Abstract
Based on herbarium and field investigations, the taxonomic status of Solenopsis laurentia (Campanulaceae), hygrophilous species belonging to the subfam. Lobelioideae showing a Mediterranean distribution is here examined. According to literature, it can be considered as a species complex, within which several taxa at specific and infraspecific rank were described. Morphological and nomenclatural studies carried out on several living populations coming from many Italian localities allowed to recognize five subspecies. For each of them, a detailed description, iconography, micromorphology of the seed testa and pollen grains, taxonomical relationships, chorology, conservation status and nomenclatural notes are provided. The recognized subspecies of Solenopsis laurentia are the following: a) subsp. laurentia, which is the most widespread in Italy and also in the Mediterranean area; b) subsp. gasparrinii (Tineo) comb. nov., which is quite common in Sicily with scattered occurrence in other Mediterranean localities; c) subsp. caespitosa Brullo, it seems currently localized in Apulia and Sardinia; d) subsp. parvula Brullo, occurring in some localities of the Italian peninsula, Sardinia and Tunisia; e) subsp. hyblaea, described as new for science from Sicily with a punctiform distribution.
References
<p>Aldasoro, J.J., Castroviejo, S., Sales, F. & Hedge, I.C. (2001) ¿Qué es <em>Lobelia minima</em> Sims? <em>Anales Jardin Botanico de Madrid</em> 59: 173.</p>
<p>Aldridge, A. & Ortega, J. (1976) Estudios en la flora de Macaronesia: algunas numeros de cromosomas II. <em>Botanica Macaronesica</em> 2: 9–18.</p>
<p>Antonelli, A. (2008) Higher level phylogeny and evolutionary trends in Campanulaceae subfam. Lobelioideae: molecular signal overshadows morphology. <em>Molecular Phylogenetics and Evolution</em> 46 (1): 1–18. https://doi.org/10.1016/j.ympev.2007.06.015</p>
<p>Bachman, S., Moat, J., Hill, A.W., Torre, de la J. & Scott, B. (2011) Supporting Red List threat as-sessments with GeoCAT: geospatial conservation assessment tool. E-Infrastructures for data publishing in biodiversity science. <em>ZooKeys</em> 150: 117–126. https://doi.org/10.3897/zookeys.150.2109</p>
<p>Bagella, S., Caria, M.C., Farris, E. & Filigheddu, R. (2007) Issue related to the classification of Mediterranean temporary wet habitats accordind with the Europaean Union Habitatas Directive. <em>Fitosociologia </em>44 (2): 249–249.</p>
<p>Barthlott, W. (1981) Epidermal and seed surface characters of plants: systematic applicability and some evolutionary aspects. <em>Nordic Journal of Botany</em> 1: 345–355. https://doi.org/10.1111/j.1756-1051.1981.tb00704.x</p>
<p>Barthlott, W. (1984) Microstructural features of seed surface. <em>In</em>: Heywood, V.H. & Moore, D.A. (Eds.) <em>Current concepts in plant taxonomy</em>. Academic Press, London, pp. 95–105.</p>
<p>Battandier, J.A. (1916) Contributions à la Flore atlantique. <em>Bulletin de la Societè Botanique de la Société Botanique de France</em> 63: 188–196. https://doi.org/10.1080/00378941.1916.10835980</p>
<p>Battandier, J.A. (1889) Dicotyléd ones. <em>In</em>: Battandier, J.A. & Trabut, L. (Eds.) <em>Flore de l’Algerie</em>. Adolphe Jourdan, Alger, 825 pp.</p>
<p>Bivona-Bernardi, A. (1806) <em>Sicularum plantarum centuria prima</em>. Philippum Barravecchia, Panormi, 73 pp.</p>
<p>Boccone, P. (1697) <em>Museo di piante rare della Sicilia, Malta, Corsica, Italia, Piemonte, e Germania</em>. Baptista Zuccato, Venetia, 196 pp.</p>
<p>Boissier, R.E. (1875) <em>Flora Orientalis</em>, 3. H. Georg Bibliopolam, Genevae et Basileae, 1033 pp.</p>
<p>Bonali, F. & Romani, E. (2020) <em>Solenopsis laurentia</em> (L.) C. Presl. <em>In</em>: Alessandrini, A., Buono, V., Longo, D., Magni, C., Manni, Q.G. & Nicolella, G. (Eds.) <em>Acta Plantarum Notes</em> 7. Araba Fenice edizioni, Boves, 111 pp.</p>
<p>Brullo, C., Brullo, S. & Giusso Del Galdo, G. (2013) <em>Solenopsis mothiana</em> (Campanulaceae), a new species from Sicily. <em>Phytotaxa</em> 145 (1): 15–26. https://doi.org/10.11646/phytotaxa.145.1.2</p>
<p>Brullo, S., Brullo, C., Sciandrello, S., Tavilla, G., Ilardi, V. Cambria, S., Tomaselli, V., Giusso del Galdo, G. & Minissale, P. (2022) The plant communities of the class <em>Isoëto-Nanojuncetea</em> in Sicily. <em>Plants</em> 11 (9) 1214: 1–73. https://doi.org/10.3390/plants11091214</p>
<p>Brullo, S. & Guarino, R. (2018) <em>Solenopsis</em> C. Presl. <em>In</em>: Pignatti, S., Guarino, R. & La Rosa, M. (Eds.) <em>Flora d’Italia</em>, ed. 2. Edagricole, Milano, 735–736 pp.</p>
<p>Brullo, S. & Minissale, P. (1998) Considerazioni sintassonomiche sulla classe Isoeto-Nanojuncetea: <em>Itinera Geobotanica</em> 11: 263–290.</p>
<p>Brullo, S., Minissale, P. & Spampinato, G. (1995) Considerazioni fitogeografiche sulla flora della Sicilia. <em>Ecolologia Mediterranea</em> 21 (1/2): 99–117. https://doi.org/10.3406/ecmed.1995.1759</p>
<p>Brullo, S., Scelsi, F. & Siracusa, G. (1994) Contributo alla conoscenza della vegetazione terofitica delle Sicilia occidentale. <em>Bollettino della Accademia Gioenia di Scienze Naturali di Catania</em> 27 (346): 341–365.</p>
<p>Brummit, R.K. (2000) Report of the Commitee for Spermatophyta: 49. <em>Taxon</em> 49: 261–278. https://doi.org/10.2307/1223840</p>
<p>Burnett, G.T. (1835) <em>Outlines of Botany</em>. J. and C. Adlaed, London, 1068 pp.</p>
<p>Cabezudo, B. (1987) <em>Solenopsis</em>. <em>In</em>: Valdés, B., Talavera, S. & Fernández-Galiano, E. (Eds.) <em>Flora Vascular de Andalucia Occidental</em>, 2. Ketres, Barcelona, 640 pp.</p>
<p>Cambria, S., Brullo, S., Bogdanovic, S. & Giusso del Galdo, G. (2019) Taxonomy and vicariance within the endemic species of <em>Solenopsis</em> (Campanulaceae) in the Mediterranean islands. XVI OPTIMA Meeting, Abstracts, 2–5 October 2019, Athens: 67.</p>
<p>Candolle, A. de (1839) Lobeliaceae. <em>In</em>: Candolle, A.P. de (Ed.) <em>Prodromus systematis naturalis regni vegetabilis</em>, 7(2). Sociorum Treuttel et Würtz, Parisiis, pp. 339–413.</p>
<p>Castro, M. & Rosselló, J.A. (2005) Chromosome numbers in plant taxa endemic to the Balearic Islands. <em>Botanical Journal of Linnaean Society</em> 148: 219–228. https://doi.org/10.1111/j.1095-8339.2005.00397.x</p>
<p>Chapman, J.L. (1966) Comparative palynology in Campanulaceae. <em>Transactions of the Kansas Academy of Science </em>69 (3–4): 197–200. https://doi.org/10.2307/3627415</p>
<p>Christodoulou, C.S., Hand, R. & Iosif, K. (2020) <em>Solenopsis annua</em> comb. nov., a new taxon for Cyprus. <em>Cypricola</em> 17: 1–8.</p>
<p>Colmeiro, M. (1887) <em>Enumeracion y revision de las plantas de la Peninsula Hispano-Lusitanas e islas Baleares</em>, 3. Imprenta dela Viuda é Hija de Fuentenebro, Madrid, 545 pp.</p>
<p>Conti, F., Abbate, G., Alessandrini, A. & Blasi, C. (2005) <em>An annotated checklist of the Italian vascular flora</em>. Palombi Editore, Roma, 420 pp.</p>
<p>Coste, H. (1903) <em>Flore descriptive et illustrée de la France de la Corse et des contrées limitrophes</em>. Paul Klincksieck, Paris, 627 pp. https://doi.org/10.1080/00378941.1903.10831075</p>
<p>Coutinho, A.X.P. (1913) <em>A Flora de Portugal (Plantäs vasculares)</em>. Aillaud Alves & C, Paris, 766 pp.CRESP M. B., SERRA, L., TURLAND, N., 1996: Lec–toe<em> liaceae—</em>Taxon 45: 117–120. https://doi.org/10.2307/1222596</p>
<p>Crespo, M.B., Serra, L. & Juan, A. (1998) <em>Solenopsis</em> (Lobeliaceae): a genus endemic in the Mediterranean Region. <em>Plant Systematic and Evolution</em> 210: 211–229. https://doi.org/10.1007/BF00985669</p>
<p>Crespo, M.B., Serra, L. & Juan, A. (2007) Notes on <em>Lobelia minima</em> Sims (Campanulaceae—Lobelioideae): what it is and is not. <em>Taxon</em> 56: 243–246.</p>
<p>Damboldt, J. (1978) <em>Laurentia</em>. <em>In</em>: Davis, P.H. (Ed.) <em>Flora of Turkey and east Aegean Islands</em> 6. Edinburgh University Press, Edinburgh, p. 1.</p>
<p>Deyuan, H. & Lammers, T.G. (2011) <em>Lobelia.</em> <em>In</em>: Wu, Z.Y., Raven, P.H. & Hong, D.Y. (Eds.) <em>Flora of China</em>, 11. Missouri Botanical Garden Press, St.Louis, pp. 554–562.</p>
<p>Dimopoulos, P., Raus, T., Bergmeier, E., Costantinidis, T., Iatrou, G., Kokkini, S., Strid, A. & Tizanoudakis, D. (2013) Vascular plants of Greece. An annotated checklist. <em>Englera</em> 31: 1–372.</p>
<p>Dunbar, A. (1975) On the pollen of Campanulaceae and related families with special reference to the surface ultrastructure. II. Campaulaceae Subfam. Cyphoideae and Subfam. Lobelioideae; Goodeniaceae, Sphenocleaceae. <em>Botaniska Notiser</em> 128: 102–118.</p>
<p>Dunbar, A. (1984) Pollen morphology in Campanulaceae. IV. <em>Nordic Journal of Botany</em> 4: 1–19. https://doi.org/10.1111/j.1756-1051.1984.tb01961.x</p>
<p>Emberger, M.L. & Maire, R. (1930) Plantae maroccanae novae vel minus cognitae (Fasciculus tertius). <em>In</em>: Litardier, R. de & Maire, R. (Eds.) <em>Plantae maroccanae novae</em>. Rabat, pp. 1–8.</p>
<p>Endlicher, S. (1838) <em>Genera plantarum secundum ordines naturales disposita</em>. F.R. Beck, Universitatis Bibliopolam, Vindobonae, pp. 401–604.</p>
<p>European Commission (1992) Council Directive 92/43/EEC on the conservation of natural habitats and of wild faunaand flora. <em>European Community Gazette</em> 206: 1–50.</p>
<p>European Commission (2007) <em>Interpretation manual of European Union habitats. Eur 27.</em> European Commission, DG Environment, Strasbourg, 142 pp.</p>
<p>Evans, D. (2006) The habitats of the European Union Habitats Directive. Biology and Environment: <em>Proceedings of the Royal Irish Academy</em> 106 B (3): 167–173. https://doi.org/10.1353/bae.2006.0032</p>
<p>Fennane, M. & Mathez, J. (2014) Campanulaceae. <em>In</em>: Fennane, M., Ibn-Tattou, M. & El Qualidi, J. (Eds.) <em>Flora pratique du Maroc. Manuel de détermination des plantes vasculaires</em>, 3. Cana Print, Rabat, pp. 65–76.</p>
<p>Fiori, A. (1933) <em>Iconographia florae Italicae ossia Flora Italiana illustrata</em>. Mariano Ricci, Firenze, 549 pp.</p>
<p>Gargano, D. (2011) Verso la redazione di nuove Liste Rosse della flora d’Italia: una griglia standard per la misura dell’<em>Area of Occupancy (AOO). Informatore Botanico Italiano </em>43 (2): 455–458.</p>
<p>Gontcharova, S.B., Gontcharov, A.A., Yakubov, V.V. & Kondo, K. (2009) Seed surface morphology in some representatives of the genus <em>Rhodiola</em> sect. <em>Rhodiola</em> (<em>Crassulaceae</em>) in the Russian Far East. <em>Flora</em> 204: 17–24. https://doi.org/10.1016/j.flora.2008.01.009</p>
<p>Greuter, W. (1981) Med-Checklist notulae, 3. <em>Willdenowia </em>11: 23–43.</p>
<p>Greuter, W., Matthäs, U. & Risse, H. (1984) Additions to the flora of Crete, 1973–1983, I. <em>Willdenowia </em>14: 27–36.</p>
<p>Hand, R. (2006) Supplementary notes to the flora of Cyprus V. <em>Willdenowia</em> 36: 761–809. https://doi.org/10.3372/wi.36.36211</p>
<p>Haridasan, V.K. & Mukherjee, P.K. (1993) Seed surface features in Indian Lobeliaceae. <em>Phytomorphology</em> 43: 287–294.</p>
<p>Hesse, M., Halbritter, H., Zetter, R., Weber, M., Buchner, R., FroschRadivo, A. & Ulrich, S. (2009) <em>Pollen terminology. An illustrated handbook. </em>Springer, Wein - New York, 264 pp.</p>
<p>Hoffmannsegg Comte de, J.C. & Link, H.F. (1820) <em>Flora Portugaise</em>, 2. Imprimeria des’Auters, Berlin, 504 pp.</p>
<p>IUCN (2012) <em>IUCN Red List Categories and Criteria: Version 3.1. Second edition. Gland, </em>Switzerland and Cambridge, UK, IUCN. iv + 32 pp.</p>
<p>Lange, J. (1861) <em>Pugillus plantarum imprimis hispanicarum, quas in itinere 1851</em>–<em>52 legit</em>, 2. Bianco Luno, Hafniae, pp. 83–164. https://doi.org/10.5962/bhl.title.10857</p>
<p>Jussieu de, A.L. (1789) <em>Genera Plantarum secundum ordines naturales disposita</em>. Viduam Herissant et Theophilum Barrois, Parisiis, pp. 1–498.</p>
<p>Kagame, S.P., Gichira, A.W., Chen, L.-Y. & Wang, Q.-F. (2021) Systematics of Lobelioideae (Campanulaceae): review, phylogenetic and biogeographic analyses. <em>PhytoKeys</em> 174: 13–45. https://doi.org/10.3897/phytokeys.174.59555</p>
<p>Knox, E.B. & Li, C. (2017) The East Asian origin of the giant Lobelioids. <em>American Journal of Botany </em>104 (6): 924–938. https://doi.org/10.3732/ajb.1700025</p>
<p>Lammers, T.G. (1993) Chromosome numbers of Campanulaceae. Review and integration of data for subfamily LobeIioideae. <em>American Journal of Botany</em> 80: 660–675. https://doi.org/10.1002/j.1537-2197.1993.tb15237.x</p>
<p>Lammers, T.G. (2007) <em>World checklist and bibliography of Campanulaceae</em>: The Board of Trustees of the Royal Botanic Gardens, Kew, 685 pp.</p>
<p>Lammers, T.G. (2011) Revision of the infrageneric classification of <em>Lobelia</em> L. (Campanulaceae: Lobelioideae). <em>Annals of the Missouri Botanical Garden</em> 98: 37–62. https://doi.org/10.3417/2007150</p>
<p>Lange, J. (1861) <em>Pugillus plantarum imprimis hispanicarum, quas in itinere 1851–52 legit,</em> 2. Bianco Luno, Hafniae, pp. 83–164. https://doi.org/10.5962/bhl.title.10857</p>
<p>Lee, S., Chung, Y.J. & Lee, J. (1988) A palynotaxonomic study of the Korean Campanulaceae. <em>Korean Journal of Plant Taxonomy</em> 18: 115–131. https://doi.org/10.11110/kjpt.1988.18.2.115</p>
<p>Le Floc’H, E., Boulos, L. & Vela, E. (2010) Catalogue synonymique commenté de la Flore de Tunisie. Banque Nationale de Gènes de la Tunisie, Tunis, 500 pp.</p>
<p>Linnaeus, C. (1753) <em>Species Plantarum,</em> 2. Laurentii Salvii, Holmiae, pp. 561–1200.</p>
<p>Linnaeus, C. (1754) <em>Genera Plantarum,</em> ed. 5. Laurentii Salvii, Holmiae, pp. 1–522.</p>
<p>Linnaeus, C. (1771) <em>Mantissa Plantarum Altera</em>. Laurentii Salvii, Holmiae, pp. 1–588.</p>
<p>Meikle, R.D. (1979) Some notes on <em>Laurentia </em>Adanson (Campanulaceae). <em>Kew Bulletin</em> 34: 373–375. https://doi.org/10.2307/4110000</p>
<p>Micheli, P.A. (1729) <em>Nova plantarum genera iuxta Tournefortii methodum disposita</em>. Bernardi Peperinii, Florentiae, 234 pp.</p>
<p>Merino, B. (1906) <em>Flora descriptiva é illustrata de Galicia</em>. Tipografía Galaica, Santiago, 634 pp. https://doi.org/10.5962/bhl.title.10911</p>
<p>Mouterde, P. (1978) <em>Nouvelle Flore du Liban et de la Syrie, </em>3(3). Dar El-Machreq, Beyrouth, pp. 205–364.</p>
<p>Murata, J. (1992) Systematic implications of seed coat morphology in <em>Lobelia </em>(Campanulaceae—Lobelioideae). <em>Journal of the Faculty of Science, University of Tokyo, Section III, Botany</em> 15: 155–172.</p>
<p>Murata, J. (1995) A revision of infrageneric classification of <em>Lobelia</em> (Campanulaceae—Lobelioideae) with special reference to seed coat morphology. <em>Journal of the Faculty of Science, University of Tokyo, Section III, Botany</em> 15: 349–371.</p>
<p>Peruzzi, L. & Gargano, D. (2003) Reports 13: 1357–1361. Mediterranean chromosome number. <em>Flora Mediterranea</em> 13: 382–387.</p>
<p>Petermann, W.L. (1845) Das Pflanzenreich in vollständigen Beschreibungen dargestellt, nach dem natürlichen Systeme geordnet und in naturgetreuen Abbildungen gezeichnet. Eduard Eisenach, Leipzig, 1010 pp.</p>
<p>Pignatti, S. (1982) <em>Laurentia.</em> <em>In</em>: Pignatti, S. (Ed.) <em>Flora d’Italia</em>, vol. 2. Edagricole, Bologna. p. 712.</p>
<p>Portal to the Flora of Italy (2022) Portal to the Flora of Italy. Available from: http://dryades.units.it/floritaly (accessed 7 March 2022)</p>
<p>Presl, C.B. (1831) <em>Symbolae botanicae, sive, Descriptiones et icones plantarum novarum aut minus cognitarum</em>. J. Spurny, Pragae, 205 pp. https://doi.org/10.5962/bhl.title.12413</p>
<p>Presl, C.B. (1836) <em>Prodromus monographiae Lobeliacearum</em>. Filiorum Theophili Haase, Prague, pp. 1–52.</p>
<p>Punt, W., Blackmore, S., Nilsson, S. & Le Thomas, A. (1994) <em>Glossary of pollen and spore terminology</em>. Lpp Foundation, Utrecht.</p>
<p>Punt, W., Hoen, P.P., Blackmore, S., Nilsson, S. & Le Thomas, A. (2007) Glossary of pollen and spore terminology. <em>Review of Palaeob otanyand Palynology</em> 143: 1–81. https://doi.org/10.1016/j.revpalbo.2006.06.008</p>
<p>Quezel, P. & Santa, S. (1963) <em>Nouvelle Flore d’Algérie et des Régions Désertiques Méridionales</em>, 2. Editions CNRS, Paris, pp. 571–1170.</p>
<p>Ray, J. (1704) <em>Historia plantarum tomus tertius: qui est Supplementum duorum precedentium</em>. Sam. Smith & Benj. Walford., Londini, 666 pp.</p>
<p>R Core Team (2021) <em>R: A language and environment for statistical computing</em>. R Foundation for Statistical Computing, Vienna, Austria. [https://www.R-project.org/]</p>
<p>Requien, E. (1825) Observations sur quelques plantes rares ou nouvelles de la Flore Française. <em>Annales des Sciences Naturelles</em> 5: 381–387.</p>
<p>Rohlf, F.J. (2000) <em>NTSYSpc Numerical Taxonomy and Multivariate Analysis System</em>, vers. 2.1. New York: Exeter Software.</p>
<p>Romano, S., Mazzola, R. & Raimondo, E.M. (1986) Numeri cromosomici per la flora Italiana: 1070–1081. <em>Informatore Botanico Italiano</em> 18: 159–167.</p>
<p>Sales, F. & Hedge, I.C. (2001) <em>Solenopsis</em> C. Presl. <em>In</em>: Paiva, J., Sales, F., Hedge, I.C., Aedo, C., Aldasoro, J.J., Castroviejo, S., Herrero, A. & Velayos, M. (Eds.) <em>Flora Iberica</em>, 14. Real Jardín Botánico, CSIC, Madrid, pp. 173–175.</p>
<p>Salisbury, R.A. (1796) <em>Prodromus Stirpium in Horto ad Chapel Allerton vigentium</em>. Londini, 422 pp. https://doi.org/10.5962/bhl.title.427</p>
<p>Sapienza, G., Hilton D.R. & Scribano, V. (2005) Helium isotope in pteridotite mineral phade from Hyblean Plateau xenoliths (south-eastern Sicily, Italy) <em>Chemical Geology</em> 219: 115–129. https://doi.org/10.1016/j.chemgeo.2005.02.012</p>
<p>Scheidweiler, M.J.F. (1841) Einiger neuen Pflanzen. <em>Allgemeine Gartenzeitung</em> 9: 1–414.</p>
<p>Serra, L. & Crespo, M.B. (1997) An outline revision of the subtribe Siphocampylinae (Lobeliaceae). <em>Lagascalia </em>19: 881–888.</p>
<p>Sibthorp, J. & Smith, J.E. (1806)<em> Florae Graecae prodromus</em>, 1. Richardi Taylor et Socii, Londini, 442 pp.</p>
<p>Strobl, G. (1883) Flora der Nebroden (Fortsetzung). <em>Flora</em> 66 (35): 547–554.</p>
<p>Takhtajan, A. (2009) <em>Flowering Plants</em>, 2 ed. Springer, Dordrecht, 871 pp. https://doi.org/10.1007/978-1-4020-9609-9</p>
<p>Thiers, B. (2021) <em>Index Herbariorum: A global directory of public herbaria and associated staff</em>. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/ih/ (continuously updated; accessed 8 May 2021)</p>
<p>Tineo, V. (1827) <em>Catalogus plantarum horti regii panormitani ad annum 182</em>7. Regali Tipographia, Panormi, pp.1–284.</p>
<p>Tison, J.M. & de Foucault, B. (2014) <em>Flora Gallica: flore de France</em>. Biotope éditions, Mèze, 1195 pp.</p>
<p>Todaro, A. (1873) Adnotationes ad Indicem Seminum horti regii botanici Panormitani ann. MDCCCLXXII. <em>Nuovo Giornale Botanico Italiano</em> 5: 156–160.</p>
<p>Tomaselli, V., Beccarisi, L., Brullo, S., Cambria, S., Forte, L., Minissale, P. & Veronico, G. (2020) Phytosociological research on temporary ponds in Apulia (southern Italy). <em>Mediterranean Botany</em> 41 (1): 15–41. https://doi.org/10.5209/mbot.63617</p>
<p>Torrey, J. (1857) Explorations and surveys for a railroad route from the Mississippi River to the Pacific Ocean, route near the thirty-fifth parallel, explored by lieutenant A. W. Whipple, topographical engineers, in 1853 and 1854. Report on the botany of the expedition. <em>Pacific Railroad Survey Report </em>4: 59–182. </p>
<p>Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) <em>Regnum Vegetabile 159. Glashütten: Koeltz Botanical Books.</em> https://doi.org/10.12705/Code.2018</p>
<p>Tutin, T.G. (1976) <em>Laurentia</em>. <em>In</em>: Tutin, T.G., Heywood, V.H., Burges, N.A., Moore, D.M., Valentine, D.H., Walters, S.M., Webb, D.A., (Eds.) <em>Flora Europaea</em>, 4. Cambridge University Press, Cambridge, p. 102.</p>
<p>Verlaque, R., Aboucaya, A., Noble, V., Hardion, L. & Vila, B. (2016) Reports. IAPT/IOPB chromosome data 22. <em>Taxon</em> 65 (5): E24–E27. https://doi.org/10.12705/655.40</p>
<p>Wimmer, F.E. (1948) Vorarbeiten zur Monographie der Campanulaceae—Lobelioideae. II. Trib. Lobelieae. <em>Annalen des Naturhistorischen Museums in Wien</em> 56: 317–347.</p>
<p>Walker, J.W. & Doyle, J.A. (1975) The bases of angiosperm phylogeny: palynology. <em>Annales of Missouri Botanical Garden</em> 62: 664–723. https://doi.org/10.2307/2395271</p>