Abstract
Argostemma tortilobum, a new species endemic to Vietnam, is described and illustrated. The species is morphologically most similar to A. sarmentosum, but differs in having erect stem without runners, densely pilose peduncle and twisted corolla lobes with ciliate margins.
References
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Beentje, H. (2016) The Kew Plant Glossary: an illustrated dictionary of plant terms, 2nd Edition. Royal Botanical Gardens, Kew, Richmand, 184 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Bremer, B. (1989) The genus Argostemma (Rubiaceae-Argostemmateae) in Borneo. Missouri Botanic Garden 76: 7–49. [https://www.jstor.org/stable/2399341]</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Chen, T., Zhu, H., Chen, J., Taylor, C.M., Ehrendorfer, F., Funston, A.M. & Puff, C. (2011) Rubiaceae. In: Wu, Z.Y., Raven, P.H. & Hong, D.Y. (eds.) Flora of China, Vol. 19. Science Press, Beijing and Missouri Botanical Garden, St. Louis, pp. 57–368.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Das, S.C. & Ranman, M.A. (2010) Notes on the Rubiaceae. 3. Five new records for Bangladesh. Bangladesh Journal of Botany 39 (2): 215–222.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">IUCN Standards and Petitions Committee. (2022) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15. Prepared by the Standards and Petitions Committee, 116 pp. [https://www.iucnredlist.org/documents/RedListGuidelines.pdf]</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Kress, W.J., Defilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003) A checklist of trees, shrubs, herbs, and climbers of Myanmar. Systematic Botany 29 (2): 462–463. https://doi.org/10.1600/036364404774195674</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lanorsavanh, S. & Souvannakhoummane, K. (2020) Six new species of Argostemma Wall. (Rubiaceae) from Laos. Nordic Journal of Botany 2020: e02714. https://doi.org/10.1111/njb.02714</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pitard, J. (1922-1924) RUBIACEES. In: Lecomte, H., Humbert, H. & Gagnepain, F. (eds.) Flore Génerale de l’Indo-Chine, Vol. 3, fasc 1–4. Masson, Paris, pp. 20–442.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ridley, H.N. (1923) LXXIV. RUBIACEAE. In: Ridley, H.N. (ed.) The Flora of the Malay Peninsula, Vol. 2. Baptist Mission Press, London, pp. 3–177.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Roxburgh, W. (1824) Flora Indica: or description of Indian plants, Vol. 2. Mission Press, Serampore, 588 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sridith, K. (1999) A synopsis of the genus Argostemma Wall. (Rubiaceae) in Thailand. Thai Forest Bulletin 27: 86–138.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sridith, K. (2007) Notes on the genus Argostemma (Rubiaceae) of the Malay Peninsula and Peninsula Thailand. Blumea 52: 367–377.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sridith, K. & Puff, C. (2000) Distribution of Argostemma Wall. (Rubiaceae), with special reference to Thailand and surrounding areas. Thai Forest Bulletin 28: 123–137.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Thiers, B. (2022) Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 29 September 2022)</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Vu, H.T., Nguyen, T.T., Nguyen, H.Q., Le, K.D. & Do, T.V. (2020) Notes on taxonomy of the genus Argostemma (Rubiaceae) from Vietnam. Thai Forest Bulletin 48: 234–238. https://doi.org/10.20531/tfb.2020.48.2.16</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Bremer, B. (1989) The genus Argostemma (Rubiaceae-Argostemmateae) in Borneo. Missouri Botanic Garden 76: 7–49. [https://www.jstor.org/stable/2399341]</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Chen, T., Zhu, H., Chen, J., Taylor, C.M., Ehrendorfer, F., Funston, A.M. & Puff, C. (2011) Rubiaceae. In: Wu, Z.Y., Raven, P.H. & Hong, D.Y. (eds.) Flora of China, Vol. 19. Science Press, Beijing and Missouri Botanical Garden, St. Louis, pp. 57–368.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Das, S.C. & Ranman, M.A. (2010) Notes on the Rubiaceae. 3. Five new records for Bangladesh. Bangladesh Journal of Botany 39 (2): 215–222.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">IUCN Standards and Petitions Committee. (2022) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15. Prepared by the Standards and Petitions Committee, 116 pp. [https://www.iucnredlist.org/documents/RedListGuidelines.pdf]</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Kress, W.J., Defilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003) A checklist of trees, shrubs, herbs, and climbers of Myanmar. Systematic Botany 29 (2): 462–463. https://doi.org/10.1600/036364404774195674</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lanorsavanh, S. & Souvannakhoummane, K. (2020) Six new species of Argostemma Wall. (Rubiaceae) from Laos. Nordic Journal of Botany 2020: e02714. https://doi.org/10.1111/njb.02714</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pitard, J. (1922-1924) RUBIACEES. In: Lecomte, H., Humbert, H. & Gagnepain, F. (eds.) Flore Génerale de l’Indo-Chine, Vol. 3, fasc 1–4. Masson, Paris, pp. 20–442.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Ridley, H.N. (1923) LXXIV. RUBIACEAE. In: Ridley, H.N. (ed.) The Flora of the Malay Peninsula, Vol. 2. Baptist Mission Press, London, pp. 3–177.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Roxburgh, W. (1824) Flora Indica: or description of Indian plants, Vol. 2. Mission Press, Serampore, 588 pp.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sridith, K. (1999) A synopsis of the genus Argostemma Wall. (Rubiaceae) in Thailand. Thai Forest Bulletin 27: 86–138.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sridith, K. (2007) Notes on the genus Argostemma (Rubiaceae) of the Malay Peninsula and Peninsula Thailand. Blumea 52: 367–377.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sridith, K. & Puff, C. (2000) Distribution of Argostemma Wall. (Rubiaceae), with special reference to Thailand and surrounding areas. Thai Forest Bulletin 28: 123–137.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Thiers, B. (2022) Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 29 September 2022)</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Vu, H.T., Nguyen, T.T., Nguyen, H.Q., Le, K.D. & Do, T.V. (2020) Notes on taxonomy of the genus Argostemma (Rubiaceae) from Vietnam. Thai Forest Bulletin 48: 234–238. https://doi.org/10.20531/tfb.2020.48.2.16</span></span></span>
