Abstract
Swertia drassensis (Gentianaceae) is described as a new species from Drass, an area located in the Trans-Himalaya region of Ladakh, India. This new species resembles in few characters to its congeneric Swertia petiolata, but clearly differs from it in possessing a pair of fused nectaries, fimbriae characteristically absent, margin of sepals slightly membranous, greenish-yellow, ovoid-ellipsoid capsule and winged seeds. Photo-illustrations and comparison of diagnostic characters of the new species with other congeneric species are provided to facilitate field identification.
References
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Banoo, S., Gulzar, R., Islam, T., Ganie, A.H. & Khuroo, A.A. Gentianopsis paludosa var. alpine (Gentianaceae) a new record for India from Ladakh, Trans-Himalaya. Rheedea 31 (2): 71–76. https://dx.doi.org/10.22244/rheedea.2021.31.02.06</span></span></span>
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<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Cheek, M., Nic Lughadha, E., Kirk, P., Lindon, H., Carretero, J., Looney, Carretero, J., Looney, B., Douglas, B., Haelewaters, D., Gaya, E., Llewellyn, T., Ainsworth, A.M., Gofforov, Y., Hyde, K., Crous, P., Hughes, M., Walker, B.E.., Forzza, R.C., Wong, K.M. & Niskanen, T. (2020) New scientific discoveries: Plants and fungi. Plants People Planet 2 (5): 371–388. https://doi.org/10.1002/ppp3.10148</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Chowdhery, H.J. & Wadhwa, B.M. (1984) Flora of Himachal Pradesh. Vols.1–3. Botanical Survey of India, Calcutta, India.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Dar, G. H & Khuroo, A.A. (2020) Biodiversity of the Himalaya: Jammu and Kashmir State (Vol. 18). Springer Nature, Singapore, pp. 3–26. https://doi.org/10.1007/978-981-32-9174-4 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Don, D. (1837) Descriptions of Indian Gentianeae. Transactions of the Linnean Society of London 17, 519 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Dvorský, M., Klimeš, L., Doležal, J., Wild, J. & Dickoré, W.B. (2018) A field guide to the flora of Ladakh. Academia, Prague, Czech Republic, pp. 172–174.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">FOC (2008) e Flora of China. Available from: http://www.efloras.org/. (accessed 13 July 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">FOP (2002) e Flora of Pakistan. Available from: http://www.efloras.org/. (accessed 13 July 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ganie, A.H., Butt, T.A., Khuroo, A.A., Rasool, N., Ahmad, R., Basharat, S. & Reshi, Z.A. (2022) Taxonomy and threat assessment of Lagotis kunawurensis Rupr. (Plantaginaceae), an endemic medicinal plant species of the Himalaya, India. Journal of Threatened Taxa 14 (6): 21239–21245. https://doi.org/10.1038/ s41598-018-36773-7 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Garg, S. (1987) Gentianaceae of the North West Himalaya, a revision. Today & Tomorrow’s Printers and Publishers, 342 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gogoi, R., Sherpa, N., Benjamin, J.H.F., Agrawala, D.K., Rai, S.K. & Dash, S.S. (2021) Flora of Sikkim. Botanical Survey of India, Kolkata, India.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gupta, P., Sangwan, N., Lal, R. & Vakhlu, J. (2015) Bacterial diversity of Drass, cold desert in Western Himalaya, and its comparison with Antarctic and Arctic. Archives of microbiology 197 (6): 851–860.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hamid, M., Khuroo, A.A., Charles, B., Ahmad, R., Singh, C.P. & Aravind, N.A. (2019) Impact of climate change on the distribution range and niche dynamics of Himalayan birch, a typical treeline species in Himalayas. Biodiversity and Conservation 28 (8): 2345–2370. https://doi.org/10.1007/s10531-018-1641-8 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hamid, M., Khuroo, A.A., Malik, A.H., Ahmad, R., Singh, C.P., Dolezal, J. & Haq, S.M. (2020) Early evidence of shifts in alpine summit vegetation: a case study from Kashmir Himalaya. Frontiers in Plant Science 11: 421. https://doi.org/10.3389/fpls.2020.00421 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hooker, J.D. (1883) The Flora of British India. Vol. IV. Reeve & Co. Ltd. Ashford. Kent. London, pp. 121–130.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ho, T.N. & Liu, S.W. (1980) New taxa of Swertia L. from China. Acta Phytotaxonomica Sinica 18 (1): 75–85.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ho, T.N. & Liu, S.W. (2015) A world wide monograph of Swertia and its allies. Science Press, Beijing, 430 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hortal, J., Bello, F.D., Diniz-Filho, J.A.F., Lewinsohn, T.M., Lobo, J.M. & Ladle, R.J. (2015) Seven Shortfalls that Beset Large-Scale Knowledge of Biodiversity. Annual Review of Ecology, Evolution, and Systematics (46): 523–549. https://doi.org/10.1146/annurev-ecolsys-112414-054400 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Islam, T., Khuroo, A.A. & Nawchoo, I.A. (2022) Swertia pahalgamensis (Gentianaceae), a new species from Kashmir Himalaya, India. Phytotaxa 547 (2): 201–207. https://doi.org/10.11646/phytotaxa.547.2.8.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Jain, S.K. & Rao, R.R. (1976) A handbook of field and herbarium methods. Today and Tomorrow’s Printers and Publishers, New Delhi.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Maiti, G. & Banerji, M.L. (1976) Exomorphic seed structure of the Himalayan species of Swertia Linn. (Gentianaceae). Proceedings of the Indian Academy of Science Sec B 6: 231–237. https://doi.org/10.1007/BF03045546 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nampy, S., Shahina, P.M., Haseena, T. & & Ashwini, H.S. (2015) A taxonomic revision of Swertia L. (Gentianaceae) in South India, with one new species and seven lectotypifications. Phytotaxa 195 (1): 31–52. https://doi.org/10.11646/phytotaxa.195.1.2 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">POWO (2021) Plants of the world online.Royal Botanic Gardens, Kew, UK. Available from: http://www.plantsoftheworldonline.org/taxon/urn:lsid:ipni.org/ (accessed 12 July 2022)</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pringle, J.S. (2014) Morphological characteristics of the family Gentianaceae. In: The Gentianaceae-Volume 1: Characterization and Ecology. Springer, Berlin, Heidelberg pp. 1–12.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">QGIS Version 3.20 (2021) Geographic Information System. Open Source Geospatial Foundation Project. Available from: http://qgis.osgeo.org/ (accessed 10 September 2021)</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rather, Z.A., Hussain, K., Dwivedi, M.D., Dar, T.U.H., Dar, A.R. & Khuroo, A.A. (2022) ‘New’ species are not always new: a case study of Ephedra sumlingensis and E. khurikensis (Ephedraceae). Plant Systematics and Evolution 308 (4): 1–16. https://doi.org/10.1007/s00606-022-01815-1 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Shah, J. (1990) Taxonomic studies in the genus Swertia L. (Gentianaceae): Monograph. Part I. Scientific Khyber 3: 17–114.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Shah, J. (1992) Taxonomic studies in the genus Swertia L. (Gentianaceae): Monograph. Part II. Scientific Khyber 5: 127–231.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Singh, B. (2021) Swertia patnitopiansis, a new species of Gentianaceae from North-Western Himalaya, India. Phytotaxa 498 (4): 273–280. https://doi.org/10.11646/phytotaxa.498.4.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Singh, N.P., Singh, D.K. & Uniyal, B.P. (2002) Flora of Jammu and Kashmir, Vol.1. Botanical Survey of India, Kolkata, India.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Smith, H. (1970) New or little known Himalayan species of Swertia and Veratrilla (Gentianaceae). Bulletin of the British Museum (Natural History), Botany 4: 237–258.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Stewart, R.R., Nasir, E. & Ali, S.I. (1972) Flora of West Pakistan: an annotated catalogue of the vascular plants of West Pakistan and Kashmir. Fakhri Printing Press, Karachi, Pakistan, pp. 561–563.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Uniyal, B.P., Sharma, J.R., Choudhery, U. & Singh, D.K. (2007) Flowering Plants of Uttarakhand: A Checklist. Bishen Singh Mahendra Pal Singh, Dehradun.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, J.C. & Lu, C.T. (1998) Revision of the genus Swertia L. (Gentianaceae) in Taiwan. Taiwania 43: 273–288.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wani, B.A., Islam, T., Khuroo, A.A., Ganie, A.H. & Nawchoo, I.A. (2022) Swertia kashmirensis, a new species from Bangus Valley of Kashmir Himalaya, India. Phytotaxa 532 (1): 93–100. https://doi.org/10.11646/phytotaxa.532.1.8 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Bridson, D. & Forman, L. (1998) The herbarium handbook. Royal Botanic Gardens, Kew.Richmond, Surrey, United Kingdom. 261 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Cheek, M., Nic Lughadha, E., Kirk, P., Lindon, H., Carretero, J., Looney, Carretero, J., Looney, B., Douglas, B., Haelewaters, D., Gaya, E., Llewellyn, T., Ainsworth, A.M., Gofforov, Y., Hyde, K., Crous, P., Hughes, M., Walker, B.E.., Forzza, R.C., Wong, K.M. & Niskanen, T. (2020) New scientific discoveries: Plants and fungi. Plants People Planet 2 (5): 371–388. https://doi.org/10.1002/ppp3.10148</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Chowdhery, H.J. & Wadhwa, B.M. (1984) Flora of Himachal Pradesh. Vols.1–3. Botanical Survey of India, Calcutta, India.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Dar, G. H & Khuroo, A.A. (2020) Biodiversity of the Himalaya: Jammu and Kashmir State (Vol. 18). Springer Nature, Singapore, pp. 3–26. https://doi.org/10.1007/978-981-32-9174-4 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Don, D. (1837) Descriptions of Indian Gentianeae. Transactions of the Linnean Society of London 17, 519 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Dvorský, M., Klimeš, L., Doležal, J., Wild, J. & Dickoré, W.B. (2018) A field guide to the flora of Ladakh. Academia, Prague, Czech Republic, pp. 172–174.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">FOC (2008) e Flora of China. Available from: http://www.efloras.org/. (accessed 13 July 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">FOP (2002) e Flora of Pakistan. Available from: http://www.efloras.org/. (accessed 13 July 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ganie, A.H., Butt, T.A., Khuroo, A.A., Rasool, N., Ahmad, R., Basharat, S. & Reshi, Z.A. (2022) Taxonomy and threat assessment of Lagotis kunawurensis Rupr. (Plantaginaceae), an endemic medicinal plant species of the Himalaya, India. Journal of Threatened Taxa 14 (6): 21239–21245. https://doi.org/10.1038/ s41598-018-36773-7 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Garg, S. (1987) Gentianaceae of the North West Himalaya, a revision. Today & Tomorrow’s Printers and Publishers, 342 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gogoi, R., Sherpa, N., Benjamin, J.H.F., Agrawala, D.K., Rai, S.K. & Dash, S.S. (2021) Flora of Sikkim. Botanical Survey of India, Kolkata, India.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gupta, P., Sangwan, N., Lal, R. & Vakhlu, J. (2015) Bacterial diversity of Drass, cold desert in Western Himalaya, and its comparison with Antarctic and Arctic. Archives of microbiology 197 (6): 851–860.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hamid, M., Khuroo, A.A., Charles, B., Ahmad, R., Singh, C.P. & Aravind, N.A. (2019) Impact of climate change on the distribution range and niche dynamics of Himalayan birch, a typical treeline species in Himalayas. Biodiversity and Conservation 28 (8): 2345–2370. https://doi.org/10.1007/s10531-018-1641-8 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hamid, M., Khuroo, A.A., Malik, A.H., Ahmad, R., Singh, C.P., Dolezal, J. & Haq, S.M. (2020) Early evidence of shifts in alpine summit vegetation: a case study from Kashmir Himalaya. Frontiers in Plant Science 11: 421. https://doi.org/10.3389/fpls.2020.00421 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hooker, J.D. (1883) The Flora of British India. Vol. IV. Reeve & Co. Ltd. Ashford. Kent. London, pp. 121–130.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ho, T.N. & Liu, S.W. (1980) New taxa of Swertia L. from China. Acta Phytotaxonomica Sinica 18 (1): 75–85.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ho, T.N. & Liu, S.W. (2015) A world wide monograph of Swertia and its allies. Science Press, Beijing, 430 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hortal, J., Bello, F.D., Diniz-Filho, J.A.F., Lewinsohn, T.M., Lobo, J.M. & Ladle, R.J. (2015) Seven Shortfalls that Beset Large-Scale Knowledge of Biodiversity. Annual Review of Ecology, Evolution, and Systematics (46): 523–549. https://doi.org/10.1146/annurev-ecolsys-112414-054400 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Islam, T., Khuroo, A.A. & Nawchoo, I.A. (2022) Swertia pahalgamensis (Gentianaceae), a new species from Kashmir Himalaya, India. Phytotaxa 547 (2): 201–207. https://doi.org/10.11646/phytotaxa.547.2.8.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Jain, S.K. & Rao, R.R. (1976) A handbook of field and herbarium methods. Today and Tomorrow’s Printers and Publishers, New Delhi.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Maiti, G. & Banerji, M.L. (1976) Exomorphic seed structure of the Himalayan species of Swertia Linn. (Gentianaceae). Proceedings of the Indian Academy of Science Sec B 6: 231–237. https://doi.org/10.1007/BF03045546 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nampy, S., Shahina, P.M., Haseena, T. & & Ashwini, H.S. (2015) A taxonomic revision of Swertia L. (Gentianaceae) in South India, with one new species and seven lectotypifications. Phytotaxa 195 (1): 31–52. https://doi.org/10.11646/phytotaxa.195.1.2 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">POWO (2021) Plants of the world online.Royal Botanic Gardens, Kew, UK. Available from: http://www.plantsoftheworldonline.org/taxon/urn:lsid:ipni.org/ (accessed 12 July 2022)</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pringle, J.S. (2014) Morphological characteristics of the family Gentianaceae. In: The Gentianaceae-Volume 1: Characterization and Ecology. Springer, Berlin, Heidelberg pp. 1–12.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">QGIS Version 3.20 (2021) Geographic Information System. Open Source Geospatial Foundation Project. Available from: http://qgis.osgeo.org/ (accessed 10 September 2021)</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rather, Z.A., Hussain, K., Dwivedi, M.D., Dar, T.U.H., Dar, A.R. & Khuroo, A.A. (2022) ‘New’ species are not always new: a case study of Ephedra sumlingensis and E. khurikensis (Ephedraceae). Plant Systematics and Evolution 308 (4): 1–16. https://doi.org/10.1007/s00606-022-01815-1 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Shah, J. (1990) Taxonomic studies in the genus Swertia L. (Gentianaceae): Monograph. Part I. Scientific Khyber 3: 17–114.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Shah, J. (1992) Taxonomic studies in the genus Swertia L. (Gentianaceae): Monograph. Part II. Scientific Khyber 5: 127–231.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Singh, B. (2021) Swertia patnitopiansis, a new species of Gentianaceae from North-Western Himalaya, India. Phytotaxa 498 (4): 273–280. https://doi.org/10.11646/phytotaxa.498.4.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Singh, N.P., Singh, D.K. & Uniyal, B.P. (2002) Flora of Jammu and Kashmir, Vol.1. Botanical Survey of India, Kolkata, India.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Smith, H. (1970) New or little known Himalayan species of Swertia and Veratrilla (Gentianaceae). Bulletin of the British Museum (Natural History), Botany 4: 237–258.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Stewart, R.R., Nasir, E. & Ali, S.I. (1972) Flora of West Pakistan: an annotated catalogue of the vascular plants of West Pakistan and Kashmir. Fakhri Printing Press, Karachi, Pakistan, pp. 561–563.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Uniyal, B.P., Sharma, J.R., Choudhery, U. & Singh, D.K. (2007) Flowering Plants of Uttarakhand: A Checklist. Bishen Singh Mahendra Pal Singh, Dehradun.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, J.C. & Lu, C.T. (1998) Revision of the genus Swertia L. (Gentianaceae) in Taiwan. Taiwania 43: 273–288.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wani, B.A., Islam, T., Khuroo, A.A., Ganie, A.H. & Nawchoo, I.A. (2022) Swertia kashmirensis, a new species from Bangus Valley of Kashmir Himalaya, India. Phytotaxa 532 (1): 93–100. https://doi.org/10.11646/phytotaxa.532.1.8 </span></span></span>