp. 164–206.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Liang, S. & Tamura, M.N. (2000) Aspidistra. In: Wu, Z.Y. & Raven, P.H. (Eds.) Flora of China 24. Science Press, Beijing, and Missouri Botanical Garden Press, St. Louis, pp. 240–250.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lin, C.R., Liu, Y. (2011) Aspidistra punctatoides sp. nov. (Ruscaceae) from limestone areas in Guangxi, China. Nordic Journal of Botany 29: 189–193. https://doi: 10.1111/j.1756-1051.2011.00858.x</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lin, C.R., Wang, B.M., Liu, J. & Liu, Y. (2019) Aspidistra synpetala and A. pulchella, two new species of Aspidistra (Asparagaceae) from Guangxi, China. Taiwania 64: 80–85. https://doi.org/10.6165/tai.2019.64.80</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lindley, J. (1826) Aspidistra punctata Lindley. Botanical Register 12: t. 977.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lu, C.T., Chuang, K.C., Tseng, Y.H. Wang, C.C. & Wang, J.C. (2020) Taxonomic revision of Aspidistra Ker-Gawl. (Asparagaceae) in Taiwan. Taiwania 65 (3): 277–285. https://doi.org/10.6165/tai.2020.65.277</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lý, N.S., Nguyen, K.S., Hoàng, T.S., ??, V.N. & Tillich, H.J. (2021) Aspidistra magnifica (Asparagaceae), a new species from Central Vietnam. Phytotaxa 480: 22–28. https://doi.org/10.11646/phytotaxa.480.1.2</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Mo, F.Y., Zhang, S.Z., Lin, C.R. & Liu, Y. (2021) Aspidistra latistyla (Asparagaceae), a new species from China. Phytotaxa 512 (1): 73–78. https://doi.org/10.11646/phytotaxa.512.1.6</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nguyen, K.S., Averyanov, L.V., Tillich, H.J., Pham, V.T., Maisak, T.V. & Ly, N.S. (2017) Aspidistra cyathiflora var. bifolia and A. neglecta spp. nov. (Convallariaceae) from northern Vietnam. Nordic Journal of Botany 35: 482–487. https://doi.org/10.1111/njb.01504 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nguyen, K.S., Averyanov, L.V., Tillich, H.J., Le, T.M.L. & Nguyen, T.H. (2020) Aspidistra babensis (Asparagaceae), a new species from northern Vietnam. Annales Botanici Fennici 57: 17–22. https://doi.org/10.5735/085.057.0103</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nong, D.X., Huang, X.Y., Huang, B.Y. & Lu, H.Z. (2018) Aspidistra cleistantha, a new species of Asparagaceae from Guangxi, China. Phytotaxa 374: 178–180. https://doi.org/10.11646/phytotaxa.374.2.12</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tillich, H.J. (2005) A key for Aspidistra (Ruscaceae), including fifteen new species from Vietnam. Feddes Repertorium 116: 313–338. https://doi.org/10.1002/fedr.200511076</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tillich, H.J. (2014) The genus Aspidistra Ker-Gawl. (Asparagaceae) in Vietnam. Taiwania 59: 1–8. https://doi.org/10.6165/tai.2014.59.1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tillich, H.J. (2023) 200 years Aspidistra (Asparagaceae), and now more than 200 species: a new comprehensive determination key, and an annotated bibliography of the genus. Nordic Journal of Botany 2023: e03818. https://doi: 10.1111/njb.03818</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tillich, H.J. & Averyanov, L.V. (2012) Four new species of Aspidistra Ker Gawl. (Asparagaceae) from China and Vietnam with a comment on A. longifolia Hook.f. and A. hainanensis W.Y.Chun & F.C.How. Gardens’ Bulletin Singapore 64 (1): 201–209.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Trinh, N.B., Pham, V.T., Nguyen, T.S., Do, T.T.H., Nguyen, K.S., Anh, L.T., Nuraliev, M.S. & Vislobokov, N.A. (2021) Aspidistra sonlaensis (Asparagaceae, Nolinoideae), a new species from northern Vietnam. Phytotaxa 528 (1): 010–018. https://doi.org/10.11646/phytotaxa.528.1.2</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Vislobokov, N.A., Nuraliev, M.S., Kuznetsov, A.N. & Kuznetsova, S.P. (2016) Aspidistra globosa (Asparagaceae, Nolinoideae), a new species with erect stem from southern Vietnam. Phytotaxa 282: 46–52. https://doi.org/10.11646/phytotaxa.282.1.5 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Vislobokov, N.A., Kuznetsov, A.N., Kuznetsova, S.P. & Kuzmicheva, E.A. (2019) Aspidistra corniculata (Asparagaceae, Nolinoideae), a new species from Vietnam. Phytotaxa 397: 125–128. https://doi.org/10.11646/phytotaxa.397.1.15</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, Z., Peng, S.A., Peng, Y., Hu, G.W. & Wang, Q.F. (2021) Aspidistra longhuiensis (Asparagaceae), a new speices from Hunan, China. Phytotaxa 510 (1): 89–93. https://doi.org/10.11646/phytotaxa.510.1.10</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wu, L., Huang, Y.F., Nong, D.X. & Gao, Q. (2012) A new variety of Aspidistra from Guangxi, China. Guihaia 32 (2): 321–323. https://doi.org/10.3969/j.issn.1000-3142.2012.05.008</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhou, H., Yi, S.R., Gao, Q., Huang, J. & Wei, Y.J. (2016) Aspidistra revoluta, a new species from limestone areas in Chongqing, China. Phytotaxa 257 (3): 280–286. https://dx.doi.org/10.11646/phytotaxa.257.3.6</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Averyanov, L.V., Nguyen, K.S., Son, H.T., Tillich, H.J., Wynn-Jones, B. & Maisak, T.V. (2020) New taxa and new records in Aspidistra (Convallariaceae s.s.) of Laos and Vietnam. Nordic Journal of Botany 38: e02877. https://doi.org/10.1111/njb.02877</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Baillon, H. (1894) Etude d` un nouvel Aspidistra. Bulletin Mensuel de la Société Linnéenne de Paris 2: 1129–1132.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Cai, L., Peng, S., Tian, J., Dao, Z.L., Wei, N., Hu, G.W. & Wang, Q.F. (2018) Aspidistra austroyunnanensis (Asparagaceae), a new species from southern Yunnan, China. Phytotaxa 356: 233–237. https://doi.org/10.11646/phytotaxa.356.3.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ding, H.B., Xiong, K.C., Yang,B., Yin,J.T., Bouamanivong, S. & Tan, Y.H. (2021) New species and taxonomic notes of Aspidistra for the flora of China and Laos. Taiwania 66 (4): 439–449. https://doi.org/10.6165/tai.2021.66.439</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Erdtman, G. (1978) Handbook of Palynology. Science press. Beijing, 429 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Huang, J.L. & Hong, D.Y. (1996) Pollen morphology in the subtribe of Aspidistrinae (Liliaceae, s.l.). Journal of Systematis and Evolution 35 (2): 117–124.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Huang, X.Y., Peng, Y.D., Nong, D.X., Huang, B.Y. & Yu, L.Y. (2022) Aspidistra saccata (Asparagaceae), a new species with erect stem from limestone areas in Guangxi, China. Phytotaxa 541 (3): 297–300. https://doi.org/10.11646/phytotaxa.541.3.10</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hsu, T.C., Luu, H.T. & Li, C.W. (2022) Aspidistra heterotepala (Asparagaceae), a new species from southern Vietnam. Taiwania 67 (1): 21?24. https://doi.org/ 10.6165/tai.2022.67.21</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ker Gawler, J.B. (1822) Aspidistra lurida. The Botanical Register 8: 628. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Li, G.Z. (2004) The genus Aspidistra. Guangxi Science & Technology Publishing House, Nanning, 229 pp. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Li, G.Z. & Lin, C.R. (2016) Aspidistra. In: Li, S.G. (Ed.) Flora of Guangxi 5. Guangxi Science & Technology Publising House, Nanning, pp. 164–206.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Liang, S. & Tamura, M.N. (2000) Aspidistra. In: Wu, Z.Y. & Raven, P.H. (Eds.) Flora of China 24. Science Press, Beijing, and Missouri Botanical Garden Press, St. Louis, pp. 240–250.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lin, C.R., Liu, Y. (2011) Aspidistra punctatoides sp. nov. (Ruscaceae) from limestone areas in Guangxi, China. Nordic Journal of Botany 29: 189–193. https://doi: 10.1111/j.1756-1051.2011.00858.x</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lin, C.R., Wang, B.M., Liu, J. & Liu, Y. (2019) Aspidistra synpetala and A. pulchella, two new species of Aspidistra (Asparagaceae) from Guangxi, China. Taiwania 64: 80–85. https://doi.org/10.6165/tai.2019.64.80</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lindley, J. (1826) Aspidistra punctata Lindley. Botanical Register 12: t. 977.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lu, C.T., Chuang, K.C., Tseng, Y.H. Wang, C.C. & Wang, J.C. (2020) Taxonomic revision of Aspidistra Ker-Gawl. (Asparagaceae) in Taiwan. Taiwania 65 (3): 277–285. https://doi.org/10.6165/tai.2020.65.277</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lý, N.S., Nguyen, K.S., Hoàng, T.S., ??, V.N. & Tillich, H.J. (2021) Aspidistra magnifica (Asparagaceae), a new species from Central Vietnam. Phytotaxa 480: 22–28. https://doi.org/10.11646/phytotaxa.480.1.2</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Mo, F.Y., Zhang, S.Z., Lin, C.R. & Liu, Y. (2021) Aspidistra latistyla (Asparagaceae), a new species from China. Phytotaxa 512 (1): 73–78. https://doi.org/10.11646/phytotaxa.512.1.6</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nguyen, K.S., Averyanov, L.V., Tillich, H.J., Pham, V.T., Maisak, T.V. & Ly, N.S. (2017) Aspidistra cyathiflora var. bifolia and A. neglecta spp. nov. (Convallariaceae) from northern Vietnam. Nordic Journal of Botany 35: 482–487. https://doi.org/10.1111/njb.01504 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nguyen, K.S., Averyanov, L.V., Tillich, H.J., Le, T.M.L. & Nguyen, T.H. (2020) Aspidistra babensis (Asparagaceae), a new species from northern Vietnam. Annales Botanici Fennici 57: 17–22. https://doi.org/10.5735/085.057.0103</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nong, D.X., Huang, X.Y., Huang, B.Y. & Lu, H.Z. (2018) Aspidistra cleistantha, a new species of Asparagaceae from Guangxi, China. Phytotaxa 374: 178–180. https://doi.org/10.11646/phytotaxa.374.2.12</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tillich, H.J. (2005) A key for Aspidistra (Ruscaceae), including fifteen new species from Vietnam. Feddes Repertorium 116: 313–338. https://doi.org/10.1002/fedr.200511076</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tillich, H.J. (2014) The genus Aspidistra Ker-Gawl. (Asparagaceae) in Vietnam. Taiwania 59: 1–8. https://doi.org/10.6165/tai.2014.59.1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tillich, H.J. (2023) 200 years Aspidistra (Asparagaceae), and now more than 200 species: a new comprehensive determination key, and an annotated bibliography of the genus. Nordic Journal of Botany 2023: e03818. https://doi: 10.1111/njb.03818</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tillich, H.J. & Averyanov, L.V. (2012) Four new species of Aspidistra Ker Gawl. (Asparagaceae) from China and Vietnam with a comment on A. longifolia Hook.f. and A. hainanensis W.Y.Chun & F.C.How. Gardens’ Bulletin Singapore 64 (1): 201–209.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Trinh, N.B., Pham, V.T., Nguyen, T.S., Do, T.T.H., Nguyen, K.S., Anh, L.T., Nuraliev, M.S. & Vislobokov, N.A. (2021) Aspidistra sonlaensis (Asparagaceae, Nolinoideae), a new species from northern Vietnam. Phytotaxa 528 (1): 010–018. https://doi.org/10.11646/phytotaxa.528.1.2</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Vislobokov, N.A., Nuraliev, M.S., Kuznetsov, A.N. & Kuznetsova, S.P. (2016) Aspidistra globosa (Asparagaceae, Nolinoideae), a new species with erect stem from southern Vietnam. Phytotaxa 282: 46–52. https://doi.org/10.11646/phytotaxa.282.1.5 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Vislobokov, N.A., Kuznetsov, A.N., Kuznetsova, S.P. & Kuzmicheva, E.A. (2019) Aspidistra corniculata (Asparagaceae, Nolinoideae), a new species from Vietnam. Phytotaxa 397: 125–128. https://doi.org/10.11646/phytotaxa.397.1.15</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, Z., Peng, S.A., Peng, Y., Hu, G.W. & Wang, Q.F. (2021) Aspidistra longhuiensis (Asparagaceae), a new speices from Hunan, China. Phytotaxa 510 (1): 89–93. https://doi.org/10.11646/phytotaxa.510.1.10</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wu, L., Huang, Y.F., Nong, D.X. & Gao, Q. (2012) A new variety of Aspidistra from Guangxi, China. Guihaia 32 (2): 321–323. https://doi.org/10.3969/j.issn.1000-3142.2012.05.008</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zhou, H., Yi, S.R., Gao, Q., Huang, J. & Wei, Y.J. (2016) Aspidistra revoluta, a new species from limestone areas in Chongqing, China. Phytotaxa 257 (3): 280–286. https://dx.doi.org/10.11646/phytotaxa.257.3.6</span></span></span>