Abstract
Epidendrum claustralis, a new species of the Rupestre Group is described and illustrated. It is a novelty from the Departments of Junín, San Martín and Amazonas, Peru. It is compared to the most similar species, Epidendrum excelsum. The species is characterized by the entire lip and with two prominent parallel calli that lean towards each other after pollination or the loss of the pollinarium, closing the entrance of the nectary, and thus avoiding another pollination event.
References
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;"><span lang="en-GB">Acu</span>ña-Tarazona, M.E., Hágsater, E. & Santiago, E. (2019) Epidendrum choccei (Orchidaceae), a new species from Northern Peru. Phytotaxa 394: 98–104. https://doi.org/10.11646/phytotaxa.394.1.7</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ames, O., Hubbard, F.T. & Schweinfurth, C. (1934) Three Polymorphic Alliances in Epidendrum. Botanical Museum Leaflets 2: 41–69. https://doi.org/10.5962/p.168587</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ames, O., Hubbard, F.T. & Schweinfurth, C. (1936) The Genus Epidendrum in the United States and Middle America. Botanical Museum, Cambridge, 233 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Collantes, B. (2014) Huancavelica: Amaru, paraíso de orquídeas. Gobierno Regional de Huancavelica, Lima, 338 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Collantes, B. & Hágsater, E. (2015) Epidendrum ochoae. In: Hágsater, E. & Sánchez S., L. (Eds.) The Genus Epidendrum, Part 11, Icones Orchidacearum 15: pl. 1546.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Decaisne, J. & Planchón, J.E. (1854) Esquisse d‘une monographe des Araliacées. Revue Horticole, Journal d’horticuluture pratique 4: 104–109. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gentry, A. (1993) Overview of the Peruvian Flora. In: Brako, L. & Zarucchi, J. (Eds.) Catalogue of the Flowering plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45: 29–40. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">GFW (2022) Global Forest World, Washington, USA. Available from: https://www.globalforestwatch.org/ (accessed 1 Sept 2022). </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Goicochea, R.A., Gutiérrez, R.A.D., Ruiz, A.R. & Salas, M.G. (2019) Orquídeas del Perú: Relación de especies y sus sinónimos. Corporación G y G E.I.R.L, Moyobamba, 285 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. (2004) Epidendrum pachydiscum. In: Hágsater, E. & Sánchez, L. (Eds.) The Genus Epidendrum, Part. 4, Icones Orchidacearum 7: pl. 773.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. & Soto, M.A. (2005a) Epidendrum. In: Pridgeon, A.M., Cribb, P.J., Chase, M.W. & Rasmussen, F.N. (Eds.) Genera Orchidacearum 4. Epidendroideae, Part. 1. Oxford University Press, Oxford, pp. 236–251.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. (2005b) Epidendrum pachydiscum. In: Hagsater, E. & Salazar, G.A. (Eds.) The Genus Epidendrum, Part 3, Icones Orchidacearum 4: pl. 773.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. (2007) Epidendrum megalocoleum. In: Hágsater, E. & Sánchez, L. (Eds.) The Genus Epidendrum, Part. 6, Icones Orchidacearum 9: pl. 956.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. (2016) Epidendrum aida-alvarezii. In: Hágsater, E. & Sánchez, L. (Eds.) The Genus Epidendrum, Part 11. Icones Orchidacearum 15: pl. 1570.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E., Navarro, W.C.R. & Quispe-Melgar, H.R. (2020) Epidendrum curimarcense. In: Hágsater, E. & Santiago, E. (eds.) The Genus Epidendrum, Part 13, Icones Orchidacearum 17: pl. 1766.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E., Edquén, J.D. & Santiago, E. (2021) Epidendrum concavitridentes. In: Hágsater, E. & Santiago, E. (Eds.) The Genus Epidendrum, Part 14, Icones Orchidacearum 18: pl. 1862.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. & Santiago, E. (in press) Epidendrum rupestre. In: Hágsater, E. & Santiago, E. (eds.) The Genus Epidendrum, Part 16, Icones Orchidacearum.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Harms, H.A. (1931) Oreopanax oroyanus. In: Bornmüller, J. (Ed.) Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem 11: 146–149. https://doi.org/10.2307/3994596</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Jacquin, N. (1760) Epidendrum difforme. In: Enumeratio Systematica Plantarum. Christiani de Lavnoy, Leiden. 29 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lindley, J. (1835) Cavendishia. Edwards’s Botanical Register 21: t. 1791. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lindley, J. (1841) Notes upon the Genus Epidendrum. Hooker’s Journal of Botany 3: 81–89.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Linnaeus, C. (1753) Species Plantarum. Ray Society’s Facsimile, London, 395 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Linnaeus, C. (1759) Systema naturae per regna tria naturae: secundum clases, ordines, géneros, especies, cum characteribus, differentiis, synomiis, locis. Tomusi, London, 1367 pp. https://doi.org/10.5962/bhl.title.542</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Linnaeus f., C. (1782) Supplementum Plantarum Systematis Vegetabilium. Ed. 2. Impensis Orphanotrophei, Brunsvigae, pp. 1–468. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Mendoza, W. (2005) Especie nueva de Polylepis (Rosaceae) de la cordillera Vilcabamba (Cusco, Peru). Revista Peruana de Biología 12: 103–106. https://doi.org/10.15381/rpb.v12i1.2364</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Navarro-Romo, W.C., Quispe-Melgar, H.R. & Hágsater, E. (2020) Epidendrum curimarcense (Orchidaceae), a new species from Central Peru. Lankesteriana 20: 7–13. https://doi.org/ 10.15517/lank.v20i1.40781 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pessoa, E.M, Cordeiro, J.M.P, Felix, L.P, Almeida, E.M, Costa, L., Nepomuceno, A., Souza, G., Chase, M.W, Alves, M. & Van Den Berg, C. (2021) Too many species: morphometrics, molecular phylogenetics and genome structure of a Brazilian species complex in Epidendrum (Laeliinae; Orchidaceae) reveal fewer species than previously thought. Botanical Journal of the Linnean Society 195: 161–188. https://doi.org/10.1093/botlinnean/boaa071</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Quispe-Melgar, H.R., Llacua-Tineo, Y.S., Ames-Martínez, F.N., David Huayta, D., Lagones Poma, K.L. & Hágsater, E. (2022) Epidendrum alejandrinae (Orchidaceae), a new species from the high Andean forests of central Peru. Phytotaxa 541: 217–224. https://doi.org/10.11646/phytotaxa.541.3.1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">R Core Team. (2022) R: A Language and Environment for Statistical Computing. R Foundation for Statistical Computing, Vienna. 2630 pp. Available from: https://www.R-project.org/ (accessed 1 September 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">RStudio. (2022) RStudio: Integrated Development Environment for R. R Studio, Boston. Available from: http://www.rstudio.com/ (accessed 1 September 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ruiz López, H. & Pavón, J.A. (1794) Florae Peruvianae, et Chilensis Prodromus, sive novorum generum plantrum peruvianum, et chilensium descriptiones et icones. Typographio Paleariniano, Madrid, pp. 1–153.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sánchez Saldaña, L.M. & Hágsater, E. (1998) Taxonomic Study of Epidendrum difforme Group (Orchidaceae). In: Raynal-Roques, A., Roguenant, A. & Prat, D. (Eds.) Proceedings of the 15th World Orchid Conference, Rio de Janeiro 1996. Naturalia Publications, Turriers, pp. 235–244.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schlechter, F.R. (1921) Die Orchideenfloren der Suedamerikanischen Kordillerenstaaten III. Ecuador. Repertorium Specierum Novarum Regni Vegetabilis. Beihefte 8: 1–172. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schlechter, F.R. (1917) XII Orchidaceae nova et criticae. Decas XLIX-L. Repertorium Specierum Novarum Regni Vegetabilis. Centralblatt für Sammlung und Veroffentlichung von Einzeldiagnosen neuer Pflanzen 15: 50–59. https://doi.org/10.1002/fedr.4870150402</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schweinfurth, C. (1970) First supplement to the Orchids of Peru. Fieldiana, Botany 33: 1–80. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Thiers, B. (2021) Index Herbariorum: a global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 29 August 2021).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Williams, L.O. (1940) Orchid Studies XIII, New or interesting orchids from Ecuador and Colombia. Botanical Museum Leaflets 9: 1–25. https://doi.org/10.5962/p.295149</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ames, O., Hubbard, F.T. & Schweinfurth, C. (1934) Three Polymorphic Alliances in Epidendrum. Botanical Museum Leaflets 2: 41–69. https://doi.org/10.5962/p.168587</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ames, O., Hubbard, F.T. & Schweinfurth, C. (1936) The Genus Epidendrum in the United States and Middle America. Botanical Museum, Cambridge, 233 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Collantes, B. (2014) Huancavelica: Amaru, paraíso de orquídeas. Gobierno Regional de Huancavelica, Lima, 338 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Collantes, B. & Hágsater, E. (2015) Epidendrum ochoae. In: Hágsater, E. & Sánchez S., L. (Eds.) The Genus Epidendrum, Part 11, Icones Orchidacearum 15: pl. 1546.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Decaisne, J. & Planchón, J.E. (1854) Esquisse d‘une monographe des Araliacées. Revue Horticole, Journal d’horticuluture pratique 4: 104–109. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gentry, A. (1993) Overview of the Peruvian Flora. In: Brako, L. & Zarucchi, J. (Eds.) Catalogue of the Flowering plants and Gymnosperms of Peru. Monographs in Systematic Botany from the Missouri Botanical Garden 45: 29–40. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">GFW (2022) Global Forest World, Washington, USA. Available from: https://www.globalforestwatch.org/ (accessed 1 Sept 2022). </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Goicochea, R.A., Gutiérrez, R.A.D., Ruiz, A.R. & Salas, M.G. (2019) Orquídeas del Perú: Relación de especies y sus sinónimos. Corporación G y G E.I.R.L, Moyobamba, 285 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. (2004) Epidendrum pachydiscum. In: Hágsater, E. & Sánchez, L. (Eds.) The Genus Epidendrum, Part. 4, Icones Orchidacearum 7: pl. 773.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. & Soto, M.A. (2005a) Epidendrum. In: Pridgeon, A.M., Cribb, P.J., Chase, M.W. & Rasmussen, F.N. (Eds.) Genera Orchidacearum 4. Epidendroideae, Part. 1. Oxford University Press, Oxford, pp. 236–251.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. (2005b) Epidendrum pachydiscum. In: Hagsater, E. & Salazar, G.A. (Eds.) The Genus Epidendrum, Part 3, Icones Orchidacearum 4: pl. 773.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. (2007) Epidendrum megalocoleum. In: Hágsater, E. & Sánchez, L. (Eds.) The Genus Epidendrum, Part. 6, Icones Orchidacearum 9: pl. 956.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. (2016) Epidendrum aida-alvarezii. In: Hágsater, E. & Sánchez, L. (Eds.) The Genus Epidendrum, Part 11. Icones Orchidacearum 15: pl. 1570.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E., Navarro, W.C.R. & Quispe-Melgar, H.R. (2020) Epidendrum curimarcense. In: Hágsater, E. & Santiago, E. (eds.) The Genus Epidendrum, Part 13, Icones Orchidacearum 17: pl. 1766.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E., Edquén, J.D. & Santiago, E. (2021) Epidendrum concavitridentes. In: Hágsater, E. & Santiago, E. (Eds.) The Genus Epidendrum, Part 14, Icones Orchidacearum 18: pl. 1862.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hágsater, E. & Santiago, E. (in press) Epidendrum rupestre. In: Hágsater, E. & Santiago, E. (eds.) The Genus Epidendrum, Part 16, Icones Orchidacearum.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Harms, H.A. (1931) Oreopanax oroyanus. In: Bornmüller, J. (Ed.) Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem 11: 146–149. https://doi.org/10.2307/3994596</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Jacquin, N. (1760) Epidendrum difforme. In: Enumeratio Systematica Plantarum. Christiani de Lavnoy, Leiden. 29 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lindley, J. (1835) Cavendishia. Edwards’s Botanical Register 21: t. 1791. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lindley, J. (1841) Notes upon the Genus Epidendrum. Hooker’s Journal of Botany 3: 81–89.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Linnaeus, C. (1753) Species Plantarum. Ray Society’s Facsimile, London, 395 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Linnaeus, C. (1759) Systema naturae per regna tria naturae: secundum clases, ordines, géneros, especies, cum characteribus, differentiis, synomiis, locis. Tomusi, London, 1367 pp. https://doi.org/10.5962/bhl.title.542</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Linnaeus f., C. (1782) Supplementum Plantarum Systematis Vegetabilium. Ed. 2. Impensis Orphanotrophei, Brunsvigae, pp. 1–468. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Mendoza, W. (2005) Especie nueva de Polylepis (Rosaceae) de la cordillera Vilcabamba (Cusco, Peru). Revista Peruana de Biología 12: 103–106. https://doi.org/10.15381/rpb.v12i1.2364</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Navarro-Romo, W.C., Quispe-Melgar, H.R. & Hágsater, E. (2020) Epidendrum curimarcense (Orchidaceae), a new species from Central Peru. Lankesteriana 20: 7–13. https://doi.org/ 10.15517/lank.v20i1.40781 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pessoa, E.M, Cordeiro, J.M.P, Felix, L.P, Almeida, E.M, Costa, L., Nepomuceno, A., Souza, G., Chase, M.W, Alves, M. & Van Den Berg, C. (2021) Too many species: morphometrics, molecular phylogenetics and genome structure of a Brazilian species complex in Epidendrum (Laeliinae; Orchidaceae) reveal fewer species than previously thought. Botanical Journal of the Linnean Society 195: 161–188. https://doi.org/10.1093/botlinnean/boaa071</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Quispe-Melgar, H.R., Llacua-Tineo, Y.S., Ames-Martínez, F.N., David Huayta, D., Lagones Poma, K.L. & Hágsater, E. (2022) Epidendrum alejandrinae (Orchidaceae), a new species from the high Andean forests of central Peru. Phytotaxa 541: 217–224. https://doi.org/10.11646/phytotaxa.541.3.1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">R Core Team. (2022) R: A Language and Environment for Statistical Computing. R Foundation for Statistical Computing, Vienna. 2630 pp. Available from: https://www.R-project.org/ (accessed 1 September 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">RStudio. (2022) RStudio: Integrated Development Environment for R. R Studio, Boston. Available from: http://www.rstudio.com/ (accessed 1 September 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ruiz López, H. & Pavón, J.A. (1794) Florae Peruvianae, et Chilensis Prodromus, sive novorum generum plantrum peruvianum, et chilensium descriptiones et icones. Typographio Paleariniano, Madrid, pp. 1–153.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sánchez Saldaña, L.M. & Hágsater, E. (1998) Taxonomic Study of Epidendrum difforme Group (Orchidaceae). In: Raynal-Roques, A., Roguenant, A. & Prat, D. (Eds.) Proceedings of the 15th World Orchid Conference, Rio de Janeiro 1996. Naturalia Publications, Turriers, pp. 235–244.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schlechter, F.R. (1921) Die Orchideenfloren der Suedamerikanischen Kordillerenstaaten III. Ecuador. Repertorium Specierum Novarum Regni Vegetabilis. Beihefte 8: 1–172. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schlechter, F.R. (1917) XII Orchidaceae nova et criticae. Decas XLIX-L. Repertorium Specierum Novarum Regni Vegetabilis. Centralblatt für Sammlung und Veroffentlichung von Einzeldiagnosen neuer Pflanzen 15: 50–59. https://doi.org/10.1002/fedr.4870150402</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Schweinfurth, C. (1970) First supplement to the Orchids of Peru. Fieldiana, Botany 33: 1–80. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Thiers, B. (2021) Index Herbariorum: a global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 29 August 2021).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Williams, L.O. (1940) Orchid Studies XIII, New or interesting orchids from Ecuador and Colombia. Botanical Museum Leaflets 9: 1–25. https://doi.org/10.5962/p.295149</span></span></span>
