Abstract
Revisions of Tillandsioideae have divided the genus Vriesea into nine genera since its last monograph, published in Flora Neotropica. Despite these efforts, the taxonomic placement of some species remains uncertain. We review the 21 species of Vriesea “incertae sedis”, to assist in a new circumscription of this genus and contribute to the nomenclatural stability of related taxa. We present 17 redescriptions, eight new combinations, two lectotypifications, one revalidation, and notes for the all taxa.
References
<p>André, E.F. (1889) <em>Bromeliaceae Andreanae. Description et histoire des Broméliacées récoltées dans la Colombie, l’Ecuador et le Venezuela</em>. Librairie Agricole, Paris, 118 pp., 40 pl. https://doi.org/10.5962/bhl.title.78580</p>
<p>Baker, J.G. (1888) A synopsis of Tillandsieae. <em>Journal of Botany, Britsh and Foreign</em> 26: 39–50.</p>
<p>Baker, J.G. (1889) <em>Handbook of the Bromeliaceae</em>. G. Bell & Sons, London, 244 pp. https://doi.org/10.5962/bhl.title.51681</p>
<p>Barfuss, M.H.J., Samuel, R., Till, W. & Stuessy, T.F. (2005) Phylogenetic relationships in subfamily Tillandsioideae (Bromeliaceae) based on DNA sequence data from seven plastid regions. <em>American Journal of Botany</em> 92: 337–351. https://doi.org/10.3732/ajb.92.2.337</p>
<p>Barfuss, M.H.J., Till, W., Leme, E.M.C., Pinzón, J.P., Manzanares, J.M., Halbritter, H., Samuel, R. & Brown, G.K. (2016) Taxonomic revision of Bromeliaceae subfam. Tillandsioideae based on a multi-locus DNA sequence phylogeny and morphology. <em>Phytotaxa</em> 279 (1): 1–97. https://doi.org/10.11646/phytotaxa.279.1.1</p>
<p>Beer, J.G. (1856 ‘1857’) <em>Die Familie der Bromeliaceen. Nach ihrem habituellen Charakter bearbeitet mit besonderer Berücksichtigung der Ananassa</em>. Tendler & Co., Vienna, 272 pp. https://doi.org/10.5962/bhl.title.15461</p>
<p>Brown, G.K. & Gilmartin, A.J. (1984) Stigma structure and variation in Bromeliaceae—neglected taxonomic characters. <em>Brittonia</em> 36: 364–374. https://doi.org/10.2307/2806597</p>
<p>Brown, G.K. & Gilmartin, A.J. (1988) Comparative ontogeny of bromeliaceous stigmas. <em>In:</em> Leins, O., Tucker, S.C. & Endress, P.K. (Eds.) <em>Aspects of floral development</em>. Lubrecht & Cramer Ltd., Berlim-Stuttgart, pp. 191–204.</p>
<p>Brown, G.K. & Gilmartin, A.J. (1989) Stigma types in Bromeliaceae-A systematic survey. <em>Systematic Botany</em> 14: 110–132. https://doi.org/10.2307/2419055</p>
<p>Christenhusz, M.J.M., Fay, M.F. & Byng, J.W. (2018) <em>The Global Flora</em>. Vol. 4: Special Edition, GLOVAP Nomenclature Part 1. <em>Pl</em>ant Gateway Ltd., Bradford, United Kingdom.</p>
<p>Costa, A.F., Gomes-da-Silva J. & Wanderley, M.G.L. (2014) <em>Vriesea</em> (Bromeliaceae, Tillandsioideae): taxonomic history, and morphology of the Brazilian lineage. <em>Journal of the Torrey Botanical Society</em> 141: 338–352. https://doi.org/10.3159/TORREY-D-13-00070.1</p>
<p>Evans, T.M., Jabaily, R.S., de Faria, A.P.G., de Sousa, L.D.O.F., Wendt, T. & Brown, G.K. (2015) Phylogenetic relationships in Bromeliaceae subfamily Bromelioideae based on chloroplast DNA sequence data. <em>Systematic Botany</em> 40: 116–128. https://doi.org/10.1600/036364415X686413</p>
<p>Givnish, T.J., Barfuss, M.H.J., Ee, B.V., Riina, R., Schulte, K., Horres, R., Gonsiska, P.A., Jabaily, R.S., Crayn, D.M., Smith, A.C., Winter, K., Brown, G.K., Evans, T.M., Holst, B.K., Luther, H., Till, W., Zizka, G., Berry, P.E. & Sytsma, K.J. (2011) Phylogeny, adaptive radiation, and historical biogeography in Bromeliaceae: insights from an eight-locus plastid phylogeny. <em>American Journal of Botany </em>98: 872–895. https://doi.org/10.3732/ajb.1000059</p>
<p>Givnish, T.J. (2017) A New World of plants. <em>Science</em> 358: 1535–1536. https://doi.org/10.1126/science.aar4191</p>
<p>Givnish, T.J., Zuluaga, A., Spalink, D., Gomez, M.S., Lam, V.K.Y., Saarela, J.M., Sass, C., Iles, W.J.D., Lima de Sousa, D.J., Leebens-Mack, J., Pires, J.C., Zomlefer, W.B., Gandolfo, M.A., Davis, J.I., Stevenson, D.W., dePamphilis, C., Specht, C.D., Graham, S.W., Barrett, C.F. & Ane, C. (2018) Monocot plastid phylogenomics, timeline, net rates of species diversification, the power of multi-gene analyse and a functional model for the origin of monocots. <em>American Journal of Botany </em>105: 1888–1910. https://doi.org/10.1002/ajb2.1178</p>
<p>Gomes-da-Silva, J. & Costa, A.F. (2013) An updated overview of taxonomy and phylogenetic history of Tillandsioideae genera (Bromeliaceae: Poales). <em>Global Journal of Botanical Science </em>1: 1–8. https://doi.org/10.12974/2311-858X.2013.01.01.1</p>
<p>Gomes-da-Silva, J. & Souza-Chies, T.T. (2018) What actually is <em>Vriesea</em>? A total evidence approach in a polyphyletic genus of Tillandsioideae (Bromeliaceae, Poales). <em>Cladistics</em> 34: 181–199. https://doi.org/10.1111/cla.12200</p>
<p>Gouda, E.J. (1987) <em>Flora of the Guianas: Phanerogams. 189. Bromeliaceae subfamily Tillandsioideae</em> (No. 3). Koeltz Scientific Books.</p>
<p>Gouda, E.J., Butcher, D. & Gouda, C.S. (Continuously updated) <em>Encyclopaedia of Bromeliads, Version 3</em>. Available from<em>:</em> http://encyclopedia.florapix.nl/ (accessed 20 June 2019)</p>
<p>Gouda, E.J. (2017) A new species of <em>Cipuropsis</em>, and some remarks about this recently resurrected genus. <em>Journal of the Bromeliad Society</em> 66: 75–86.</p>
<p>Grant, J.R. (1992) <em>Vriesea lutheriana</em>, a new species with tripinnate inflorescence from Costa Rica. <em>Journal of the Bromeliad Society</em> 42: 114–117.</p>
<p>Grant, J.R. (1995a) Bromelienstudien. The resurrection of <em>Alcantarea</em> and <em>Werauhia</em>, a new genus. <em>Tropische und Subtropische Pflanzenwelt</em> 91: 1–57.</p>
<p>Grant, J.R. (1995b) New combinations and new taxa in the Bromeliaceae. <em>Phytologia</em> 79: 254–256.</p>
<p>Grisebach, A.H.R. (1864) Uber die von Fendler in Venezuela gesammelten Bromeliaceen. <em>Nachrichten von der Königl. Gesellschaft der Wissenschaften und der Georg Augusts Universität zu Göttingen Nr</em>. 1: 1–21.</p>
<p>Harms, H. (1929) Bromeliaceae novae III. <em>Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem</em> 10: 784–805. https://doi.org/10.2307/3994800</p>
<p>Harms, H. (1930) Bromeliaceae. <em>In</em>: Engler, A. & Prantl, K. (Eds.) <em>Die natürlichen Pflanzenfamilien nebst ihren Gattungen und wichtigsten Arten, insbesondere den Nutzpflanzen,</em> ed. 2. Verlag von Wilhem Engelmen, Leipzig, pp. 65–169.</p>
<p>Harms, H. (1935) Bromeliaceae Novae V. <em>Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem</em> 12: 529–539. https://doi.org/10.2307/3994848</p>
<p>Jussieu, A.L. (1789) <em>Genera Plantarum secundum ordines naturales disposita, juxta methodum in horto region parisiensi exaratam</em>. Herissant, Paris, 498 pp. https://doi.org/10.5962/bhl.title.284</p>
<p>Kessous I.M., Neves B., Couto D.R., Paixão-Souza B., Pederneiras L.C., Moura R.L., Barfuss M.H.J., Salgueiro F. & Costa A.F. (2020) Historical biogeography of a Brazilian lineage of Tillandsioideae (subtribe Vrieseinae, Bromeliaceae): the Paranaean Sea hypothesized as the main vicariant event. <em>Botanical Journal of the Linnean Society </em>192: 625–641. https://doi.org/10.1093/botlinnean/boz038</p>
<p>Lindley, J. (1843) <em>Vriesia psittacina</em>, Parrot flowered <em>Vriesia</em>. <em>Edwards’s Botanical Register</em> 29: 10–11.</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum</em>. Laurentius Salvius, Stockholm, 1200 pp.</p>
<p>Luther, H.E. & Norton, K.F. (2009) A new <em>Vriesea</em> species from Northern Peru. <em>Journal of the Bromeliad Society</em> 59: 204–206.</p>
<p>Luther, H.E. & Norton, K.F. (2010) A new grey <em>Vriesea </em>from Northern Peru. <em>Journal of the Bromeliad Society</em> 60: 53–56.</p>
<p>Luther, H.E. & Norton, K.F. (2011) <em>Vriesea skotakii</em>, a new ornamental bromeliad from Panama. <em>Journal of the Bromeliad Society</em> 61: 53–56.</p>
<p>Machado, T.M., Loiseau, O., Paris, M., Weigand, A., Versieux, L.M., Stehmann, J.R., Lexer, C. & Salamin, N. (2020) Systematics of <em>Vriesea</em> (Bromeliaceae): phylogenetic relationships based on nuclear gene and partial plastome sequences. <em>Botanical Journal of the Linnean Society</em> 192: 656–674. https://doi.org/10.1093/botlinnean/boz102</p>
<p>Mez, C.C. (1894–1895) Bromeliaceae. <em>In:</em> von Martius, C.F.P (Ed.) <em>Flora brasiliensis</em>, vol. 3 (3). R. Oldenbourg, Munich, Vien & Leipzig, pp. 511–674. https://doi.org/10.5962/bhl.title.454</p>
<p>Mez, C.C. (1896) Bromeliaceae. <em>In</em>: de Candolle, C. (Ed.) <em>Monographiae phanerogamarum.</em> Vol. 9. Masson & Cie, Paris, pp. i–lxxvii,1–990. Available from: http://www.biodiversitylibrary.org/item/145130 (accessed 12 August 2019)</p>
<p>Mez, C.C. (1919) XVII - Additamenta Monographica. <em>Repertorium Specierum Novarum Regni Vegetabilis</em> 16: 65–79. https://doi.org/10.1002/fedr.19190160502</p>
<p>Mez, C.C. (1935) Bromeliaceae. <em>In</em>: Engler, A. (Ed.) <em>Das Pflanzenreich</em>. Vol. 4. W. Engelmann, Leipzig, pp. 362–667. Available from: http://www.biodiversitylibrary.org/item/56490 (accessed 12 August 2019)</p>
<p>Morren, É. (1875) Notice sur le <em>Vriesea platzmanni</em> sp. nov. ou <em>Vriesea</em> de M. Julius Platzmann. <em>La Belgique Horticole</em> 25: 349–351.</p>
<p>Morren, É. (1881) Description du <em>Vriesea chrysostachis</em>, sp. nov. de MM. J. Veitch, A. Londres. <em>La Belgique horticole </em>31: 87.</p>
<p>Niesler, I.M. (2016) Scanned Bromeliads – a Flash of Colours!. <em>Die Bromelie</em> 2016: 4–14.</p>
<p>Radford, A.E. (1986) <em>Fundamentals of Plant Systematics</em>. New York: Harper & Row, Publ. Inc., 498 pp.</p>
<p>Ramírez N.I. & Bevilacqua, M. (1990) Una nueva especie de <em>Tillandsia </em>(Bromeliaceae) Para la ciencia. <em>Acta Botánica Venezuélica</em> 15: 149–152.</p>
<p>Rauh, W. (1976) Neue <em>Vriesea</em>-Arten aus Peru und Ecuador. <em>Tropische und Subtropische Pflanzenwelt </em>16: 19–22.</p>
<p>Rauh, W. (1976) Bromelienstudien (Mitteilung 5). I. Neue und wenig bekannte Arten aus Peru und anderen Ländern. <em>Tropische und subtropische Pflanzenwelt</em> 16: 207–259.</p>
<p>Reed, C.F. & Read, R.W. (1981) Two New Species of Bromeliads from Venezuela. <em>Journal of the Bromeliad Society</em> 31: 59–61.</p>
<p>Ruiz, H.L. & Pavon, J.V. (1802) <em>Flora peruviana, et chilensis, sive descriptiones, et icones plantarum peruvianarum, et chilensium, secundum systema linnaeanum digestae, cum characteribus plurium generum evulgatorum reformatis.</em> Tomus 3. Madrid, Typis Gabrielis de Sancha, 95 pp.</p>
<p>Scharf, U. & Gouda, E.J. (2008) Bringing Bromeliaceae back to homeland botany. <em>Journal of the Bromeliad Society</em> 58: 123–129.</p>
<p>Smith, L.B. (1931) Bromeliaceae in H. A. Gleason. The Tyler-Duida-Expedition. <em>Bulletin of the Torrey Botanical Club</em> 58: 337–341. https://doi.org/10.2307/2480694</p>
<p>Smith, L.B. (1935) Studies in the Bromeliaceae - VI. <em>Contributions of the Gray Herbarium</em> 106: 147–220. https://doi.org/10.5962/p.336160</p>
<p>Smith, L.B. (1946) Bromeliaceae <em>In</em> Schultes, R. W. Plantae Austro-Americanae-III. <em>Botanical Museum Leaflets </em>12: 118–122. https://doi.org/10.5962/p.168447</p>
<p>Smith, L.B. (1948) Bromeliáceas notables de Colombia - IV. <em>Caldasia</em> 5: 1–15.</p>
<p>Smith, L.B. (1953) Notes on Bromeliaceae - I. <em>Phytologia</em> 4: 213–221.</p>
<p>Smith, L.B. (1955) Notes on Bromeliaceae -VI. <em>Phytologia</em> 5: 281–288. https://doi.org/10.5962/bhl.part.12982</p>
<p>Smith, L.B. (1955) Bromeliaceae. Plantae Colombianae XIII. <em>Botanical Museum Leaflets</em> 17: 67–77.</p>
<p>Smith, L.B. (1957) Bromeliaceae. The Botany of the Guiana Highland – Part II. <em>Memoirs of the New York Botanical Garden </em>9: 283–318.</p>
<p>Smith, L.B. (1959) Notes on Bromeliaceae - XI. <em>Phytologia </em>6 (8): 433–443<em>. </em>https://doi.org/10.5962/bhl.part.9778</p>
<p>Smith, L.B. (1963) <em>Vriesea soderstromii</em>. <em>Bulletin of the Bromeliad Society </em>13: 52–53. https://doi.org/10.1093/aibsbulletin/13.6.52</p>
<p>Smith, L.B. (1967) Notes on Bromeliaceae -XXVI. <em>Phytologia</em> 15: 163–200. https://doi.org/10.5962/bhl.part.21069</p>
<p>Smith, L.B. (1968) Notes on Bromeliaceae - XXVIII. <em>Phytologia</em> 16: 459–463. https://doi.org/10.5962/bhl.part.21075</p>
<p>Smith, L.B. (1969) Bromeliaceae. <em>In: Maguire </em>B. Botany of the Guayana Highland –Part VIII. <em>Memoirs of the New York Botanical Garden </em>18: 29–32.</p>
<p>Smith, L.B. (1970) Notes on Bromeliaceae - XXXI. <em>Phytologia</em> 20: 121–183. https://doi.org/10.5962/bhl.part.7116</p>
<p>Smith, L.B. & Read R.W. (1977) Notes on Bromeliaceae -XXXIX. <em>Phytologia</em> 38: 135–148.</p>
<p>Smith, L.B. & Downs, R.J. (1974) Pitcairnioideae (Bromeliaceae).<em> Flora Neotropica</em>. Monograph 14: 1–662.</p>
<p>Smith, L.B. & Downs, R.J. (1977) Bromeliaceae (Tillandsioideae). <em>Flora Neotropica Monograph </em>14: 663–1492.</p>
<p>Spencer, M.A. & Smith, L.B. (1993) <em>Racinaea</em>, a new genus of Bromeliaceae (Tillandsioideae). <em>Phytologia</em> 74: 151–160.</p>
<p>Stearn, W.T. (1973) <em>Botanical Latin,</em> ed. 2. London: David & Charles Newton Abbot, 566 pp.</p>
<p>Till, W. (2008) <em>Vriesea speckmaieri</em> – a New Lower-altitude Cloud Forest Species from Northern Venezuela. <em>Journal of the Bromeliad Society</em> 58: 250–254.</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (Eds.) (2018) <em>International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. <em>Regnum Vegetabile</em> 159. Glasshütten: Koeltz Botanical Books. https://doi.org/10.12705/Code.2018</p>
<p>Ule, E. (1907 ‘1906’) II. Beiträge zur Flora der Hylaea nach den Sammlungen von Ule’s Amazonas-Expedition. <em>Verhandlungen des Botanischen Vereins für die Provinz Brandenburg </em>48: 117–208.</p>
<p>Versieux, L.M. & Wanderley, M.D.G.L. (2010) Delimitation of the <em>Alcantarea extensa complex</em> (Bromeliaceae) and a new species from Espírito Santo, Brazil. <em>Rodriguésia</em> 61: 421–429. https://doi.org/10.1590/2175-7860201061306</p>
<p>Wittmack, L. (1890) Plantae Lehmannianae in Guatemala, Costarica, Columbia, Ecuador etc. collectae. Bromeliaceae. <em>Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie </em>11: 52–71.</p>
<p>Wright, C.H. (1871) Flora Cubana. <em>Anales de la Academia de Ciencias Médicas, Físicas y Naturales de la Habana</em> 8: 48–56. [https://www.biodiversitylibrary.org/page/14603136#page/65/mode/1up]</p>
<p>Baker, J.G. (1888) A synopsis of Tillandsieae. <em>Journal of Botany, Britsh and Foreign</em> 26: 39–50.</p>
<p>Baker, J.G. (1889) <em>Handbook of the Bromeliaceae</em>. G. Bell & Sons, London, 244 pp. https://doi.org/10.5962/bhl.title.51681</p>
<p>Barfuss, M.H.J., Samuel, R., Till, W. & Stuessy, T.F. (2005) Phylogenetic relationships in subfamily Tillandsioideae (Bromeliaceae) based on DNA sequence data from seven plastid regions. <em>American Journal of Botany</em> 92: 337–351. https://doi.org/10.3732/ajb.92.2.337</p>
<p>Barfuss, M.H.J., Till, W., Leme, E.M.C., Pinzón, J.P., Manzanares, J.M., Halbritter, H., Samuel, R. & Brown, G.K. (2016) Taxonomic revision of Bromeliaceae subfam. Tillandsioideae based on a multi-locus DNA sequence phylogeny and morphology. <em>Phytotaxa</em> 279 (1): 1–97. https://doi.org/10.11646/phytotaxa.279.1.1</p>
<p>Beer, J.G. (1856 ‘1857’) <em>Die Familie der Bromeliaceen. Nach ihrem habituellen Charakter bearbeitet mit besonderer Berücksichtigung der Ananassa</em>. Tendler & Co., Vienna, 272 pp. https://doi.org/10.5962/bhl.title.15461</p>
<p>Brown, G.K. & Gilmartin, A.J. (1984) Stigma structure and variation in Bromeliaceae—neglected taxonomic characters. <em>Brittonia</em> 36: 364–374. https://doi.org/10.2307/2806597</p>
<p>Brown, G.K. & Gilmartin, A.J. (1988) Comparative ontogeny of bromeliaceous stigmas. <em>In:</em> Leins, O., Tucker, S.C. & Endress, P.K. (Eds.) <em>Aspects of floral development</em>. Lubrecht & Cramer Ltd., Berlim-Stuttgart, pp. 191–204.</p>
<p>Brown, G.K. & Gilmartin, A.J. (1989) Stigma types in Bromeliaceae-A systematic survey. <em>Systematic Botany</em> 14: 110–132. https://doi.org/10.2307/2419055</p>
<p>Christenhusz, M.J.M., Fay, M.F. & Byng, J.W. (2018) <em>The Global Flora</em>. Vol. 4: Special Edition, GLOVAP Nomenclature Part 1. <em>Pl</em>ant Gateway Ltd., Bradford, United Kingdom.</p>
<p>Costa, A.F., Gomes-da-Silva J. & Wanderley, M.G.L. (2014) <em>Vriesea</em> (Bromeliaceae, Tillandsioideae): taxonomic history, and morphology of the Brazilian lineage. <em>Journal of the Torrey Botanical Society</em> 141: 338–352. https://doi.org/10.3159/TORREY-D-13-00070.1</p>
<p>Evans, T.M., Jabaily, R.S., de Faria, A.P.G., de Sousa, L.D.O.F., Wendt, T. & Brown, G.K. (2015) Phylogenetic relationships in Bromeliaceae subfamily Bromelioideae based on chloroplast DNA sequence data. <em>Systematic Botany</em> 40: 116–128. https://doi.org/10.1600/036364415X686413</p>
<p>Givnish, T.J., Barfuss, M.H.J., Ee, B.V., Riina, R., Schulte, K., Horres, R., Gonsiska, P.A., Jabaily, R.S., Crayn, D.M., Smith, A.C., Winter, K., Brown, G.K., Evans, T.M., Holst, B.K., Luther, H., Till, W., Zizka, G., Berry, P.E. & Sytsma, K.J. (2011) Phylogeny, adaptive radiation, and historical biogeography in Bromeliaceae: insights from an eight-locus plastid phylogeny. <em>American Journal of Botany </em>98: 872–895. https://doi.org/10.3732/ajb.1000059</p>
<p>Givnish, T.J. (2017) A New World of plants. <em>Science</em> 358: 1535–1536. https://doi.org/10.1126/science.aar4191</p>
<p>Givnish, T.J., Zuluaga, A., Spalink, D., Gomez, M.S., Lam, V.K.Y., Saarela, J.M., Sass, C., Iles, W.J.D., Lima de Sousa, D.J., Leebens-Mack, J., Pires, J.C., Zomlefer, W.B., Gandolfo, M.A., Davis, J.I., Stevenson, D.W., dePamphilis, C., Specht, C.D., Graham, S.W., Barrett, C.F. & Ane, C. (2018) Monocot plastid phylogenomics, timeline, net rates of species diversification, the power of multi-gene analyse and a functional model for the origin of monocots. <em>American Journal of Botany </em>105: 1888–1910. https://doi.org/10.1002/ajb2.1178</p>
<p>Gomes-da-Silva, J. & Costa, A.F. (2013) An updated overview of taxonomy and phylogenetic history of Tillandsioideae genera (Bromeliaceae: Poales). <em>Global Journal of Botanical Science </em>1: 1–8. https://doi.org/10.12974/2311-858X.2013.01.01.1</p>
<p>Gomes-da-Silva, J. & Souza-Chies, T.T. (2018) What actually is <em>Vriesea</em>? A total evidence approach in a polyphyletic genus of Tillandsioideae (Bromeliaceae, Poales). <em>Cladistics</em> 34: 181–199. https://doi.org/10.1111/cla.12200</p>
<p>Gouda, E.J. (1987) <em>Flora of the Guianas: Phanerogams. 189. Bromeliaceae subfamily Tillandsioideae</em> (No. 3). Koeltz Scientific Books.</p>
<p>Gouda, E.J., Butcher, D. & Gouda, C.S. (Continuously updated) <em>Encyclopaedia of Bromeliads, Version 3</em>. Available from<em>:</em> http://encyclopedia.florapix.nl/ (accessed 20 June 2019)</p>
<p>Gouda, E.J. (2017) A new species of <em>Cipuropsis</em>, and some remarks about this recently resurrected genus. <em>Journal of the Bromeliad Society</em> 66: 75–86.</p>
<p>Grant, J.R. (1992) <em>Vriesea lutheriana</em>, a new species with tripinnate inflorescence from Costa Rica. <em>Journal of the Bromeliad Society</em> 42: 114–117.</p>
<p>Grant, J.R. (1995a) Bromelienstudien. The resurrection of <em>Alcantarea</em> and <em>Werauhia</em>, a new genus. <em>Tropische und Subtropische Pflanzenwelt</em> 91: 1–57.</p>
<p>Grant, J.R. (1995b) New combinations and new taxa in the Bromeliaceae. <em>Phytologia</em> 79: 254–256.</p>
<p>Grisebach, A.H.R. (1864) Uber die von Fendler in Venezuela gesammelten Bromeliaceen. <em>Nachrichten von der Königl. Gesellschaft der Wissenschaften und der Georg Augusts Universität zu Göttingen Nr</em>. 1: 1–21.</p>
<p>Harms, H. (1929) Bromeliaceae novae III. <em>Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem</em> 10: 784–805. https://doi.org/10.2307/3994800</p>
<p>Harms, H. (1930) Bromeliaceae. <em>In</em>: Engler, A. & Prantl, K. (Eds.) <em>Die natürlichen Pflanzenfamilien nebst ihren Gattungen und wichtigsten Arten, insbesondere den Nutzpflanzen,</em> ed. 2. Verlag von Wilhem Engelmen, Leipzig, pp. 65–169.</p>
<p>Harms, H. (1935) Bromeliaceae Novae V. <em>Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem</em> 12: 529–539. https://doi.org/10.2307/3994848</p>
<p>Jussieu, A.L. (1789) <em>Genera Plantarum secundum ordines naturales disposita, juxta methodum in horto region parisiensi exaratam</em>. Herissant, Paris, 498 pp. https://doi.org/10.5962/bhl.title.284</p>
<p>Kessous I.M., Neves B., Couto D.R., Paixão-Souza B., Pederneiras L.C., Moura R.L., Barfuss M.H.J., Salgueiro F. & Costa A.F. (2020) Historical biogeography of a Brazilian lineage of Tillandsioideae (subtribe Vrieseinae, Bromeliaceae): the Paranaean Sea hypothesized as the main vicariant event. <em>Botanical Journal of the Linnean Society </em>192: 625–641. https://doi.org/10.1093/botlinnean/boz038</p>
<p>Lindley, J. (1843) <em>Vriesia psittacina</em>, Parrot flowered <em>Vriesia</em>. <em>Edwards’s Botanical Register</em> 29: 10–11.</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum</em>. Laurentius Salvius, Stockholm, 1200 pp.</p>
<p>Luther, H.E. & Norton, K.F. (2009) A new <em>Vriesea</em> species from Northern Peru. <em>Journal of the Bromeliad Society</em> 59: 204–206.</p>
<p>Luther, H.E. & Norton, K.F. (2010) A new grey <em>Vriesea </em>from Northern Peru. <em>Journal of the Bromeliad Society</em> 60: 53–56.</p>
<p>Luther, H.E. & Norton, K.F. (2011) <em>Vriesea skotakii</em>, a new ornamental bromeliad from Panama. <em>Journal of the Bromeliad Society</em> 61: 53–56.</p>
<p>Machado, T.M., Loiseau, O., Paris, M., Weigand, A., Versieux, L.M., Stehmann, J.R., Lexer, C. & Salamin, N. (2020) Systematics of <em>Vriesea</em> (Bromeliaceae): phylogenetic relationships based on nuclear gene and partial plastome sequences. <em>Botanical Journal of the Linnean Society</em> 192: 656–674. https://doi.org/10.1093/botlinnean/boz102</p>
<p>Mez, C.C. (1894–1895) Bromeliaceae. <em>In:</em> von Martius, C.F.P (Ed.) <em>Flora brasiliensis</em>, vol. 3 (3). R. Oldenbourg, Munich, Vien & Leipzig, pp. 511–674. https://doi.org/10.5962/bhl.title.454</p>
<p>Mez, C.C. (1896) Bromeliaceae. <em>In</em>: de Candolle, C. (Ed.) <em>Monographiae phanerogamarum.</em> Vol. 9. Masson & Cie, Paris, pp. i–lxxvii,1–990. Available from: http://www.biodiversitylibrary.org/item/145130 (accessed 12 August 2019)</p>
<p>Mez, C.C. (1919) XVII - Additamenta Monographica. <em>Repertorium Specierum Novarum Regni Vegetabilis</em> 16: 65–79. https://doi.org/10.1002/fedr.19190160502</p>
<p>Mez, C.C. (1935) Bromeliaceae. <em>In</em>: Engler, A. (Ed.) <em>Das Pflanzenreich</em>. Vol. 4. W. Engelmann, Leipzig, pp. 362–667. Available from: http://www.biodiversitylibrary.org/item/56490 (accessed 12 August 2019)</p>
<p>Morren, É. (1875) Notice sur le <em>Vriesea platzmanni</em> sp. nov. ou <em>Vriesea</em> de M. Julius Platzmann. <em>La Belgique Horticole</em> 25: 349–351.</p>
<p>Morren, É. (1881) Description du <em>Vriesea chrysostachis</em>, sp. nov. de MM. J. Veitch, A. Londres. <em>La Belgique horticole </em>31: 87.</p>
<p>Niesler, I.M. (2016) Scanned Bromeliads – a Flash of Colours!. <em>Die Bromelie</em> 2016: 4–14.</p>
<p>Radford, A.E. (1986) <em>Fundamentals of Plant Systematics</em>. New York: Harper & Row, Publ. Inc., 498 pp.</p>
<p>Ramírez N.I. & Bevilacqua, M. (1990) Una nueva especie de <em>Tillandsia </em>(Bromeliaceae) Para la ciencia. <em>Acta Botánica Venezuélica</em> 15: 149–152.</p>
<p>Rauh, W. (1976) Neue <em>Vriesea</em>-Arten aus Peru und Ecuador. <em>Tropische und Subtropische Pflanzenwelt </em>16: 19–22.</p>
<p>Rauh, W. (1976) Bromelienstudien (Mitteilung 5). I. Neue und wenig bekannte Arten aus Peru und anderen Ländern. <em>Tropische und subtropische Pflanzenwelt</em> 16: 207–259.</p>
<p>Reed, C.F. & Read, R.W. (1981) Two New Species of Bromeliads from Venezuela. <em>Journal of the Bromeliad Society</em> 31: 59–61.</p>
<p>Ruiz, H.L. & Pavon, J.V. (1802) <em>Flora peruviana, et chilensis, sive descriptiones, et icones plantarum peruvianarum, et chilensium, secundum systema linnaeanum digestae, cum characteribus plurium generum evulgatorum reformatis.</em> Tomus 3. Madrid, Typis Gabrielis de Sancha, 95 pp.</p>
<p>Scharf, U. & Gouda, E.J. (2008) Bringing Bromeliaceae back to homeland botany. <em>Journal of the Bromeliad Society</em> 58: 123–129.</p>
<p>Smith, L.B. (1931) Bromeliaceae in H. A. Gleason. The Tyler-Duida-Expedition. <em>Bulletin of the Torrey Botanical Club</em> 58: 337–341. https://doi.org/10.2307/2480694</p>
<p>Smith, L.B. (1935) Studies in the Bromeliaceae - VI. <em>Contributions of the Gray Herbarium</em> 106: 147–220. https://doi.org/10.5962/p.336160</p>
<p>Smith, L.B. (1946) Bromeliaceae <em>In</em> Schultes, R. W. Plantae Austro-Americanae-III. <em>Botanical Museum Leaflets </em>12: 118–122. https://doi.org/10.5962/p.168447</p>
<p>Smith, L.B. (1948) Bromeliáceas notables de Colombia - IV. <em>Caldasia</em> 5: 1–15.</p>
<p>Smith, L.B. (1953) Notes on Bromeliaceae - I. <em>Phytologia</em> 4: 213–221.</p>
<p>Smith, L.B. (1955) Notes on Bromeliaceae -VI. <em>Phytologia</em> 5: 281–288. https://doi.org/10.5962/bhl.part.12982</p>
<p>Smith, L.B. (1955) Bromeliaceae. Plantae Colombianae XIII. <em>Botanical Museum Leaflets</em> 17: 67–77.</p>
<p>Smith, L.B. (1957) Bromeliaceae. The Botany of the Guiana Highland – Part II. <em>Memoirs of the New York Botanical Garden </em>9: 283–318.</p>
<p>Smith, L.B. (1959) Notes on Bromeliaceae - XI. <em>Phytologia </em>6 (8): 433–443<em>. </em>https://doi.org/10.5962/bhl.part.9778</p>
<p>Smith, L.B. (1963) <em>Vriesea soderstromii</em>. <em>Bulletin of the Bromeliad Society </em>13: 52–53. https://doi.org/10.1093/aibsbulletin/13.6.52</p>
<p>Smith, L.B. (1967) Notes on Bromeliaceae -XXVI. <em>Phytologia</em> 15: 163–200. https://doi.org/10.5962/bhl.part.21069</p>
<p>Smith, L.B. (1968) Notes on Bromeliaceae - XXVIII. <em>Phytologia</em> 16: 459–463. https://doi.org/10.5962/bhl.part.21075</p>
<p>Smith, L.B. (1969) Bromeliaceae. <em>In: Maguire </em>B. Botany of the Guayana Highland –Part VIII. <em>Memoirs of the New York Botanical Garden </em>18: 29–32.</p>
<p>Smith, L.B. (1970) Notes on Bromeliaceae - XXXI. <em>Phytologia</em> 20: 121–183. https://doi.org/10.5962/bhl.part.7116</p>
<p>Smith, L.B. & Read R.W. (1977) Notes on Bromeliaceae -XXXIX. <em>Phytologia</em> 38: 135–148.</p>
<p>Smith, L.B. & Downs, R.J. (1974) Pitcairnioideae (Bromeliaceae).<em> Flora Neotropica</em>. Monograph 14: 1–662.</p>
<p>Smith, L.B. & Downs, R.J. (1977) Bromeliaceae (Tillandsioideae). <em>Flora Neotropica Monograph </em>14: 663–1492.</p>
<p>Spencer, M.A. & Smith, L.B. (1993) <em>Racinaea</em>, a new genus of Bromeliaceae (Tillandsioideae). <em>Phytologia</em> 74: 151–160.</p>
<p>Stearn, W.T. (1973) <em>Botanical Latin,</em> ed. 2. London: David & Charles Newton Abbot, 566 pp.</p>
<p>Till, W. (2008) <em>Vriesea speckmaieri</em> – a New Lower-altitude Cloud Forest Species from Northern Venezuela. <em>Journal of the Bromeliad Society</em> 58: 250–254.</p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (Eds.) (2018) <em>International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017</em>. <em>Regnum Vegetabile</em> 159. Glasshütten: Koeltz Botanical Books. https://doi.org/10.12705/Code.2018</p>
<p>Ule, E. (1907 ‘1906’) II. Beiträge zur Flora der Hylaea nach den Sammlungen von Ule’s Amazonas-Expedition. <em>Verhandlungen des Botanischen Vereins für die Provinz Brandenburg </em>48: 117–208.</p>
<p>Versieux, L.M. & Wanderley, M.D.G.L. (2010) Delimitation of the <em>Alcantarea extensa complex</em> (Bromeliaceae) and a new species from Espírito Santo, Brazil. <em>Rodriguésia</em> 61: 421–429. https://doi.org/10.1590/2175-7860201061306</p>
<p>Wittmack, L. (1890) Plantae Lehmannianae in Guatemala, Costarica, Columbia, Ecuador etc. collectae. Bromeliaceae. <em>Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie </em>11: 52–71.</p>
<p>Wright, C.H. (1871) Flora Cubana. <em>Anales de la Academia de Ciencias Médicas, Físicas y Naturales de la Habana</em> 8: 48–56. [https://www.biodiversitylibrary.org/page/14603136#page/65/mode/1up]</p>