A checklist of Iranian Grasses

MAHMOUD BIDARLORD; FARROKH GHAHREMANINEJAD

doi:10.11646/phytotaxa.574.1.1

Issue: Vol. 574 No. 1: 25 November 2022

Type: Article

Published: 2022-11-25

DOI: 10.11646/phytotaxa.574.1.1

Page range: 1-31

Abstract views: 579

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A checklist of Iranian Grasses

MAHMOUD BIDARLORD⁺⁻
FARROKH GHAHREMANINEJAD⁺⁻

Department of Forests and Rangelands Research, GilanAgricultural and Natural Resources Research and Education Center, Agricultural Research, Education and Extension Organization (AREEO), Rasht, Iran

Department of Plant Sciences, Faculty of Biological Sciences, Kharazmi University; Postal Code 15719-14911, Tehran, Iran

Flora Iranica Persia southwest Asia true grasses Poaceae Gramineae Monocots

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Abstract

This paper provides a taxonomic treatment for the large angiosperm family, the Poaceae, in Iran. The last comprehensive review of Iranian Grasses (Poaceae) dates back to half a century ago in Flora Iranica. Since then, a number of taxa have been described as new species or newly recorded from Iran, and some names have been placed in synonym. This study presents a comprehensive and updated account of all reported Iranian grasses and their classification. The results include 128 genera, 477 species, 64 subspecies and 44 varieties belonging to 37 subtribes, 19 tribes, and seven subfamilies, of which 28 taxa are endemic for Iran. The genera, Elymus, with 7 species and Festuca, with 5 species have the highest number of endemic species. The most diverse genera in Iran are, Bromus (39 spp.), Festuca (27 spp.), Stipa (27 spp.), and Poa (26 spp.). Poeae is the largest tribe, and Pooideae is the largest subfamily in Iran. This paper provides a new name, Festuca neopersica a new combination, Leymus europaeus as well as nomenclatural modifications on the basis of recent taxonomic and molecular investigation of Poaceae taxa related to Iran. The comparison of the species distribution in Iran with that in neighboring countries indicates that Turkey has the most and the Arabian Peninsula the least species in common with Iran, with 305 (64%) and 156 (32%) species, respectively. The species shared with Iran, Pakistan and Saudi Arabia are placed mainly in PACMAD clade, while the species shared with Iran, Turkey and Transcaucasia are mainly placed in PEB clade. Grasslands in Iran are in need of more protection as a result of climatic change and human actions.

References

Abbasi, M. & Assadi, M. (2014) Evaluation of generic concept in Colpodium Trin. and Catabrosella (Tzvel) Tzvel: a new subspecies in Poaceae family in Iran. International Journal of Biosciences 5: 50–57. https://doi.org/10.12692/ijb/5.7.50-57

Abbasi, M., Assadi, M., Nejadsattari, T. & Attar, F. (2010) Micromorphological and anatomical featuers of Puccinellia dolicholepis (Poaceae), a new record for the flora of Iran. The Iranian Journal of Botany 16: 267–272. 

Aghaei, M.J., Mozafari, J., Taleei, A.R., Naghavi, M.R. & Omid, M. (2008) Distribution and diversity of Aegilops tauschii in Iran. Genetic Resources and Crop Evolution 55: 341–349. https://doi.org/10.1007/s10722-007-9239-0

Akhani, H. & Joharchi, M.R. (1995) New and interesting records for the flora of Iran. Willdenowia 25: 261–266.

Akhani, H. & Scholz, H. (1998) Studies on the flora and vegetation of the Golestan National Park, NE Iran II: A new Poa and some new and noteworthy grass records for Iran. Edinburgh Journal of Botany 55: 443–453. https://doi.org/10.1017/S0960428600003280

Akhani, H. & Deil, U. (2012) First observations on the flora and vegetation of three islands in the NW Persian Gulf (Iran). Phyton (Horn) 52: 73–99.

Alexeev, E.B. (1971) Byulleten’ Moskovskogo Obshchestva Ispytatelei Prirody. Otdel Biologicheskii 76: 119.

Alexeev, E.B. (1977) Bjulleten Moskovskogo Oba?estva Ispytatelej Prirody. Otdel Biologi?eskij 82: 101.

Alexeev, E.B. (1979) Genus Festuca L. Florae Iranicae et territoriorum confinium. Novosti Sistematiki Vysshikh Rastenii 16: 7–35.

Amini Rad, M. (2015) A study on the genus Digitaria (Poaceae) in Iran. Taxonomy and Biosystematics 7: 39–50.

Amini Rad, M. & Scholz, H. (2011) Urochloa panicoides and a subspecies of Eragrostis cilianensis, new to the flora of Iran. Rostaniha 12: 195–198.

Amini Rad, M., Sonboli, A., Ghasemnezhad, A. & Hadian, J. (2012) Hierrochloa odorata (Poaceae), a new report from Iran. The Iranian Journal of Botany 18: 224–225.

Amiri, N. (2016) Molecular phylogeny of Poa L. sensu lato (Poaceae) with a focus on West Asian species. M.Sc. Thesis. University of Ottawa.

Asri, Y. & Rabie, M. (2010) Taxonomical and phytogeographical studies on Centropodia forskalii subsp. forskalii (Poaceae), a new record for the flora of Iran. The Iranian Journal of Botany 16: 229–232.

Assadi, M. (1983) Plant from N. Azarbayejan, new to Iran. The Iranian Journal of Botany 2: 1–8. 

Assadi, M. (1984a) New species and new plant records from Iran. The Iranian Journal of Botany 2: 83–94. 

Assadi, M. (1984b) Study on the autumn plant of Kavir. The Iranian Journal of Botany 2: 125–148.

Assadi, M. (chief editor) (1987–2021) Flora of Iran, vols. 1–151. Research Institute of Forests and Rangelands of Iran, Tehran.

 Assadi, M. (1988) Plant of Arasbaran Protected Area, NW, Iran (Part II). The Iranian Journal of Botany 4: 1–59.

Assadi, M. (1994) Crossing experiments in Elymus transhyrcanus group, a new subspecies and a new species. The Iranian Journal of Botany 6: 185–195.

Assadi, M. (1996a) A new species of Ephedra L. and reports of two new or interesting grasses from Iran. The Iranian Journal of Botany 7: 1–6.

Assadi, M. (1996b) A taxonomic revision of Elymus sect. Caespitosae and sect. Elytrigia (Poaceae, Triticeae) in Iran. Willdenowia 26: 251–271. https://doi.org/10.3372/wi.26.2612

Assadi, M. (1998a) Biosystematic studies of the Elymus hispidus (Poaceae: Triticeae) group in Iran. Nordic Journal of Botany 18: 483–491. https://doi.org/10.1111/j.1756-1051.1998.tb01526.x

Assadi, M. (1998b) Noteworthy plant records for the flora of Iran. The Iranian Journal of Botany 7: 217–220. 

Assadi, M. (2017) New species, new combination and an identification key of the genus Elymus (Poaceae: Triticeae) in Iran. The Iranian Journal of Botany 23: 77–83. 

Assadi, M. & Runemark, H. (1983) Note on the flora and vegetation of S. Baluchistan, Iran. The Iranian Journal of Botany 2: 69–78.

Assadi, M. & Runemark, H. (1995) Hybridisation, genomic constitution and generic delimitation in Elymus s.l. (Poaceae: Triticeae). Plant Systematics and Evolution 194: 189–205. https://doi.org/10.1007/BF00982855

Assadi, M. & Wendelbo, P. (1977) New and interesting plant records from NW Iran. The Iranian Journal of Botany 1: 97–108. 

Baden, C. (1991) A taxonomic revision of Psathyrostachys (Poaceae). Nordic Journal of Botany 2: 3–26. https://doi.org/10.1111/j.1756-1051.1991.tb01790.x

Barberá, P., Romero-Zarco, C. & Aedo, C. (2017) Taxonomic revision of Trisetum section Sibirica (Poaceae: Pooideae: Aveninae). Systematic Botany 42 (4): 782–806. https://doi.org/10.1600/036364417X696564

Baum, B.R., Fleischmann, G., Martens, J.W., Rajhathy, T. & Thomas, H. (1972) Notes on the habitat and distribution of Avena species in the Mediterranean and Middle East. Canadian Journal of Botany 50: 1385–1397. https://doi.org/10.1139/b72-167

Behmanesh, B., Abedi Sarvestani, A., Sharafatmandrad, M., Shahraki, M.R. & Hajili-Davaji, A. (2016) Assessment of rangeland degradation indicators using exploiters’ view between authorized and unauthorized exploiters (Case study: Saryqmish Winter Rangelands, Golestan Province, Iran). Desert 21 (2): 105–113.

Behroozian, M., Ejtehadi, H., Memariani, F., Joharchi, M.R. & Mesdaghi, M. (2019) Stipa richteriana (Poaceae) and Galium songaricum (Rubiaceae): two new additions of the Central Asian species to the flora of Iran. Nova Biologica Reperta 6 (3): 326–333. https://doi.org/10.29252/nbr.6.3.326

Blaner, A., Schneider, J. & Roser, M. (2014) Phylogenetic relationships in the grass family (Poaceae) based on the nuclear single copy locus topoisomerase 6 compared with chloroplast DNA. Systematics and Biodiversity 12: 111–124. https://doi.org/10.1080/14772000.2014.890137

Blattner, F.R. (2009) Progress in phylogenetic analysis and a new infrageneric classification of the barley genus Hordeum (Poaceae: Triticeae). Breeding Science 59: 471–480. https://doi.org/10.1270/jsbbs.59.471

Boissier, E. (1846) Diagnoses Plantarum Orientalium Novarum, Ser. 1: 129.

Bor, N.L. (1968) Gramineae. In: Townsend, C.C., Guest, E. & Al-Rawi, A. (Eds.) Flora of Iraq, vol. 9. Baghdad, Iraq Ministry of Agriculture, 588 pp.

Bor, N.L. (1970) Gramineae. In: Rechinger, K.H. (Ed.) Flora Iranica no. 70. Akademische Druck- u. Verlagsanstalt, Graz-Austria, 573 pp.

Borchers, E., Frey, W. & Hilger, H. (1982) C4 species in the highlands of Iran and Afghanistan. Notes from the Royal Botanic Garden, Edinburgh 40: 99–113. 

Bordbar, F. & Rahiminejad, M.R. (2010) A taxonomic revision of Aegilops crassa Boiss. (Poaceae) in Iran. The Iranian Journal of Botany 16: 30–35.

Bothmer, R. von, Flink, J., Jacobsen, N. & Jorgensen, R.B. (1989) Variation and differentiation in Hordeum marinum (Poaceae). Nordic Journal of Botany 9: 1–10. https://doi.org/10.1111/j.1756-1051.1989.tb00975.x

Bothmer, R. von, Flink, J., Jacobsen, N., Baden, C., Jorgensen, R.B. & Linde-Laursen, I.B. (1991) An ecogeographical study of the genus Hordeum, IBPGR (International Board for Plant Genetic Resources), Rome, pp. 71–75.

Bothmer, R.V. (1996) Distribution and habitat preferences in the genus Hordeum in Iran and Turkey. Annalen des Naturhistorischen Museums in Wien 98 B Suppl.: 107–116.

Boulos, L. (2005) Flora of Egypt vol. 4. Al Hadara Publishing, Cairo. pp. 1–617.

Bouchenak-Khelladi, Y., Verboom, G.A., Savolainen, V. & Hodkinson, T.R. (2010) Biogeography of the grasses (Poaceae): a phylogenetic approach to reveal evolutionary history in geographical space and geological time. Botanical Journal of the Linnean Society 162: 543–557. https://doi.org/10.1111/j.1095-8339.2010.01041.x

Brummitt, R.K. & Powell, C.E. (1992) Authors of Plant Names. A list of authors of scientific names of Plants, with recommended standard forms of their names, including abbreviations. Royal Botanic Gardens, Kew, pp. 732. 

Brummitt, R.K. (comp.) (1992) Vascular plant families and genera. Royal Botanic Gardens, Kew, 804 pp.

Cabi, E., Soreng, R.J., Çýngay, B. & Karabacak, E. (2015) A new species of Bellardiochloa, B. doganiana (Poaceae), from the Taurus Mountains of Turkey. Phytotaxa 205: 123–128. https://doi.org/10.11646/phytotaxa.205.2.6

Catalán, P., López-Álvarez, D., Bellosta, C. & Villar, L. (2016) Updated taxonomic descriptions, iconography, and habitat preferences of Brachypodium distachyon, B. stacei, and B. hybridum (Poaceae). Anales Jardin Botánico Madrid 73 (1): 1–14. https://doi.org/10.3989/ajbm.2428

Catalán, P., Torrecilla, P., López-Rodríguez, J.A., Müller, J.& Stace, C.A. (2007) A systematic approach to subtribe Loliinae (Poaceae: Pooideae) based on phylogenetic evidence. Journal of Systematic and Evolutionary 23: 380–405. https://doi.org/10.5642/aliso.20072301.31

Catalán, P., Torrecilla, P., Rodrýguez, J.A.L. & Olmstead, R.G. (2004) Phylogeny of the festucoid grasses of subtribe Loliinae and allies (Poeae, Pooideae) inferred from ITS and trnL–F sequences. Molecular phylogenetics and evolution 31: 517–541. https://doi.org/10.1016/j.ympev.2003.08.025

Chemisquy, M.A., Giussani, L.M., Scataglini, M.A., Kellogg, E.A.& Morrone, O. (2010) Phylogenetic studies favour the unification of Pennisetum, Cenchrus and Odontelytrum (Poaceae): a combined nuclear, plastid and morphological analysis, and nomenclatural combinations in Cenchrus. Annals of Botany 106: 107–30. https://doi.org/10.1093/aob/mcq090

Choo, M.K., Soreng, R.J. & Davis, J. (1994) Phylogenetic relationships among Puccinellia and allied genera of Poaceae as inferred from chloroplast DNA restriction site variation. American Journal of Botany 81: 119–126. https://doi.org/10.1002/j.1537-2197.1994.tb15417.x

Clayton, W.D, Govaerts, R., Harman, K.T., Williamson, H. & Vorontsova, M. (2020) World Checklist of Poaceae (Gaminae). Facilitated by the Royal Botanic Gardens, Kew.Available from: http://wcsp.science.kew.org/ (Retrieved 10 February 2021)

Clayton, W.D. & Snow, N. (2010) A key to Pacific Grasses. Kew Publishing, Royal Botanic Gardens, Kew. pp. 1–107.

Clayton, W.D. & Renvoize, S.A. (1986) Genera Graminum: Grasses of the World. Kew Bulletin Additional Series 13, Royal Botanical Garden, Kew. Her Majesty’s Stationery Office, London. pp. 389 

Clayton, W.D., Vorontsova, M.S., Harman, K.T. & Williamson, H. (2006 onwards) GrassBase – The Online World Grass Flora. Available from: http://www.kew.org/data/grasses-db.html (accessed 8 February 2021).

Cope, T.A. (1983) Centropodia: an earlier name for Asthenatherum (Gramineae). Kew Bulletin 37: 657–658. https://doi.org/10.2307/4109739

Darbyshire, S.J. (1993) Realignment of Festuca subgenus Schedonorus with the genus Lolium (Poaceae). Novon 3: 239–243. https://doi.org/10.2307/3391460

Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (Eds.) (2015) The plants of Sudan and South Sudan. Kew Publishing, Royal Botanic Gardens, Kew, 400 pp. 

Davis, P.H. (ed.) (1985) Flora of Turkey and the East Aegean Islands 9. Edinburgh University Press, Edinburgh. pp. 1–724.

Dewey, D.R. & Asay, K.H. (1975) The crested wheatgrasses of Iran. Crop Science 15: 844–849. https://doi.org/10.2135/cropsci1975.0011183X001500060028x

Dinarvand, M. & Howizeh, H. (2017) A new record of the genus Urochondra (Poaceae) from Iran. The Iranian Journal of Botany 23: 88–90.

Díaz-Pérez, A.J., Sharifi-Tehrani, M., Inda, L.A. & Catalán, P. (2014) Polyphyly, gene-duplication and extensive allopolyploidy framed the evolution of the ephemeral Vulpia grasses and other fine-leaved Loliinae (Poaceae). Molecular Phylogenetics and Evolution 79: 92–105. https://doi.org/10.1016/j.ympev.2014.06.009

Dobignard, D. & Chatelain, C. (2010) Index synonymique de la flore d’Afrique du nord 1. Éditions des Conservatoire et Jardin Botaniques, Genève. pp. 1–455.

Dobignard, A. & Chatelain, C. (2013) Index synonymique de la flore d’Afrique du nord 5. Éditions des Conservatoire et Jardin Botaniques, Genève. pp. 1–451.

Doðan, M. (1999) A concise taxonomic revision of the genus Alopecurus L. (Gramineae). Turkish Journal of Botany 23: 245–262.

Duvall, M.R., Davis, J.I., Clark, L.G., Noll, J.D., Goldman, D.H. & Sanchez-Ken, J.G. (2007) Phylogeny of the grasses (Poaceae) revisited. Aliso: A Journal of Systematic and Evolutionary Botany 23: 237–247. https://doi.org/10.5642/aliso.20072301.18

Eftekhari, T. & Ramezani, M. (2004) Introduction to plant biodiversity in Iran. In: Pushpangadan, P., Nair, K.N.& Ahmad, M.R. (eds.) Biodiversity and medicinal plantwealth of South Asian countries. National Botanical Research Institute, Lucknow, India. pp. 39–40.

Eftekhari, Z., Filehkesh, E., Bakhshi, Gh. & Assadi, M. (2005) Leymus tianschanicus (Poaceae), a new name for the flora of Iran. The Iranian Journal of Botany 11: 85–87.

Euro+Med (2006 onwards) Euro+Med PlantBase - the information resource for Euro-Mediterranean plant diversity. Available from: http://ww2.bgbm.org/EuroPlusMed/ (accessed 21 February 2017)

Fakhre Tabatabaei, S.M. & Ramak Maassoumi, T. (2001) Triticum boeoticum ssp thaoudar “exists” in Iran!. Cereal Research Communications 29 (1): 121–126. https://doi.org/10.1007/BF03543651

FAO. (2005) Fertilizer use by crop in the Islamic Republic of Iran; Land and Plant Nutrition Management Service Land and Water Development Division. Published by FAO, Rome.

Faramarzi, A., Saeidi, H. & Rahiminejad, M. (2012) A taxonomic revision of the genus Vulpia C.C.Gmel. (Poaceae, Poeae) in Iran. The Iranian Journal of Botany 18: 199–206.

Finot, V.L., Marticorena, A., Rodríguez, R. & Muñoz, R.G. (2017) Endemic species of the family Poaceae in Chile: Taxonomy, distribution, and conservation. In: Almusaed, A. & Al-Samaraee, S.M.S. (Eds.) Grasses-Benefits, diversities and functional roles. In Tech, pp. 63–86. https://doi.org/10.5772/intechopen.68973

Foggi, B., Signorini, M.A., Ardenghi, N.M.G., Pignotti, L. & Vitek, E. (2017) Types of Festuca names (Poaceae) in the herbarium W: taxa described by Johann Vetter (1865?1945). Webbia 72: 181–195. https://doi.org/10.1080/00837792.2017.1318632

Foggi, B. & Müller, J. (2009) Festuca. In: Valdés, B. & Scholz, H. (Eds.) with contributions from Raab-Straube, E. von & Parolly, G. Poaceae. Euro+Med Plantbase—the information resource for Euro-Mediterranean plant diversity. Available from: http://ww2.bgbm.org/EuroPlusMed/ (accessed February 2019).

Frederiksen, S. (1991) Taxonomic studies in Eremopyrum (Poaceae). Nordic Journal of Botany 2: 271–285. https://doi.org/10.1111/j.1756-1051.1991.tb01405.x

Frederiksen, S. (1986) Revision of Taeniatherum (Poaceae). Nordic Journal of Botany 6: 389–397. https://doi.org/10.1111/j.1756-1051.1986.tb00894.x

Fredriksen, S. & Petersen, G. (1997) Morphometrical analyses of Secale (Triticeae, Poaceae). Nordic Journal of Botany 17: 185–198. https://doi.org/10.1111/j.1756-1051.1997.tb00309.x

Freitag, H. (1975) The genus Piptatherum (Gramineae) in Southwest Asia. Notes from the Royal Botanic Garden, Edinburgh 33: 341–408.

Freitag, H. (1985) The genus Stipa (Gramineae) in southwest and south Asia. Notes from the Royal Botanical Garden Edinburgh 42: 355–489.

Farouji, A.E., Khodayari, H., Saeidi, H. & Rahiminejad, M.R. (2015) Genetic diversity of diploid Triticum species in Iran assessed using inter-retroelement amplified polymorphisms (IRAP) markers. Biologia 70: 52–60. https://doi.org/10.1515/biolog-2015-0002

Ghahreman, A., Alemi, M., Attar, F., Hamzeh’ee, B. & Columbus, J.T. (2006) Anatomical studies in some species of Bromus L. (Poaceae) in Iran. The Iranian Journal of Botany 12: 1–14.

Ghahremaninejad, F. & Nejad Falatoury, A. (2016) An update on the flora of Iran: Iranian angiosperm orders and families in accordance with APG IV. Nova Biologica Reperta 3: 80–107. https://doi.org/10.21859/acadpub.nbr.3.1.80

Ghahremaninejad, F. & Baldini, R.M. (2009) The correct author for Stipagrostis karelinii (Gramineae). Annalen des Naturhistorischen Museums in Wien 110B: 264. 

Ghahremaninejad, F., Hoseini, E. & Fereidounfar, S. (2020) Cities in drylands as artificial protected areas for plants. Biodiversity and Conservation 30: 243–248. https://doi.org/10.1007/s10531-020-02079-2

Ghahremaninejad, F., Hoseini, E. & Jalali, S. (2021) The cultivation and domestication of wheat and barley in Iran, brief review of a long history. The Botanical Review 87 (1): 1–22. https://doi.org/10.1007/s12229-020-09244-w

Ghasemkhani, M., Akhani, H., Saebi, J. & Scholz, H. (2008) The genera Aristida and Stipagrostis (Poaceae)in Iran. Willdenowia 38: 135–148. https://doi.org/10.3372/wi.38.38108

Gibson, D.J. (2009) Grasses and grassland ecology. Oxford University Press, New York, 305 pp.

Gillespie, L.J., Soreng, R.J., Cabi, E. & Amiri, N. (2018) Phylogeny and taxonomic synopsis of Poa subgenus Pseudopoa (including Eremopoa and Lindbergella) (Poaceae, Poeae, Poinae). PhytoKeys 111: 69. https://doi.org/10.3897/phytokeys.111.28081

Givnish, T.J., Ames, M., Mcneal, J.R., Mckain, M.R., Steele, P.R., Depamphilis, C.W., Graham, S.W., Pires, J.C., Stevenson, D.W., Zomlefer, W.B., Briggs, B.G., Duvall, M.R., Moore, M.J., Heaney, J.M., Soltis, D.E., Soltis, P.S., Thiele, K. & Leebens-Mack, J.H. (2010) Assembling the tree of the monocotyledons: plastom sequence phylogeny and evolution of Poales. Annals of the Missouri Botanical Garden 97: 584–616. https://doi.org/10.3417/2010023

Goncharov, N.P., Golovnina, K.A. & Kondratenko, E.Y. (2009) Taxonomy and molecular phylogeny of natural and artificial wheat species. Breeding Science 59: 492–498. https://doi.org/10.1270/jsbbs.59.492

González, T. & Morton, C.M. (2005) Molecular and morphological phylogenetic analysis of Brachiaria and Urochloa (Poaceae). Molecular Phylogenetics and Evolution 37: 36–44. https://doi.org/10.1016/j.ympev.2005.06.003

Gonzalo, R., Aedo, C. &.García, M.A. (2011a) Two new combination in stipa Sect. Smirnovia (Poaceae). Annales Botanici Fennici 48: 159–162. https://doi.org/10.5735/085.048.0209

Gonzalo, R., Aedo, C. & García, M.Á. (2011b) Taxonomic revision of Stipa section Smirnovia and section Subsmirnovia (Poaceae). Botanical Journal of the Linnean Society 167: 394–448. https://doi.org/10.1111/j.1095-8339.2011.01188.x

Gonzalo, R., Aedo, C. & García, M.A. (2013a) Taxonomic revision of Stipa subsection Pulcherrimae (Poaceae). Taxonomía y sistemática de Stipa Sect. Stipa L. (Poaceae) y grupos afines. [Tesis Doctoral].Universidad Complutense De Madrid.

Gonzalo, R., Aedo, C. & García, M.A. (2013b) Taxonomic revision of the Eurasian Stipa subsections Stipa and Tirsae (Poaceae). Systematic Botany 38: 344–378. https://doi.org/10.1600/036364413X666615

Govaerts, R.H.A. (2011) World checklist of selected plant families published update. Facilitated by the Trustees of the Royal Botanic Gardens, Kew. 

Grass Phylogeny Working Group I (GPWG I) (2001) Phylogeny and Subfamilial Classification of the Grasses (Poaceae) Annals of the Missouri Botanical Garden 88: 373–457.

Grass Phylogeny Working Group II (GPWG II) (2012) New grass phylogeny resolves deep evolutionary relationships and discovers C4 origins. New Phytologist 193: 304–312. https://doi.org/10.1111/j.1469-8137.2011.03972.x

Gutermann, W., Schneeweiss, G.M., Schonswetter, P., Staudinger, M. & Wallinger, C. (2014) The vascular plants of Kálamos (Ionian Islands, Greece): a catalogue (Materials towards a Flora Ionica). Phyton (Horn) 54: 161–196.

Hamasha, H.R., Hagen, K.B. & Röser, M. (2012) Stipa (Poaceae) and allies in the Old World: molecular phylogenetics realigns genus circumscription and gives evidence on the origin of American and Australian lineages. Plant Systematics and Evolution 298: 351–367. https://doi.org/10.1007/s00606-011-0549-5

Hamzeh’ee, B. (1992) Interesteding records of the Gramineae family from Iran. The Iranian Journal of Botany 5: 91–94.

Hamzeh’ee, B. (2000) Some new and noteworthy plant records from Iran. The Iranian Journal of Botany 8: 271–277.

Hamzeh’ee, B. & Jalili, A. (2002) A new report and an interesting record from Iran. The Iranian Journal of Botany 9: 187–190.

Hamzeh’ee, B. & Assadi, M. (2015) Taxonomical notes on the genus Piptatherum P.Beauv. (Poaceae) in Iran. The Iranian Journal of Botany 21: 1–9.

Hamzeh’ee, B., Alemi, M., Attar, F. & Ghahreman, A. (2007) Bromus catharticus Vahl and Bromus danthoniae var. uniaristatus (Poaceae), two new records from Iran. The Iranian Journal of Botany 13: 33–36. 

Hamzeh’ee, B., Ghahremaninejad, F., Bidar Lord, M. & Attar, F. (2008) Ventenata Koeler, a new genus (Gramineae: Pooideae) record for Iran. The Iranian Journal of Botany 14: 105–107. 

Hamzeh’ee, B. & Jalili, A. (2018) The importance of Poaceae in nature and human life. Iran Nature 2 (6): 46–55.

Hamzeh’ee, B. & Naqinezhad, A. (2009) Arthraxon P.Beauv (Gramineae) and Carex caryophyllea (Cyperceae): New genus and species records from Iran. The Iranian Journal of Botany 15: 68–71.

Hand, R. (Ed.) (2015) Supplementary notes to the flora of Cyprus VIII. Willdenowia 45: 245–259. https://doi.org/10.3372/wi.45.45210

Hempel, W. (2012) Revision und phylogenie der arten der gattung Melica L. (Poaceae) in Eurasien und Nordafrika. Feddes Repertorium 122: 1–253. https://doi.org/10.1002/fedr.201100029

Henderson, S. & Schäfer, H. (2003) Synopsis of the genus Rostraria (Poaceae) in the Azores. Botanical Journal of the Linnean Society 141: 125–131. https://doi.org/10.1046/j.1095-8339.2003.00114.x

Henrard, J.T. (1926) A critical revision of the genus Aristida. Mededeelingen van’s Rijks Herbarium 54: 1–220. [Leiden]

Heshmati, G.A. (2007) Vegetation characteristics of four ecological zones of Iran. International Journal of Plant Production 1: 215–224.

Heywood, V.H. (1985) Flowering Plants of the World, Cambridge, Cambridge Univ. Press.

Hodkinson, T.R. (2018) Evolution and taxonomy of the grasses (Poaceae): A model family for the study of species—rich Groups. Annual Plant Reviews 1: 1–39. https://doi.org/10.1002/9781119312994.apr0622

Hosseini, S.Z., Rahiminejad, M.R., Saeidi, H. & Assadi, M. (2013a) Taxonomic study of Festuca L. subgenus Schedonorus (P.Beauv.) Peterm. in Iran. Taxonomy and Biosystematics 16: 69–74.

Hosseini, S.Z., Rahiminejad, M.R. & Saeidi, H. (2013b) Leaf anatomical structure of Iranian narrow-leaved species of the genus Festuca L. (Poaceae, Poeae). The Iranian Journal of Botany 10: 86–93. 

IPNI (2019) International Plant Names Index. Checklist dataset. Available from: http://www.ipni.org (accessed August 2022).

Kavousi, M., Assadi, M. & Nejadsattari, T. (2015) Taxonomic revision of the genus Poa L. in Iran, new additions to Flora Iranica, and a new identification key. Turkish Journal of Botany 39: 105–127. https://doi.org/10.3906/bot-1311-31

Kavousi, M., Assadi, M. & Nejadsattari, T. (2009) Some new and noteworthy plant records from Iran. The Iranian Journal of Botany 8: 271–277. 

Kavousi, M., Assadi, M. & Nejadsattari, T. (2010) Two new records of poa L. (Poaceas) for the Flora of Iran.Journal of Sciences (Islamic Azad University) 74: 136–139.

Kellogg, E.A. (2015) Flowering plants. Monocots: Poaceae. In: Kubitski, K. (Ed.) The Families and Genera of Vascular Plants vol. 13. Springer International, Publishing AG, pp. 1–416.

Kellogg, E.A., Abbott, J.R., Bawa, K.S., Gandhi, K.N., Kailash, B.R., Ganeshaiah, K.N., Shrestha, U.B. & Raven, P.H. (2020) Checklist of the grasses of India. PhytoKeys 163: 1–560. https://doi.org/10.3897/phytokeys.163.38393

Keshavarzi, M., Khaksar, M. & Ghadam, P. (2012) Biosystematic study of Phalaris L. species (Poaceae) in Iran. Taxonomy and Biosystematics 13: 25–30.

Keshavarzi, M., Rahiminejad, M.R. & Kheradmandnia. (2007) Discriminativa analysis of 10 species Aeagilops L. (Poaceae) native to Iran. Iranian Journal of Science & Technology 31: 321–324.

Keshavarzi, M., Ijbari, H., Mosaferi, S. & Ebrahimi, F. (2017) A Floristic Study of Hamun Lake Basin, South East of Iran. Ecologia Balkanica 9: 1–9.

Khodashenas, M. (2008) Aeluropus peterganicus (Poaceae), a new species from Iran. The Iranian Journal of Botany 14: 13–15.

Khodashenas, M. (2009) Two new records and a new combination of the genus Aeluropus Trin (Poaceae) for the flora of Iran. The Iranian Journal of Botany 15: 61–62.

Khodashenas, M. (2011) A revision of the genera Phragmites and Schismus (Poaceae) in Iran. The Iranian Journal of Botany 17: 40–42.

Khodashenas, M. & Joharchi, M. (2012) Aeluropus laciniatus (Poaceae), a new species from Iran. The Iranian Journal of Botany 18: 64–66.

Kilian, B., Mammen, K., Millet, E., Sharma, R., Graner, A., Salamini, F., Hammer, K. & Özkan, H. (2011) Aegilops, In: Wild crop relatives: genomic and breeding resources, Springer, Berlin, Heidelberg, pp. 1–76. https://doi.org/10.1007/978-3-642-14228-4_1

Kimber, G. (1974) A reassessment of the origin of polyploid wheats. Genetics 78: 487–492. https://doi.org/10.1093/genetics/78.1.487

Kranz, A.R. (1957) Populationsgenetische Untersuchungen am Iranischen Primitivroggen. Ein Beitrag zur Systematik, Evolution und Ziichtung des Roggens. Z. Pflanzenziich 38: 101–146.

Komarov, V.L. (Ed.) (1934) The flora of USSR vol. 2. Moskow & Leningrad, Nauka.

Léonard, J. (1981) Contribution à l’étude de la flore et de la végétation des deserts d’Iran, Fas 1. Brussels: Jardin botanique nacional de Belgique.

Linder, H.P., Lehmann, C.E., Archibald, S., Osborne, C.P. & Richardson, D.M. (2018) Global grass (P oaceae) success underpinned by traits facilitating colonization, persistence and habitat transformation. Biological Reviews 93 (2): 1125–1144. https://doi.org/10.1111/brv.12388

Liu, Z.W. & Wang, R.R.C. (1993) Genome analysis of Elytrigia caespitosa, Lophopyrum nodosum, Pseudoroegneria geniculata ssp. scythica, and Thinopyrum intermedium (Triticeae: Gramineae). Genome 36: 102–111. https://doi.org/10.1139/g93-014

Lu, S.l. & Phillips, S.M. (2006) Polypogon Desfontaines (Poaceae). In: Wu, Z.Y., Raven, P.H. & Hong, D.Y. (Eds.) Flora of China. Vol. 22. Science Press, Beijing and Missouri Botanical Garden, St. Louis, pp. 361–363. Available from: http://www.efloras.org (accessed 12 March 2022).

Mabberley, D.J. (2018) Mabberley’s plant-book: a portable dictionary of plants, their classification and uses (No. Ed. 4). Cambridge University Press. https://doi.org/10.1017/9781316335581

Mahelka, V., Kopecky, D. & Pastova, L. (2011) On the genome constitution and evolution of intermediate wheatgrass (Thinopyrum intermedium: Poaceae, Triticeae). BMC Evolutionary Biology 11: 127. https://doi.org/10.1186/1471-2148-11-127

Markgraf-Dannenberg, I. (1985) Festuca. In: Davis, P.H. (Ed.) Flora of Turkey and the East Aegean Islands vol. 9. Edinburgh University Press, p.p. 400–442.

Maroofi, H. (2009) Two new records of Lotus (Fabaceae) and Phalaris (Poaceae) from Iran. The Iranian Journal of Botany 15: 76–79.

Maroofi, H. (2011) A new species and a new record of Poaceae family from Iran. The Iranian Journal of Botany 17: 10–14. 

Melderis, A. (1985) Agropyron Gaertner. In: Davis, P.H. (Ed.) Flora of Turkey and the East Aegean Islands, vol. 9. University Press, Edinburgh, Scotland, pp. 204–206. 

Memariani, F. & Arjmandi, A.A. (2013) Festuca karatavica (Poaceae), a new grass record for the flora of Iran. The Iranian Journal of Botany 19: 57–61.

Memariani, F., Joharchi, M.R. & Arjmandi, A.A. (2012) A revision of Bromus sect. Triniusa (Poaceae) in Khorassan (Iran). Rostaniha 13: 189–196.

Memariani, F., Joharchi, M.R. & Akhani, H. (2016) Plant diversity of Ghorkhod Protected Area, NE Iran. Phytotaxa 249: 118–158. https://doi.org/10.11646/phytotaxa.249.1.6

Memariani, F., Ejtehadi, H., Arjmandi, A.A & Joharchi, M.R. (2022) Piptatherum ferganense (Poaceae), another Middle Asian species as a new record for the flora of Iran. Nova Biologica Reperta 9: 132–138.

Mill, R.R. (1985) Bellardiochloa. In: Davis, P.H. (ed.) Flora of Turkey and the East Aegean Islands vol. 9. Edinburgh University Press, pp. 442–444.

Mobayen, S. (1975) Flora of Iran, vol. 1. University of Tehran Press, Tehran. pp. 372–381. [In Persian].

Mozaffarian, V. (1985) New species and new plant records. The Iranian Journal of Botany 3: 81–86.

Mozaffarian, V. (1991) New species and new plant records from Iran. The Iranian Journal of Botany 5: 29–39.

Mozaffarian, V. (1992) New species and intersting plant records from Iran. The Iranian Journal of Botany 5: 83–90.

Mozaffarian, V. (1994) Studies on the flora of Iran, new species and new records. The Iranian Journal of Botany 6: 235–243.

Naderi, R. & Rahiminejad, M.R. (2012) Some comments on the species of Bromus sect. Bromus (Poaceae) in Iran. The Iranian Journal of Botany 18: 226–230.

Naderi, R. & Rahiminejad, M.R. (2015) A taxonomic revision of the genus Bromus (Poaceae) and a new key to the tribe Bromeae in Iran. Annales Botanici Fennici 52: 233–248. https://doi.org/10.5735/085.052.0316

Naderi, R., Rahiminejad, M.R., Assadi, M. & Saeidi, H. (2012) Notes on the genus Bromus L. (Poaceae) in Iran. The Iranian Journal of Botany 18: 42–46.

Naderi, R., Rahiminejad, M.R., Assadi, M. & Vitek, E. (2016) A new taxonomic concept for Bromus danthoniae. Annalen des Naturhistorischen Museums in Wien, B 118: 167–180.

Naqinezhad, A. & Hamedani, H. (2017) Puccinellia poecilantha (Poaceae) in Iran. Phytotaxa 313: 185–194. https://doi.org/10.11646/phytotaxa.313.2.4

Naqinezhad, A., Ghahremaninejad, F. & Bahari, H. (2010) Monerma Beauv. (Gramineae) a new monotypic genus record for the flora of Iran. The Iranian Journal of Botany 17: 75–77.

Naqinezhad, A. (2012) Cenchrus longispinus (Poaceae), a new record from coastal sands of Caspian Sea (N Iran). Rostaniha 13: 211–214.

Nasernakhaei, F., Rahiminejad, M.R., Saeidi, H. & Tavasoli, M. (2013) Taxonomic identity of the Iranian diploid Triticum as evidenced by nrDNA ITS analysis. Phytotaxa 143: 43–53. https://doi.org/10.11646/phytotaxa.143.1.2

Nobis, M., Gudkova, P.D., Nowak, A., Sawicki, J. & Nobis, A. (2020) A Synopsis of the Genus Stipa (Poaceae) in Middle Asia, Including a Key to Species Identification, an Annotated Checklist, and Phytogeographic Analyses1. Annals of the Missouri Botanical Garden 105: 1–63. https://doi.org/10.3417/2019378

Noori, A., Nourozi, M., Azizian, D., Sheidai, M. & Termeh, F. (2004) Bromus arvensis (Poaceae), a new record for the flora of Iran. The Iranian Journal of Botany10: 173–175.

Noroozi, J., Moser, D. & Essl, F. (2016) Diversity, distribution, ecology and description rates of alpine endemic plant species from Iranian mountains. Alpine Botany 126: 1–9. https://doi.org/10.1007/s00035-015-0160-4

Nourouzi, M. Sheydaei, M., Nouri, A. & Assadi, M. (2005) Bromus secalinus L. (Poaceae), a new record for flora of Iran. The Iranian Journal of Botany 11: 71–73.

Parsa, A. (1950) Flore de l, Iran. 5 Ministere de l’ Education, Tehran. pp. 817–828.

Peterson, P.M., Romaschenko, K., Herrera Arrieta, Y. & Saarela, J.M. (2014) A molecular phylogeny and new subgeneric classification of Sporobolus (Poaceae: Chloridoideae: Sporobolinae). Taxon 63: 1212–1243. https://doi.org/10.12705/636.19

Peterson, P.M., Romaschenko, K. & Arrieta, Y.H. (2015) A molecular phylogeny and classification of the Eleusininae with a new genus, Micrachne (Poaceae: Chloridoideae: Cynodonteae). Taxon 64: 445–467. https://doi.org/10.12705/643.5

Peterson, P.M., Romaschenko, K., Snow, N. & Johnson, G. (2012) A molecular phylogeny and classi?cation of Leptochloa (Poaceae: Chloridoideae: Chlorideae) sensu lato and related genera. Annals of Botany 109: 1317–1329. https://doi.org/10.1093/aob/mcs077

Petersen, G. & Seberg, O. (2008) Phylogenetic relationships of allotetraploid Hordelymus europaeus (L.) Harz (Poaceae: Triticeae). Plant Systematics and Evolution 273: 87–95. https://doi.org/10.1007/s00606-008-0014-2

POWO (2020) Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. Available from: http://www.plantsoftheworldonline.org (accessed 25 February 2021).

Quintanar, A., Catalan, P. & Castroviejo, S. (2010) A review of the systematics and phylogenetics of the Koeleriina (Poaceae: Poeae). In: Seberg, O., Petersen, G., Barfod, A.S. & Davis, J.I. (Eds.) Diversity, phylogeny, and evolution in the monocotyledons. Aarhus, Denmark: Aarhus University Press. pp. 539–556.

Raeisi Chehrazi, S., Sharifi-Tehrani, M. & Saeidi, H. (2015) Relationships of Festuca–Lolium complex in Iran assessed using trnH-psbA sequences. Taxonomy and Biosystematics 23: 57–68.

Rahiminejad, M.R., Sahebi, J. & Nasser–Nakhaie, F. (2005) A morphological study and a taxonomic revision of the genus Secale L. (Triticeae, Poaceae) in Iran. The Iranian Journal of Botany 11: 1–14.

Rahiminejad, M.R. & Kharazian, N. (2005) Illustration of and a key to the genus Triticum L. in Iran. Wheat Information Service 99: 29–34.

Rahmanian, S., Saeidi, H. Assadi, M. & Rahiminejad, M.R. (2014) A taxonomic revision of the genus Eremopoa Roschv. (Poaceae, Poeae) in Iran. The Iranian Journal of Botany 20: 8–15.

Rahmati, F. & Saeidi, H. (2012) A cytotaxonomic study of Catabrosa P.Beauv. (Poaceae) in Iran. Taxonomy and Biosystematics 4: 37–42.

Rathakrishnan, N.C. & Nair, N.C. (1983) The correct name for Gastrochilus pulchellus (Orchidaceae). Taxon 32: 638–639. https://doi.org/10.2307/1221745

Rechinger, K.H. (1977) Plant of the Touran Protected Area. Iran [with a contribution by P.Wendelbo (Chenopodiaceae)]. The Iranian Journal of Botany 1: 97–108.

Rezaei, E., Saeidi, H. & Rahiminejad, M.R. (2011) A taxonomic revision of the genus Cynosurus L. (Poaceae) in Iran. The Iranian Journal of Botany 17: 212–219.

Romaschenko, K., Peterson, P.M., Soreng, R.J., Futorna, O. & Susanna, A. (2011) Phylogenetics of Piptatherum s.l. (Poaceae: Stipeae): Evidence for a new genus, Piptatheropsis, and resurrection of Patis. Taxon 60: 1703–1716. https://doi.org/10.1002/tax.606015

Romero-Zarco, C. (2011) Helictochloa Romero Zarco (Poaceae), a new genus of oat grass. Candollea 66: 87–103.

Röser, M. (2012) Stipellula, a new genus, and new combinations in feather grasses (Poaceae tribe Stipeae). Schlechtendalia 24: 91–94.

Saarela, J.M., Bull, R.D., Paradis, M.J., Ebata, S.N., Soreng, R.J., Peterson, P.M. & Paszko, B. (2017) Molecular phylogenetics of cool-season grasses in the subtribes Agrostidinae, Anthoxanthinae, Aveninae, Brizinae, Calothecinae, Koeleriinae and Phalaridinae (Poaceae: Pooideae: Poeae: Poeae chloroplast. PhytoKeys 87: 1–139. https://doi.org/10.3897/phytokeys.87.12774

Saarela, J.M., Liu, Q., Peterson, P.M., Soreng, R.J., Paszko, B. (2010) Phylogenetics of the grass ‘Aveneae-type plastid DNA clade’ (Poaceae: Pooideae, Poeae) based on plastid and nuclear ribosomal DNA sequence data. In: Seberg, O., Petersen, G., Barfod, A.S. & Davis, J.I. (Eds.) Diversity, phylogeny, and evolution in the monocotyledons. Aarhus, Denmark: Aarhus University Press. pp. 557–587.

Saeidi, H. & Rahmati, F. (2012) A cytotaxonomic study of Catabrosa P.Beauv. (Poaceae) in Iran. Taxonomy and Biosystematics 12: 37–42.

Sahebi, J., Rahiminejad, M.R & Ghahreman, A. (2004) A taxonomic review of the Hordeum marinum sensu lato (Poaceae: Triticeae) in Iran. Iranian Journal of Science & Technology 28: 127–135.

Salimi, A., Ebrahimzadeh, H. & Taeb, M. (2005) Description of Iranian diploid wheat resources. Genetic Resources and Crop Evolution 52 (4): 351–361. https://doi.org/10.1007/s10722-005-2256-y

Salimi, S. & Rahiminejad, M.R. (2016) A new record of the genus Sporobolus (Poaceae) for the flora of Iran. The Iranian Journal of Botany 22: 6–7.

Scholz, H. (1979) Quanten-Artbildung auch bei Stipagrostis (Gramineae). Willdenowia 9: 209–214.

Scholz, H. (1981a) Der Bromus-pectinatus-Komplex(Gramineae) im Nahen und Mittleren Osten. Botanische Jahrbücher für Systematik 102: 471–495.

Scholz, H. (1981b) Drei neue Gramineen aus Iran und Libyen. Willdenowia 11: 95–100.

Scholz, H. (1981c) Ergänzungen und Verbesserungen zur “Flora iranica”: Gramineae. Willdenowia 11: 259–266.

Scholz, H. (1985) Stipa L. In: Davis, P.H. (Ed.) Flora of Turkey and the East Aegean Islands 9. Edinburgh University Press, Edinburgh. pp. 541–552.

Scholz, H. (1987) Delimitation and classification of Bromus fasciculatus (Poaceae). Plant Systematics and Evolution 155: 277–282. https://doi.org/10.1007/BF00936304

Scholz, H. (1988) Eine neue Stipagrostis (Gramineae) aus den Sand-Wüsten Irans und Afghanistan. Willdenowia 17: 107–109.

Scholz, H. (1989) Floristisches und Taxonomisches über einige Gramineae von Rhodos und Symi (Ost-Ägäis, Griechenland). Willdenowia 19: 105–110.

Scholz, H. (1990) Centropodia forskalii subsp. persica (Gramineae): eine Neubewertung = Centropodia forskalii subsp. persica (Gramineae): a revaluation. Willdenowia 19: 445–44

Scholz, H. (1998) Notes on Bromus danthoniae and relatives (Poaceae). Willdenowia 28: 143–150. https://doi.org/10.3372/wi.28.2813

Shouliang, Ch., Phillips, S.M. & Renvoize, S.A. (2006) Tribe Paniceae. In: Wu, Z., Raven, P.H., Hong, D. (Eds.) Flora of China. Beijing: Science Press. pp. 499–554.

Shu, T.M. (2006) Glyceria. In: Wu, Z., Raven, P.H. & Hong, D. (Eds.) Flora of China, vol. 22, Poaceae, 213–216. Science Press, Beijing, China and Missouri Botanical Garden Press, St. Louis, Missouri, USA.

Smith, P.M. (1968) The Bromus mollis aggregate in Britain. Watsonia 6: 327–344.

Soreng, R.J., Peterson, P.M, Romaschenko K, Davidse, G., Zuloaga, F.O., Judziewicz, E.J., Filgueiras, T.S., Davis, J.I. & Morrone, O. (2015) A worldwide phylogenetic classification of the Poaceae (Gramineae). Journal of Systematics and Evolution 53: 117–137. https://doi.org/10.1111/jse.12150

Soreng, R.J. Peterson, P.M, Romaschenko, K, Davidse, G, Teisher, J.K.,Clark, L.G., Barber, P., Gillespie, L.J. & Zuloag, F.O. (2017) A worldwide phylogenetic classification of the Poaceae (Gramineae) II: An update and a comparison of two 2015 classifications. Journal of Systematics and Evolution 55: 235–410. https://doi.org/10.1111/jse.12262

Stace, C.A. (1985) Catapodium. In: Davis, P.H. (Ed.) Flora of Turkey and the East Aegean Islands vol. 9. Edinburgh University Press, p.p. 461–465.

Taheri, Gh., Assadi, M., Nejadsattari, T., Fallahian, F., Joharchi, M.R. & Bahrami, A.R. (2006) Apera interrupta (Poaceae), a new record for the flora of Iran. The Iranian Journal of Botany 12: 67–68. 

Taheri, Gh. (2013) New species and new records of Agrostis for the flora of Iran. The Iranian Journal of Botany 19: 40–43.

Talebi, A., Naqinezhad, A. & Attar, F. (2022) Trisetum spicatum (Poaceae: Pooideae): an interglacial relict in the Iranian mountains. Phytotaxa 532 (1): 85–92. https://doi.org/10.11646/phytotaxa.532.1.7

Takhtajan, A.L. (1986) Floristic Regions of the World. University of California Press. California.

 Termeh, F. (1975) Contribution á l’étude de quelques Graminées nouvelles pour la flora de l’Iran. Institut de Recherches Entomologiques et Phytopathologiques d’Evine, Departement de Botanique No. 5. Teheran. 

Termeh, F. (1987) Contribution á l’étude de quelques Graminées nouvelles pour la flora de l’Iran. Fasc. 2. Ministry Agriculture, Plant Pests & Diseaeses Research Institute, Teheran. 

Termeh, F. (2000) New records of the family Gramineae from Iran (3). Rostaniha 1: 21–25

Tirgan, S., Naqinezhad, A. & Hoseinzadeh, Z. (2019) Cutandia rigescens (Poaceae) a new species to the flora of Iran: distribution, taxonomy and ecology. Phytotaxa 419: 287–300. https://doi.org/10.11646/phytotaxa.419.3.3

Torrecilla, P., Angel, J., Rodríguez, L. & Catalán, P. (2004) Phylogenetic relationships of Vulpia and related genera (Poeae, Poaceae) based on analysis of ITS and trnL-F sequences. Annals of the Missouri Botanical Garden 91: 124–158.

Torrecilla, P. & Catalán, P. (2002) Phylogeny of broad-leaved and fine-leaved Festuca lineages (Poaceae) based on nuclear ITS sequences. Systematic Botany 27: 241–251.

Torrecilla, P., José-Angel, López-Rodríguez & Catalán, P. (2004) Phylogenetic relationships of Vulpia and related genera (Poeae, Poaceae) based on analysis of ITS and trnL-F sequences. Annals of the Missouri Botanical Garden 91: 124–158.

Tropicos.org (2018–present) Tropicos.org. Missouri Botanical Garden. Available from: http://www.tropicos.org (accessed 17 July 2018)

Tzvelev, N.N. (1997) A new species of the genus Festuca (Poaceae) from Iran. Botanicheskii Zhurnal 82: 118.

Tzvelev, N.N. (984) Grasses of the Soviet Union, Part I, II. A.A. Fedorov (Chief editor). Russian Translaction series 8, AA. Balkema, Rotterdam. pp. 1121.

Uosofi, M. (2002) Elymus lazicus subsp. attenuatiglumis, a new record for Iran. The Iranian Journal of Botany 9: 249–251.

Valdés, B. & Scholz, H. (2006) The Euro+ Med treatment of Gramineae—a generic synopsis and some new names. Willdenowia 36 (2): 657–669. https://doi.org/10.3372/wi.36.36202

Watson, L. & Dallwitz, M.J. (1999) Grass genera of the world: descriptions, illustrations, identification, and information retrieval, including synonyms, morphology, anatomy, physiology, phytochemistry, cytology, classification, pathogens, world and local distribution, and references. Available from: http://biodiversityunoedu/delta/Version (accessed 15 November 2019). 

Witcombe, J.R. (1983) A guide to the species of Aegilops L. Their taxonomy, morphology and distribution. International Board for Plant Genetic Resources Wheat Programme, Rome, Italy.

Wölk, A. & Röser, M. (2014) Polyploid evolution, intercontinental biogeographical relationships and morphology of the recently described African oat genus Trisetopsis (Poaceae). Taxon 63: 773–788. https://doi.org/10.12705/634.1

Wölk, A. & Röser, M. (2017) Hybridization and long-distance colonization in oat-like grasses of South and East Asia, including an amended circumscription of Helictotrichon and the description of the new genus Tzveleviochloa (Poaceae). Taxon 66: 20–43. https://doi.org/10.12705/661.2

Wölk, A., Winterfeld, G. & Röser, M. (2015) Genome evolution in a Mediterranean species complex: phylogeny and cytogenetics of Helictotrichon (Poaceae) allopolyploids based on nuclear DNA sequences (rDNA, topoisomerase gene) and FISH. Systematics and Biodiversity 13: 326–345. https://doi.org/10.1080/14772000.2015.1023867

Wu, Z. & Phillips, S.M. (2006) Tribe Stipeae. In: Wu, Z., Raven, P.H. & Hong, D. (eds.) Flora of China. Beijing: Science Press. pp. 188–212.

Yen, C. & Yang, J. (1990) Kengyilia gobicola, a new taxon from west China. Canadian Journal of Botany 68: 1894–1897. https://doi.org/10.1139/b90-248

Yen, C. & Yang, J. (1998) New taxa, new combinations, and observations in Kengyilia (Poaceae: Triticeae). Novon 8: 94–100. https://doi.org/10.2307/3391904

Yen, C. & Yang, J. (2020) Biosystematics of Triticeae vol I. Triticum-Aegilops complex. China Agriculture Press & Springer Nature Singapore Pte Ltd. pp. 246. https://doi.org/10.1007/978-981-13-9931-2

Yen, C., Yang, J. & Baum, B.R. (2009) Synopsis of Leymus Hochst. (Triticeae: Poaceae). Journal of Systematics and Evolution 47: 67–86. https://doi.org/10.1111/j.1759-6831.2009.00004.x

Yousofi, M. & Assadi, M. (2006) Tow new taxa of Agropyron cristatum (Poaceae), for the flora of Iran. The Iranian Journal of Botany 12: 47–50.

Zarco, C.R. (2015) Las gramíneas de la Península Ibérica e Islas Baleares: claves ilustradas para la determinación de los géneros y catálogo preliminar de las especies, vol. 15. Jolube Consultor Botánico y Editor, 170 pp.

Zhang, X. & Sun, G. (2010) RPB2 sequences reveal a close phylogenetic relationship between tetraploid Hordelymus and diploid Hordeum species in Triticeae (Poaceae). Biochemical Systematics and Ecology 38: 789–795. https://doi.org/10.1016/j.bse.2010.08.001

Zhu, G., Liu, L., Soreng, R.J. & Olonova, M.V. (2006) Poa. In: Wu, Z.Y., Raven, P.H. & Hong, D.Y. (Eds.) Flora of China, Vol. 22. Science Press, Beijing and Missouri Botanical Garden, St. Louis, pp. 257–309. Available from: http://www.efloras. org (accessed 12 March 2022).

Zohary, M. (1963) On the Geobotanical Stucture of Iran. Bulletin of the Research Council of Palestina suppl. to vol. 11/D (Botany). https://doi.org/10.24157/ARC_13414

Zohdi, M., Arzani, H., Javadi, S.A., Jalili, A. & Khorshidi, G. (2018) Government and Range Management in Iran (Policy, Laws, and Plans). Applied Ecology and Environmental Research 16 (4): 4637–4654. https://doi.org/10.15666/aeer/1604_46374654

Zoughi, S., Saeidi, H. & Rahiminejad, M.R. (2011) A taxonomical, morphological and cytological revision of the genus Nardurus Reichenb. (Poeae, Poaceae) in Iran. The Iranian Journal of Botany 17: 189–199.

How to Cite

Bidarlord, M. & Ghahremaninejad, F. (2022)

A checklist of Iranian Grasses

. Phytotaxa 574 (1): 1–31. https://doi.org/10.11646/phytotaxa.574.1.1

A checklist of Iranian Grasses

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