Abstract
Thalictrum hengduanshanense and T. longistipitatum (Ranunculaceae), two new species from the southern Hengduan Mountains region in southeastern Xizang and northwestern Yunnan, China, are illustrated and described. Morphologically T. hengduanshanense is most closely similar to T. tsawarungense in having narrowly oblanceolate-linear filaments of stamens and apically slightly recurved styles, but differs by having laxly branched, corymb-like (vs. narrowly paniculiform) inflorescences and larger (ca. 9 mm vs. 4.5‒7 mm long), narrowly lunate (vs. broadly lunate) achenes (body width/body length = ca. 1/3 vs. ca. 1/2). Thalictrum longistipitatum is closely similar to T. hengduanshanense in habit and in having narrowly oblanceolate-linear filaments of stamens, apically slightly recurved styles, and long stipitate achenes, but differs by having apically abruptly recurved (vs. straight) pedicels and shorter styles (ca. 1 mm vs. ca. 2 mm long). Chromosome numbers of T. longistipitatum (2n = 14), T. rostellatum (2n = 28), T. tsawarungense (2n = 14) and T. wangii (2n = 14) are reported here for the first time.
References
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Boivin, B. (1945) Notes on some Chinese and Korean species of <em>Thalictrum</em>. <em>Journal of the Arnold Arboretum </em>26: 111–118. https://doi.org/10.5962/p.185520</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Boufford, D.E. & van Dyck, P.P. (1999) South-Central China. <em>In</em>: Mittermeier, R.A. & Mittermeier, C.G. (Eds.) <em>Hotspots: Earth’s Biologically Richest and Most Endangered Terrestrial Ecoregions</em>. CEMEX, Mexico City, pp. 338–351.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Emura, H.K.M. (1972) Cytotaxonomical studies on the genus <em>Thalictrum</em> in Eurasia with special reference to Japanese species. <em>Journal of the Faculty of Science, The University of Tokyo, Section 3, Botany </em>11: 93–136.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Fu, D.Z. & Zhu, G.H. (2001) <em>Thalictrum</em> Linnaeus. <em>In</em>: Wu, Z.Y., Raven, P.H. & Hong, D.Y. (Eds.) <em>Flora of China</em>, vol. 6. Science Press, Beijing & Missouri Botanical Garden Press, St. Louis, pp. 282–302.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Gregory, W.C. (1941) Phylogenetic and Cytological Studies in the Ranunculaceae Juss. <em>Transactions of the American Philosophical Society</em> 31: 443–521. https://doi.org/10.2307/1005611</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hooker, J.D. & Thomson, T. (1855) Ranunculaceae. <em>In</em>: Hooker, J.D. & Thomson, T. (Eds.) <em>Flora Indica</em>, vol. 1. W. Pamplin, London, pp. 1–60.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">IUCN Standards and Petitions Committee (2019) <em>Guidelines for Using the IUCN Red List Categories and Criteria</em>. Version 14. IUCN Standards and Petitions Committee. </span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Langlet, O (1927) Beiträge zur Zytologie der Ranunculazeen. <em>Svensk Botanisk Tidskrift</em> 21: 1–17.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lecoyer, J.C. (1885) Monographie du genre <em>Thalictrum</em>. <em>Bulletin de la Société Royale de Botanique de Belgique</em> 24: 78–324.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Li, X.W. & Li, J. (1993) A preliminary floristic study on the seed plants from the region of Hengduan Mountain. <em>Acta Botanica Yunnanica</em> 15: 217‒231.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Linnaeus, C. (1753) <em>Species Plantarum</em>. Laurentii Salvii, Holmiae, 1200 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Liu, Y., Ye, J.F., Hu, H.H., Peng, X.D., Zhao, L.N., Lu, L.M., Zaman, W. & Chen, Z.D. (2021) Influence of elevation on bioregionalisation: A case study of the Sino-Himalayan flora. <em>Journal of Biogeography</em> 48: 2578‒2587. https://doi.org/10.1111/jbi.14222</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Luo, D., Yue, J.P., Sun, W.G., Xu, B. Li, Z.M., Comes, H.P. & Sun, H. (2016) Evolutionary history of the subnival flora of the Himalaya-Hengduan Mountains: first insights from comparative phylogeography of four perennial herbs.<em> Journal of Biogeography</em> 43: 31‒43.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Maximowicz, C.J. (1877) Diagnoses plantarum novarum Asiaticarum. <em>Bulletin de l’Académie Impériale des Sciences de St-Pétersbourg </em>23: 305‒391. https://doi.org/10.5962/bhl.title.46308</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Myers, N., Mittermeier, R.A., Mittermeier, C.G., Fonseca, G.A.B. da & Kant, J. (2000) Biodiversity hotspots for conservation priorities. <em>Nature</em> 403: 853‒858. https://doi.org/10.1038/35002501</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rau, M.A. (1993) Ranunculaceae. <em>In</em>: Sharma, B.D., Balakrishnan, N.P., Rao, R.R. & Hajra, P.K. (Eds.) <em>Flora of India</em>, vol. 1. Botanical Survey of India, Calcutta, pp. 1–145.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tamura, M. (1995) <em>Thalictrum</em> L. <em>In</em>: Hiepko, P. (Ed.) <em>Die Natürlichen Pflanzenfamilien</em>, Band 17 a IV. Duncker und Humblot, Berlin, pp. 474–494.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wallich, N. (1831) <em>Plantae Asiaticae Rariores; or Descriptions and Figures of a Select Number of Unpublished East Indian Plants</em>, vol. 2. Treuttel and Würtz, London, 200 pp. https://doi.org/10.5962/bhl.title.468</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, W.T. (1982) <em>Thalictrum</em> L. <em>In</em>: Wang, W.T. (Ed.) <em>Iconographia Cormophytorum Sinicorum</em>, suppl. 1. Science Press, Beijing, pp. 410‒431.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, W.T. (1985) <em>Thalictrum</em> L. <em>In</em>: Wu, C.Y. (Ed.) <em>Flora Xizangica</em>, vol. 2. Science Press, Beijing, pp. 63‒76.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, W.T. (2000a) <em>Thalictrum</em> L.<em> In</em>: Wu, Z.Y. (Ed.) <em>Flora Yunnanica</em>, vol. 11. Science Press, Beijing, pp. 153–177.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, W.T. (2000b) <em>Thalictrum</em> L. <em>In</em>: Fu, L.K. & Hong, T. (Eds.) <em>Higher Plants of China</em>, vol. 3. Qingdao Publishing House, Qingdao, pp. 459‒481.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, W.T. (2018a) <em>Thalictrum (Ranunculaceae) in China</em>. Peking University Press, Beijing, 376 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, W.T. (2018b) <em>Thalictrum nepalense</em>, a new species of Ranunculaceae from Nepal. <em>Bulletin of Botanical Research</em> 38: 641‒643.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, W.T. (2020) Three new species of <em>Thalictrum</em> (Ranunculaceae) from Sichuan Province. <em>Bulletin of Botanical Research</em> <em>(Harbin)</em> 40: 5–9.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, W.T. & Wang, S.H. (1979a) <em>Thalictrum</em> L. <em>In</em>: Anonymous (Ed.) <em>Flora Reipublicae Popularis Sinicae</em>, vol. 27. Science Press, Beijing, pp. 502–592.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wang, W.T. & Wang, S.H. (1979b) Addenda. <em>In</em>: Anonymous (Ed.) <em>Flora Reipublicae Popularis Sinicae, </em>vol. 27. Science Press, Beijing, pp. 603–622.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Wu, Z.Y. (1988) Hengduan mountain flora and her significance. <em>The Journal of Japanese Botany</em> 63: 297‒311.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Xie, L., Wang, Z.T., Liu, Q., Li, S.M., Li, Y.S., Song, R.K., Zhao, S.M., Han, L.T., Wang, W., Yu, L., Qian, J.S., Yang, X.W., Zhang, Y.T., Qiao, X., Yang, X.B., Cong, X., Ma, T., Dong, S.Z., Wang, A.N., He, Z.Y. & Pei, J. (2016) <em>Thalictrum</em> L. <em>In</em>: Wei, F.N., Wang, Z.T., Zhang, W.S., Pei, J. & Yang, X.W. (Eds.) <em>Medical Flora of China</em>, vol. 3. Peking University Medical Press, Beijing, pp. 231‒287.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Yonekura, K. (2008) Ranunculaceae. <em>In</em>: Ohba, H., Iokawa, Y. & Sharma, L.R. (Eds.) <em>Flora of Mustang, Nepal</em>. Kodansha Scientific Ltd., Tokyo, pp. 61‒94.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Yuan, Q. & Yang, Q.E. (2017a) Neotypification of the name <em>Thalictrum umbricola</em> (Ranunculaceae). <em>Phytotaxa</em> 323: 199‒200. https://doi.org/10.11646/phytotaxa.323.2.10</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Yuan, Q. & Yang, Q.E. (2017b) Proposal to conserve the name <em>Thalictrum delavayi</em> against <em>T. thibeticum</em> (Ranunculaceae). <em>Taxon</em> 66: 761. https://doi.org/10.12705/663.29</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Yuan, Q. & Yang, Q.E. (2017c)<em> Thalictrum callianthum </em>(Ranunculaceae) is merged with <em>T. diffusiflorum</em>.<em> Phytotaxa </em>307: 285‒291. https://doi.org/10.11646/phytotaxa.307.4.6</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Yuan, Q., Ren, C. & Yang, Q.E. (2018a) <em>Thalictrum argyi</em> (Ranunculaceae) is conspecific with <em>T. fortunei</em>, not <em>T. javanicum</em>. <em>Phytotaxa</em> 346: 197‒200. https://doi.org/10.11646/phytotaxa.346.2.7</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Yuan, Q., Ren, C. & Yang, Q.E. (2018b) Lectotypification of the name <em>Thalictrum purdomii </em>(Ranunculaceae). <em>Phytotaxa</em> 340: 98‒100. https://doi.org/10.11646/phytotaxa.340.1.9</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Yuan, Q., Ren, C. & Yang, Q.E. (2018c) Clarification of the taxonomic identity of<em> Thalictrum neurocarpum</em> (Ranunculaceae), with the reduction of <em>T. austrotibeticum</em> to its synonymy. <em>Phytotaxa</em> 338: 195‒201. https://doi.org/10.11646/phytotaxa.338.2.4</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P. (2022) Taxonomic studies of the genus <em>Thalictrum</em> L. (Ranunculaceae) from China. Ph.D. thesis, University of Chinese Academy of Sciences. Beijing, 701 pp.</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2018) Reinstatement of the independent specific status of <em>Thalictrum hamatum</em> (Ranunculaceae), with <em>T. macrorhynchum</em> reduced to its synonymy. <em>Phytotaxa</em> 369: 278‒286. https://doi.org/10.11646/phytotaxa.369.4.5</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2020a) <em>Thalictrum punduanum</em> var. <em>hirtellum</em> (Ranunculaceae), another new synonym of <em>T. neurocarpum</em>. <em>Phytotaxa</em> 442: 89‒100. https://doi.org/10.11646/phytotaxa.442.2.4</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2020b). <em>Thalictrum semiscandens</em> (Ranunculaceae), almost totally overlooked since its publication, is merged with <em>T. trichopus</em>. <em>Phytotaxa</em> 451: 283‒293. https://doi.org/10.11646/phytotaxa.451.4.3</span></span></span></p>
<p align="justify"><span style="color: #000000;"> <span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2020c) <em>Thalictrum mairei</em> (Ranunculaceae) is synonymous with <em>T. foliolosum</em>, not <em>T. leuconotum</em>. <em>Phytotaxa</em> 452: 137‒154. https://doi.org/10.11646/phytotaxa.452.2.3</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2021a) <em>Thalictrum nanhuaense</em> (Ranunculaceae), a new species from central Yunnan, China. <em>Phytotaxa </em>479: 121‒128. https://doi.org/10.11646/phytotaxa.479.1.10</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2021b) <em>Thalictrum brevisericeum</em> var. <em>pentagynum</em> (Ranunculaceae), a new synonym of <em>T. uncinulatum</em> from China. <em>Phytotaxa</em> 497: 97‒112. https://doi.org/10.11646/phytotaxa.497.2.3</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2021c) <em>Thalictrum kangdingense</em> (Ranunculaceae), a new synonym of <em>T. megalostigma</em> from China. <em>Phytotaxa</em> 498: 87‒103. https://doi.org/10.11646/phytotaxa.498.2.2</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2021d) <em>Thalictrum minshanicum</em> and <em>T. pseudoramosum</em> (Ranunculaceae), two new species from China. <em>Phytotaxa</em> 502: 133‒148. https://doi.org/10.11646/phytotaxa.502.2.2</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2021e) <em>Thalictrum jilongense</em> and <em>T. tenuicaule</em> (Ranunculaceae), described from China, are both merged with <em>T. leuconotum</em>. <em>Phytotaxa</em> 509: 168‒184. https://doi.org/10.11646/phytotaxa.509.2.2</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2021f) <em>Thalictrum bouffordii</em> (Ranunculaceae), a new species from the Qionglai mountains region in western Sichuan, China. <em>Phytotaxa</em> 510: 18‒28. https://doi.org/10.11646/phytotaxa.510.1.2</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2021g) Clarification of some imperfectly understood characters in the Chinese species <em>Thalictrum przewalskii</em> (Ranunculaceae), with reduction of <em>T. lasiogynum</em> and <em>T. latistylum</em> to its synonymy. <em>Phytotaxa</em> 511: 187‒210. https://doi.org/10.11646/phytotaxa.511.3.1</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2021h) <em>Thalictrum panzhihuaense</em> (Ranunculaceae) is placed in synonymy with <em>T. glandulosissimum</em>, with reduction of <em>T. tsaii</em> to <em>T. glandulosissimum</em> var. <em>tsaii</em>. <em>Phytotaxa</em> 520: 1‒20. https://doi.org/10.11646/phytotaxa.520.1.1</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Fei, W.Q., Yuan, Q. & Yang, Q.E. (2021i) <em>Thalictrum changii</em> (Ranunculaceae), a new species from southern Xizang (Tibet), China. <em>Phytotaxa</em> 511: 248‒260. https://doi.org/10.11646/phytotaxa.511.3.4</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2022a). <em>Thalictrum spiristylum</em> (Ranunculaceae), described from northwestern Yunnan, China, is merged with <em>T. atriplex</em>. <em>Phytotaxa</em> 538: 1‒20. https://doi.org/10.11646/phytotaxa.538.1.1</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2022b). <em>Thalictrum daguanense</em> (Ranunculaceae) is merged with <em>T. omeiense</em>, a species distributed in Sichuan and Yunnan, China. <em>Phytotaxa</em> 539: 130‒146. https://doi.org/10.11646/phytotaxa.539.2.2</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2022c). <em>Thalictrum fusiforme</em> (Ranunculaceae), described from southeastern Xizang (Tibet), China, is merged with <em>T. rostellatum</em>, a Sino-Himalayan species. <em>Phytotaxa</em> 545: 234‒260. https://doi.org/10.11646/phytotaxa.545.3.1</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2022d) Lectotypification of <em>Thalictrum chayuense </em>(Ranunculaceae), with its reduction to the synonymy of <em>T. tsawarungense</em>, both described from Zayu in southeastern Xizang, China. <em>Phytotaxa</em> 548: 184‒198. https://doi.org/10.11646/phytotaxa.548.2.4</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2022e) <em>Thalictrum cuonaense</em> (Ranunculaceae) is merged with the Himalayan <em>T. reniforme</em>, with notes on its morphologically similar species. <em>Phytotaxa</em> 554: 221‒244. https://doi.org/10.11646/phytotaxa.554.3.2</span></span></span></p>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Zeng, Y.P., Yuan, Q. & Yang, Q.E. (2022f) Clarification of morphology and distribution of <em>Thalictrum fortunei</em> (Ranunculaceae) from China, with reduction of <em>T. fortunei </em>var. <em>bulbiliferum</em> to its synonymy on its morphologically similar species. <em>Phytotaxa</em> 561: 121‒137. https://doi.org/10.11646/phytotaxa.561.2.1</span></span></span></p>