Abstract
During recent fieldwork at Azad Jammu and Kashmir (Pakistan), many basidiomycetes of the genus Cortinarius were collected from mixed coniferous Himalayan temperate forests. Their morphological characterization and phylogenetic analyses based on ITS and LSU sequence data yielded a well-delimited, new species within C. subgenus Telamonia sect Disjungendi. It is characterized by producing small and dry basidiomata with blackish brown conico-convex pilei when young and uplifted at maturity, dark brown lamellae, a fibrillose stipe and small basidiospores (9.4 × 5.5 µm in average).
References
<p>Ahmad, S., Iqbal, S.H. & Khalid, A.N. (1997) <em>Fungi of Pakistan.</em> Lahore, Pakistan: Sultan Ahmad Mycological Society.</p>
<p>Altschul, S.F., Madden, T.L., Schäffer, A.A., Zhang, J., Zhang, Z., Miller, W. & Lipman, D.J. (1997) Gapped BLAST and PSI-BLAST: a new generation of protein database search programs. <em>Nucleic Acids Research</em> 25: 3389–3402. https://doi.org/10.1093/nar/25.17.3389</p>
<p>Brandrud, T.E., Gorbunova, I.A., Ageev, D.V., Dahlberg, A., Dima, B., Morozova, O.V. & Svetasheva, T. (2019) New data on <em>Cortinarius funga</em> (Agaricales, Basidiomycota) from Altaiskiy Nature Reserve and Gorno-Altaysk area (SW Siberia, Russia). <em>Mycology and Plant Pathology</em>: 325–341. https://doi.org/10.1134/S0026364819060035</p>
<p>Cubeta, M.A., Echandi, E., Abernethy, T. & Vilgalys, R. (1991) Characterization of anastomosis groups of binucleate Rhizoctonia species using restriction analysis of an amplified ribosomal RNA gene. <em>Phytopathology</em> 81: 1395–1400. https://doi.org/10.1094/Phyto-81-1395</p>
<p>Frøslev, T.G., Matheny, P.B. & Hibbett, D.S. (2005) Lower level relationships in the mushroom genus <em>Cortinarius</em> (Basidiomycota, Agaricales): a comparison of RPB1, RPB2, and ITS phylogenies. <em>Molecular Phylogenetics and Evolution</em> 37: 602–618. https://doi.org/10.1016/j.ympev.2005.06.016</p>
<p>Frøslev, T.G., Jeppesen, T.S., Læssøe, T. & Kjøller, R. (2007) Molecular phylogenetics and delimitation of species in <em>Cortinarius </em>section <em>Calochroi </em>(Basidiomycota, Agaricales) in Europe. <em>Molecular phylogenetics and evolution</em> 44: 217–27. https://doi.org/10.1016/j.ympev.2006.11.013</p>
<p>Gardes, M. & Bruns, T.D. (1993) ITS primers with enhanced specificity for basidiomycetes application to the identification of mycorrhizae and rusts. <em>Molecular Ecology</em> 2: 113–118. https://doi.org/10.1111/j.1365-294X.1993.tb00005</p>
<p>Garnica, S., Schön, M.E., Abarenkov, K., Riess, K., Liimatainen, K., Niskanen, T., Dima, B., Soop, K., Frøslev, T.G., Jeppesen, T.S. & Peintner, U. (2016) Determining threshold values for barcoding fungi: lessons from <em>Cortinarius </em>(Basidiomycota), a highly diverse and widespread ectomycorrhizal genus. FEMS <em>Microbiology Ecology</em> 92: 4. https://doi.org/10.1093/femsec/fiw045</p>
<p>Garnica, S., Weiß, M., Oertel, B. & Oberwinkler, F. (2005) A framework for a phylogenetic classifcation in the genus <em>Cortinarius </em>(Basidiomycota, Agaricales) derived from morphological and molecular data. <em>Canadian Journal of Botany</em> 83: 1457–1477. https://doi.org/10.1139/b05-107</p>
<p>He, M.Q., Zhao, R.L., Hyde, K.D., Begerow, D. & Kirk, P.M. (2019) Notes, outline and divergence times of Basidiomycota. <em>Fungal diversity</em> 99 (1): 105–367. https://doi.org/10.1007/s13225-019-00435-4</p>
<p>Katoh, K. & Standley, D.M. (2013) MAFFT: Multiple Sequence Alignment Software version 7: improvements in performance and usability. <em>Molecular Biology and Evolution</em> 30: 772–780. https://doi.org/10.1093/molbev/mst010</p>
<p>Khan, J., Sher, H., Hussain, S. & Khalid, A.N. (2020) <em>Cortinarius brunneocarpus</em> var. <em>microsporus,</em> an interesting variety with Eurasian distribution. <em>Plant Systematics and Evolution</em> 306 (5): 1–9. https://doi.org/10.1007/s00606-020-01703-6</p>
<p>Kirk, P.M., Cannon, P.F., Minter, D.W. & Stalpers, J.A. (2008) <em>Dictionary of the fungi</em>, 10th edn. CAB International, Wallingford, UK.</p>
<p>Li, Y., Li, T.H., Yang, Z.L., Bau, T. & Dai, Y.C. (2015) Atlas of Chinese Macrofungal Resources. Zhongyuan Farmers Press, Zhengzhou, 1351 pp.</p>
<p>Liimatainen, K. (2014) <em>Nomenclatural novelties</em>. Index Fungorum no. 196.</p>
<p>Liimatainen, K. & Ainsworth, A.M. (2018) Fifteen <em>Cortinarius </em>species associated with Helianthemum in Great Britain: results of a DNA-based analysis. <em>Field mycology</em> 19 (4): 119–135. https://doi.org/10.1016/j.fldmyc.2018.10.006</p>
<p>Liimatainen, K., Niskanen, T., Ammirati, J.F., Kytövuori, I. & Dima, B. (2015) <em>Cortinarius,</em> subgenus Telamonia, section Disjungendi, cryptic species in North America and Europe. <em>Mycological progress</em> 14 (1): 1–8. https://doi.org/10.1007/s11557-014-1016-9</p>
<p>Liimatainen, K., Niskanen, T., Dima, B., Kytövuori, I., Ammirati, J.F. & Frøslev, T.G. (2014) The largest type study of Agaricales species to date: bringing identification and nomenclature of <em>Phlegmacium (Cortinarius)</em> into the DNA era. <em>Persoonia-Molecular Phylogeny and Evolution of Fungi </em>33 (1): 98–140. https://doi.org/10.3767/003158514X684681</p>
<p>Luo, Y. & Bau, T. (2021) <em>Cortinarius jiaoheensis</em> (Cortinariaceae), a new species of <em>Cortinarius </em>subgenus <em>Telamonia </em>section <em>Flexipedes,</em> from northeast China. <em>Phytotaxa </em>494 (1): 113–121. https://doi.org/10.11646/phytotaxa.494.1.7</p>
<p>Lanfear, R., Calcott, B., Ho, S.Y. & Guindon, S. (2012) Partition Finder: combined selection of partitioning schemes and substitution models for phylogenetic analyses. <em>Molecular biology and evolution</em> 29 (6): 1695–1701. https://doi.org/10.1093/molbev/mss020</p>
<p>Miller, M.A., Pfeiffer, W. & Schwartz, T. (2010) <em>Creating the CIPRES Science Gateway for inference of large phylogenetic trees. Proceedings of the Gateway Computing Environments Workshop</em> <em>(GCE)</em>. New Orleans, Louisiana. 1–8. https://doi.org/10.1109/GCE.2010.5676129</p>
<p>Naseer, A., Garrido-Benavent, I., Khan, J., Ballarà, J., Mahiques, R., Khalid, A.N. & Sher, H. (2020) <em>Cortinarius pakistanicus</em> and <em>C. pseudotorvus</em>: two new species in oak forests in the Pakistan Himalayas. <em>MycoKeys</em> 74: 91. https://doi.org/10.3897/mycokeys.74.49734</p>
<p>Niskanen, T. (2014) <em>Nomenclatural novelties</em>. Index Fungorum no. 186.</p>
<p>Niskanen, T., Kytövuori, I. & Liimatainen, K. (2011) <em>Cortinarius </em>sect. Armillati in northern Europe. <em>Mycologia </em>103 (5): 1080–1101. https://doi.org/10.3852/10-350</p>
<p>Niskanen, T., Liimatainen, K. & Kytövuori, I. (2015) Nomenclatural novelties 256. Index Fungorum, (256). Available from: http://www. indexfungorum.org/Publications/Index%20Fungorum%20no.256.pdf (accessed 20 November 2020)</p>
<p>Niskanen, T., Liimatainen, K., Kytövuori, I. & Lindström, H. (2012) Subgen. <em>Telamonia</em> (Fr.) Trog. <em>In</em>: Knudsen, H. & Vesterholt, J. (Eds.) <em>Funga Nordica</em>, 2nd revised edition. Agaricoid, boletoid, clavarioid, cyphelloid and gastroid genera. Nordsvamp, Denmark, pp. 826–885.</p>
<p>Nuske, S.J., Anslan, S., Tedersoo, L., Congdon, B.C. & Abell, S.E. (2019) Ectomycorrhizal fungal communities are dominated by mammalian dispersed truffle-like taxa in north-east Australian woodlands. <em>Mycorrhiza </em>29 (3): 181–193. https://doi.org/10.1007/s00572-019-00886-2</p>
<p>Peintner, U., Moncalvo, J.M. & Vilgalys, R. (2004) Toward a better understanding of the infrageneric relationships in <em>Cortinarius </em>(Agaricales, Basidiomycota). <em>Mycologia </em>96: 1042–1058. https://doi.org/10.1080/15572536.2005.11832904</p>
<p>Ronquist, F., Teslenko, M., Van Der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. <em>Systematic biology</em> 61 (3): 539–542. https://doi.org/10.1093/sysbio/sys029</p>
<p>Saba. M., Khalid, A.N., Jabeen, S. & Dima, B. (2017) <em>Cortinarius longistipitatus,</em> a new species in subgenus <em>Telamonia,</em> section <em>Cinnabarini,</em> from Pakistan. <em>Phytotaxa</em> 328 (3): 257–266. https://doi.org/10.11646/phytotaxa.328.3.4</p>
<p>Song, J., Liang, J.-F., Mehrabi-Koushki, M., Krisai-Greilhuber, I., Ali, B., Bhatt, V.K., Cerna-Mendoza, A., Chen, B., Chen, Z.-X., Chu, H.-L., Corazon-Guivin, M.A., da Silva, G.A., De Kesel, A., Dima, B., Dovana, F., Farokhinejad, R., Ferisin, G., Guerrero-Abad, J.C., Guo, T., Han, L.-H., Ilyas, S., Justo, A., Khalid, A.N., Khodadadi-Pourarpanahi, S., Li, T.-H., Liu, C., Lorenzini, M., Lu, J.-K., Mumtaz, A.S., Oehl, F., Pan, X.-Y., Papp, V., Qian, W., Razaq, A., Semwal, K.C., Tang, L.-Z., Tian, X.-L., Vallejos-Tapullima, A., van der Merwe, N.A., Wang, S.-K., Wang, C.-Q., Yang, R.-H., Yu, F., Zapparoli, G., Zhang, M., Antonín, V., Aptroot, A., Aslan, A., Banerjee, A., Chatterjee, S., Dirks, A.C., Ebrahimi, L., Fotouhifar, K.-B., Ghosta, Y., Kalinina, L.B., Karahan, D., Liu, J., Maiti, M.K., Mookherjee, A., Nath, P.S., Panja, B., Saha, J., Ševčíková, H., Voglmayr, H., Yazıcı, K. & Haelewaters, D. (2019) Fungal Systematics and Evolution: FUSE 5. <em>Sydowia </em>71: 141–245. https://doi.org/10.1093/sysbio/sys029</p>
<p>Stamatakis, A. (2014) RAxML version 8: a tool for phylogenetic analysis and postanalysis of large phylogenies. Bioinformatics. https://doi.org/10.1093/bioinformatics/btu033</p>
<p>Soop, K., Dima, B., Cooper, J.A., Park, D. & Oertel, B. (2019) A phylogenetic approach to a global supraspecifc taxonomy of <em>Cortinarius </em>(Agaricales) with an emphasis on the southern mycota. <em>Persoonia</em> 42: 261–290. https://doi.org/10.3767/persoonia.2019.42.10</p>
<p>Vellinga, E.C., Sysouphanthong, P. & Hyde, K.D. (2011) The family Agaricaceae: phylogenices and two new white spored genera. <em>Mycologia</em> 103 (3): 494–509. https://doi.org/10.3852/10-204</p>
<p>Vilgalys, R. & Hester, M. (1990) Rapid genetic identification and mapping of enzymatically amplified ribosomal DNA from several Cryptococcus species. <em>Journal of Bacteriology</em> 172 (8): 4238–4246. https://doi.org/10.1128/JB.172.8.4238-4246.1990</p>
<p>Teasdale, S.E., Beulke, A.K., Guy, P.L. & Orlovich, D.A. (2013) Environmental barcoding of the ectomycorrhizal fungal genus <em>Cortinarius.</em> <em>Fungal Diversity</em> 58: 299–310. https://doi.org/10.1007/s13225-012-0218-1</p>
<p>White, T.J, Bruns T, Lee, S.J. & Taylor, J. (1990) Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. <em>PCR protocols: a guide to methods and applications</em> 18 (1): 315–322. https://doi.org/10.1016/B978-0-12-372180-8.50042-1</p>
<p>Altschul, S.F., Madden, T.L., Schäffer, A.A., Zhang, J., Zhang, Z., Miller, W. & Lipman, D.J. (1997) Gapped BLAST and PSI-BLAST: a new generation of protein database search programs. <em>Nucleic Acids Research</em> 25: 3389–3402. https://doi.org/10.1093/nar/25.17.3389</p>
<p>Brandrud, T.E., Gorbunova, I.A., Ageev, D.V., Dahlberg, A., Dima, B., Morozova, O.V. & Svetasheva, T. (2019) New data on <em>Cortinarius funga</em> (Agaricales, Basidiomycota) from Altaiskiy Nature Reserve and Gorno-Altaysk area (SW Siberia, Russia). <em>Mycology and Plant Pathology</em>: 325–341. https://doi.org/10.1134/S0026364819060035</p>
<p>Cubeta, M.A., Echandi, E., Abernethy, T. & Vilgalys, R. (1991) Characterization of anastomosis groups of binucleate Rhizoctonia species using restriction analysis of an amplified ribosomal RNA gene. <em>Phytopathology</em> 81: 1395–1400. https://doi.org/10.1094/Phyto-81-1395</p>
<p>Frøslev, T.G., Matheny, P.B. & Hibbett, D.S. (2005) Lower level relationships in the mushroom genus <em>Cortinarius</em> (Basidiomycota, Agaricales): a comparison of RPB1, RPB2, and ITS phylogenies. <em>Molecular Phylogenetics and Evolution</em> 37: 602–618. https://doi.org/10.1016/j.ympev.2005.06.016</p>
<p>Frøslev, T.G., Jeppesen, T.S., Læssøe, T. & Kjøller, R. (2007) Molecular phylogenetics and delimitation of species in <em>Cortinarius </em>section <em>Calochroi </em>(Basidiomycota, Agaricales) in Europe. <em>Molecular phylogenetics and evolution</em> 44: 217–27. https://doi.org/10.1016/j.ympev.2006.11.013</p>
<p>Gardes, M. & Bruns, T.D. (1993) ITS primers with enhanced specificity for basidiomycetes application to the identification of mycorrhizae and rusts. <em>Molecular Ecology</em> 2: 113–118. https://doi.org/10.1111/j.1365-294X.1993.tb00005</p>
<p>Garnica, S., Schön, M.E., Abarenkov, K., Riess, K., Liimatainen, K., Niskanen, T., Dima, B., Soop, K., Frøslev, T.G., Jeppesen, T.S. & Peintner, U. (2016) Determining threshold values for barcoding fungi: lessons from <em>Cortinarius </em>(Basidiomycota), a highly diverse and widespread ectomycorrhizal genus. FEMS <em>Microbiology Ecology</em> 92: 4. https://doi.org/10.1093/femsec/fiw045</p>
<p>Garnica, S., Weiß, M., Oertel, B. & Oberwinkler, F. (2005) A framework for a phylogenetic classifcation in the genus <em>Cortinarius </em>(Basidiomycota, Agaricales) derived from morphological and molecular data. <em>Canadian Journal of Botany</em> 83: 1457–1477. https://doi.org/10.1139/b05-107</p>
<p>He, M.Q., Zhao, R.L., Hyde, K.D., Begerow, D. & Kirk, P.M. (2019) Notes, outline and divergence times of Basidiomycota. <em>Fungal diversity</em> 99 (1): 105–367. https://doi.org/10.1007/s13225-019-00435-4</p>
<p>Katoh, K. & Standley, D.M. (2013) MAFFT: Multiple Sequence Alignment Software version 7: improvements in performance and usability. <em>Molecular Biology and Evolution</em> 30: 772–780. https://doi.org/10.1093/molbev/mst010</p>
<p>Khan, J., Sher, H., Hussain, S. & Khalid, A.N. (2020) <em>Cortinarius brunneocarpus</em> var. <em>microsporus,</em> an interesting variety with Eurasian distribution. <em>Plant Systematics and Evolution</em> 306 (5): 1–9. https://doi.org/10.1007/s00606-020-01703-6</p>
<p>Kirk, P.M., Cannon, P.F., Minter, D.W. & Stalpers, J.A. (2008) <em>Dictionary of the fungi</em>, 10th edn. CAB International, Wallingford, UK.</p>
<p>Li, Y., Li, T.H., Yang, Z.L., Bau, T. & Dai, Y.C. (2015) Atlas of Chinese Macrofungal Resources. Zhongyuan Farmers Press, Zhengzhou, 1351 pp.</p>
<p>Liimatainen, K. (2014) <em>Nomenclatural novelties</em>. Index Fungorum no. 196.</p>
<p>Liimatainen, K. & Ainsworth, A.M. (2018) Fifteen <em>Cortinarius </em>species associated with Helianthemum in Great Britain: results of a DNA-based analysis. <em>Field mycology</em> 19 (4): 119–135. https://doi.org/10.1016/j.fldmyc.2018.10.006</p>
<p>Liimatainen, K., Niskanen, T., Ammirati, J.F., Kytövuori, I. & Dima, B. (2015) <em>Cortinarius,</em> subgenus Telamonia, section Disjungendi, cryptic species in North America and Europe. <em>Mycological progress</em> 14 (1): 1–8. https://doi.org/10.1007/s11557-014-1016-9</p>
<p>Liimatainen, K., Niskanen, T., Dima, B., Kytövuori, I., Ammirati, J.F. & Frøslev, T.G. (2014) The largest type study of Agaricales species to date: bringing identification and nomenclature of <em>Phlegmacium (Cortinarius)</em> into the DNA era. <em>Persoonia-Molecular Phylogeny and Evolution of Fungi </em>33 (1): 98–140. https://doi.org/10.3767/003158514X684681</p>
<p>Luo, Y. & Bau, T. (2021) <em>Cortinarius jiaoheensis</em> (Cortinariaceae), a new species of <em>Cortinarius </em>subgenus <em>Telamonia </em>section <em>Flexipedes,</em> from northeast China. <em>Phytotaxa </em>494 (1): 113–121. https://doi.org/10.11646/phytotaxa.494.1.7</p>
<p>Lanfear, R., Calcott, B., Ho, S.Y. & Guindon, S. (2012) Partition Finder: combined selection of partitioning schemes and substitution models for phylogenetic analyses. <em>Molecular biology and evolution</em> 29 (6): 1695–1701. https://doi.org/10.1093/molbev/mss020</p>
<p>Miller, M.A., Pfeiffer, W. & Schwartz, T. (2010) <em>Creating the CIPRES Science Gateway for inference of large phylogenetic trees. Proceedings of the Gateway Computing Environments Workshop</em> <em>(GCE)</em>. New Orleans, Louisiana. 1–8. https://doi.org/10.1109/GCE.2010.5676129</p>
<p>Naseer, A., Garrido-Benavent, I., Khan, J., Ballarà, J., Mahiques, R., Khalid, A.N. & Sher, H. (2020) <em>Cortinarius pakistanicus</em> and <em>C. pseudotorvus</em>: two new species in oak forests in the Pakistan Himalayas. <em>MycoKeys</em> 74: 91. https://doi.org/10.3897/mycokeys.74.49734</p>
<p>Niskanen, T. (2014) <em>Nomenclatural novelties</em>. Index Fungorum no. 186.</p>
<p>Niskanen, T., Kytövuori, I. & Liimatainen, K. (2011) <em>Cortinarius </em>sect. Armillati in northern Europe. <em>Mycologia </em>103 (5): 1080–1101. https://doi.org/10.3852/10-350</p>
<p>Niskanen, T., Liimatainen, K. & Kytövuori, I. (2015) Nomenclatural novelties 256. Index Fungorum, (256). Available from: http://www. indexfungorum.org/Publications/Index%20Fungorum%20no.256.pdf (accessed 20 November 2020)</p>
<p>Niskanen, T., Liimatainen, K., Kytövuori, I. & Lindström, H. (2012) Subgen. <em>Telamonia</em> (Fr.) Trog. <em>In</em>: Knudsen, H. & Vesterholt, J. (Eds.) <em>Funga Nordica</em>, 2nd revised edition. Agaricoid, boletoid, clavarioid, cyphelloid and gastroid genera. Nordsvamp, Denmark, pp. 826–885.</p>
<p>Nuske, S.J., Anslan, S., Tedersoo, L., Congdon, B.C. & Abell, S.E. (2019) Ectomycorrhizal fungal communities are dominated by mammalian dispersed truffle-like taxa in north-east Australian woodlands. <em>Mycorrhiza </em>29 (3): 181–193. https://doi.org/10.1007/s00572-019-00886-2</p>
<p>Peintner, U., Moncalvo, J.M. & Vilgalys, R. (2004) Toward a better understanding of the infrageneric relationships in <em>Cortinarius </em>(Agaricales, Basidiomycota). <em>Mycologia </em>96: 1042–1058. https://doi.org/10.1080/15572536.2005.11832904</p>
<p>Ronquist, F., Teslenko, M., Van Der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. <em>Systematic biology</em> 61 (3): 539–542. https://doi.org/10.1093/sysbio/sys029</p>
<p>Saba. M., Khalid, A.N., Jabeen, S. & Dima, B. (2017) <em>Cortinarius longistipitatus,</em> a new species in subgenus <em>Telamonia,</em> section <em>Cinnabarini,</em> from Pakistan. <em>Phytotaxa</em> 328 (3): 257–266. https://doi.org/10.11646/phytotaxa.328.3.4</p>
<p>Song, J., Liang, J.-F., Mehrabi-Koushki, M., Krisai-Greilhuber, I., Ali, B., Bhatt, V.K., Cerna-Mendoza, A., Chen, B., Chen, Z.-X., Chu, H.-L., Corazon-Guivin, M.A., da Silva, G.A., De Kesel, A., Dima, B., Dovana, F., Farokhinejad, R., Ferisin, G., Guerrero-Abad, J.C., Guo, T., Han, L.-H., Ilyas, S., Justo, A., Khalid, A.N., Khodadadi-Pourarpanahi, S., Li, T.-H., Liu, C., Lorenzini, M., Lu, J.-K., Mumtaz, A.S., Oehl, F., Pan, X.-Y., Papp, V., Qian, W., Razaq, A., Semwal, K.C., Tang, L.-Z., Tian, X.-L., Vallejos-Tapullima, A., van der Merwe, N.A., Wang, S.-K., Wang, C.-Q., Yang, R.-H., Yu, F., Zapparoli, G., Zhang, M., Antonín, V., Aptroot, A., Aslan, A., Banerjee, A., Chatterjee, S., Dirks, A.C., Ebrahimi, L., Fotouhifar, K.-B., Ghosta, Y., Kalinina, L.B., Karahan, D., Liu, J., Maiti, M.K., Mookherjee, A., Nath, P.S., Panja, B., Saha, J., Ševčíková, H., Voglmayr, H., Yazıcı, K. & Haelewaters, D. (2019) Fungal Systematics and Evolution: FUSE 5. <em>Sydowia </em>71: 141–245. https://doi.org/10.1093/sysbio/sys029</p>
<p>Stamatakis, A. (2014) RAxML version 8: a tool for phylogenetic analysis and postanalysis of large phylogenies. Bioinformatics. https://doi.org/10.1093/bioinformatics/btu033</p>
<p>Soop, K., Dima, B., Cooper, J.A., Park, D. & Oertel, B. (2019) A phylogenetic approach to a global supraspecifc taxonomy of <em>Cortinarius </em>(Agaricales) with an emphasis on the southern mycota. <em>Persoonia</em> 42: 261–290. https://doi.org/10.3767/persoonia.2019.42.10</p>
<p>Vellinga, E.C., Sysouphanthong, P. & Hyde, K.D. (2011) The family Agaricaceae: phylogenices and two new white spored genera. <em>Mycologia</em> 103 (3): 494–509. https://doi.org/10.3852/10-204</p>
<p>Vilgalys, R. & Hester, M. (1990) Rapid genetic identification and mapping of enzymatically amplified ribosomal DNA from several Cryptococcus species. <em>Journal of Bacteriology</em> 172 (8): 4238–4246. https://doi.org/10.1128/JB.172.8.4238-4246.1990</p>
<p>Teasdale, S.E., Beulke, A.K., Guy, P.L. & Orlovich, D.A. (2013) Environmental barcoding of the ectomycorrhizal fungal genus <em>Cortinarius.</em> <em>Fungal Diversity</em> 58: 299–310. https://doi.org/10.1007/s13225-012-0218-1</p>
<p>White, T.J, Bruns T, Lee, S.J. & Taylor, J. (1990) Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. <em>PCR protocols: a guide to methods and applications</em> 18 (1): 315–322. https://doi.org/10.1016/B978-0-12-372180-8.50042-1</p>