Abstract
Botanical excursions in the mountains of Bukidnon province, Mindanao resulted in the collection of two poorly known Philippine endemic Begonia species, B. malindangensis and B. mindanaensis. These two species are lectotypified and a comprehensive description, photographs, geographical distribution information and a preliminary IUCN conservation assessment are presented for each species.
References
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Amoroso, V.B., Obsioma, L.D., Arlalejo, J.B., Aspiras, R.A., Capili, D.P., Polizon, J.J.A., Sumile, E.B. (2009) Inventory and conservation of endangered, endemic and economically important flora of Hamiguitan Range, Southern Philippines. Blumea 54: 71–76. https://doi.org/10.3767/000651909X474113</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Amoroso, V.B. & Aspiras, R.A. (2011) Hamiguitan Range: a sanctuary for native flora. Saudi Journal Biological Sciences 18: 7–15. https://doi.org/10.1016/j.sjbs.2010.07.003</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Bachman, S., Moat, J., Hill, A., de la Torre, J. & Scott, B. (2011) Supporting Red List threat assessments with GeoCAT: Geospatial Conservation Assessment Tool. ZooKeys 150: 117–126. https://doi.org/10.3897/zookeys.150.2109</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Beentje, H.J. (2010) The Kew Plant Glossary: An Illustrated Dictionary of Plant Terms. Royal Botanic Gardens, Kew, Richmond, Surrey.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Buenavista, D.P., Ang, Y.P., Pranada, M.A.K., Salas, D.A., Mollee, E. & McDonald, M. (2021) Begonia bangsamoro (Begoniaceae, section Petermannia), a new species from Mindanao Island, the Philippines. Phytotaxa 497: 39–48. https://doi.org/10.11646/phytotaxa.497.1.4</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Coritico, F.P., Amoroso, V.B., Acma, F.M., Cariño, Y.L.L. & Fritsch, P.W. (2020a) Ferns and lycophytes of Mt. Tago Range, Bukidnon, Southern Philippines: species richness, distribution, and conservation status. Philippine Journal of Science 149 (3-a): 773–790.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">de Candolle, A.L.P.P. (1859) Mémoire sur la famille des Bégoniacées. Annales des Sciences Naturelles; Botanique Sér. 4 (11): 93–149.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hughes, M., Moonlight, P.W., Jara, A. & Pullan, M. (2015–Present) Begonia Resource Centre. Available from: http://padme.rbge.org.uk/begonia (accessed 5 September 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hughes, M., Peng, C.-I., Lin, C.-W., Rubite, R.R., Blanc, P. & Chung, K.-F. (2018) Chloroplast and nuclear DNA exchanges among Begonia sect. Baryandra species (Begoniaceae) from Palawan Island, Philippines, and descriptions of five new species. PLoS ONE 13: e0194877. https://doi.org/10.1371/journal.pone.0194877</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">IUCN Standards and Petitions Subcommittee. (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Available from: http://www.iucnredlist.org/documents/RedListGuidelines.pdf (accessed 15 December 2020).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Klotzsch, J.F. (1854) Begoniaceen-Gattungen und Arten. Monatsberichte der Königlich preussischen Akademie der Wissenschaften zu Berlin 1854: 119–128.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lillo, E.P., Fernando, E.S. & Lillo, M.J.R. (2019) Plant diversity and structure of forest habitat types on Dinagat Island, Philippines. Journal of Asia-Pacific Biodiversity 12: 83–105. https://doi.org/10.1016/j.japb.2018.07.003</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lin, C.-W., Chung, S.-W. & Peng, C.-I. (2017) Eleven new species of Begonia (Begoniaceae) from Sarawak, Borneo. Taiwania 62: 219–251. https://doi.org/10.6165/tai.2017.62.219 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Merrill, E.D. (1912 [“1911”]) The Philippine species of Begonia. Philippine Journal of Science 6: 396–406.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Naive, M.A.K., Alejandro, G.J.D. & Hughes, M. (2018) Taxonomic notes on the Philippine endemic Begonia colorata (Begoniaceae, section Petermannia). Phytotaxa 381: 127–131. https://doi.org/10.11646/phytotaxa.381.1.16</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Naive, M.A.K. & Cababan, M.A. (2021) Taxonomy of the genus Begonia (Begoniaceae) in Mindanao, Philippines I: Begonia olganunezae, a new Begonia section Petermannia species discovered in the province of Bukidnon. Taiwania 66: 433–438. https://doi.org/10.6165/tai.2021.66.433</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Naive, M.A.K., Mazo, K.R. & Hughes, M. (2022) Taxonomy of the genus Begonia (Begoniaceae) in Mindanao, Philippines II: Lectotypification of two Begonia section Petermannia species from the Zamboanga Peninsula. Kew Bulletin 77: 139–146. https://doi.org/10.1007/s12225-022-10008-5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nakamura, K., Rubite, R.R., Kono, Y., Callado, J.R.C. & Peng, C.-I. (2013) Begonia tandangii (Begoniaceae, section Baryandra), a new species from Luzon Island, the Philippines. Phytotaxa 145: 27–37. https://doi.org/10.11646/phytotaxa.145.1.3</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pasion, O.P., Barrias, C.D.P., Asuncion, M.P., Angadol, A.H., Pabiling, R.R., Pasion Jr., A., Braulio, A.A. & Baysa Jr., A.M. (2021) Assessing tree diversity and carbon density of a riparian zone within a protected area in southern Philippines. Journal of Asia-Pacific Biodiversity 14: 78–86. https://doi.org/10.1016/j.japb.2020.10.006</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pelser, P.B., Barcelona, J.F. & Nickrent, D.L. (Eds.) (2011) Co’s Digital Flora of the Philippines. Available from: www.philippineplants.org (accessed 22 January 2021).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Peng, C.-I., Rubite, R.R., Lin, C.-W., Hughes, M., Kono, Y. & Chung, K.-F. (2017) Three new species of Begonia sect. Baryandra from Panay Island, Philippines. Botanical Studies 58: 28. https://doi.org/10.1186/s40529-017-0182-x</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R. & Madulid, D.A. (2009) The rediscovery of Philippine Begonia. Blumea 54: 267–268. https://doi.org/10.3767/000651909X476256</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R., Hughes, M., Alejandro, G.J.D. & Peng, C.-I. (2013) Recircumscription of Begonia sect. Baryandra (Begoniaceae): evidence from molecular data. Botanical Studies 54: 38. https://doi.org/10.1186/1999-3110-54-38</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R. (2013) Begonia section Petermannia of Luzon Island, The Philippines. Philippine Journal of Science 142: 183–197.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R., Hughes, M., Blanc, P., Chung, K.-F., Ang, H.-A., Kono, Y., Alejandro, G.J.D., De Layola, L.B., Virata, A.G.N. & Peng, C.-I. (2015) Three new species of Begonia endemic to the Puerto Princesa Subterranean River National Park, Palawan. Botanical Studies 56: 19. https://doi.org/10.1186/s40529-015-0099-1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R., Brillantes, R.Y., Tandang, D.N., Moran, C.B., Rule, M.G. & Lin, C.-W. (2021) Begonoa benitotanii (section Petermannia, Begoniaceae), a new species endemic to the Philippine island of Bucas Grande. Phytotaxa 513: 257–264. https://doi.org/10.11646/phytotaxa.513.3.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rule, M.G.Q., Ang, Y.P., Rubite, R.R., Docot, R.V.A., Bustamante, R.A.A. & Robinson, A.S. (2020) Begonia makuruyot (Begoniaceae, section Baryandra), a new species from Surigao del Norte Province, Philippines. Phytotaxa 470: 226–234. https://doi.org/10.11646/phytotaxa.470.3.2</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sangvirotjanapat, S., Denduangboriphant, J. & Newman, M.F. (2019) A taxonomic revision of Globba sect. Nudae subsect. Mediocalcaratae (Zingiberaceae). Edinburgh Journal of Botany 77 (1): 1–63. https://doi.org/10.1017/S0960428619000258</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tandang, D.N., Rubite, R.R., Angeles Jr., R.T. & de Guzman, M.C. (2016) Begonia titoevangelistae (sect. Baryandra, Begoniaceae) a new species from Catanduanes Island, the Philippines. Phytotaxa 282: 273–281. https://doi.org/10.11646/phytotaxa.282.4.4</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Thiers, B.M. (2021) Index Herbariorum: a global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 20 December 2020)</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017). Regnum Vegetabile 159. Koeltz Botanical Books, Glashu?tten. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Warburg, O. (1904) Begoniaceae. In: Perkins, J.R. Fragmenta Florae Philippinae. Contributions to the Flora of the Philipines Islands. Borntraeger, Leipzig.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Amoroso, V.B. & Aspiras, R.A. (2011) Hamiguitan Range: a sanctuary for native flora. Saudi Journal Biological Sciences 18: 7–15. https://doi.org/10.1016/j.sjbs.2010.07.003</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Bachman, S., Moat, J., Hill, A., de la Torre, J. & Scott, B. (2011) Supporting Red List threat assessments with GeoCAT: Geospatial Conservation Assessment Tool. ZooKeys 150: 117–126. https://doi.org/10.3897/zookeys.150.2109</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Beentje, H.J. (2010) The Kew Plant Glossary: An Illustrated Dictionary of Plant Terms. Royal Botanic Gardens, Kew, Richmond, Surrey.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Buenavista, D.P., Ang, Y.P., Pranada, M.A.K., Salas, D.A., Mollee, E. & McDonald, M. (2021) Begonia bangsamoro (Begoniaceae, section Petermannia), a new species from Mindanao Island, the Philippines. Phytotaxa 497: 39–48. https://doi.org/10.11646/phytotaxa.497.1.4</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Coritico, F.P., Amoroso, V.B., Acma, F.M., Cariño, Y.L.L. & Fritsch, P.W. (2020a) Ferns and lycophytes of Mt. Tago Range, Bukidnon, Southern Philippines: species richness, distribution, and conservation status. Philippine Journal of Science 149 (3-a): 773–790.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">de Candolle, A.L.P.P. (1859) Mémoire sur la famille des Bégoniacées. Annales des Sciences Naturelles; Botanique Sér. 4 (11): 93–149.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hughes, M., Moonlight, P.W., Jara, A. & Pullan, M. (2015–Present) Begonia Resource Centre. Available from: http://padme.rbge.org.uk/begonia (accessed 5 September 2022).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Hughes, M., Peng, C.-I., Lin, C.-W., Rubite, R.R., Blanc, P. & Chung, K.-F. (2018) Chloroplast and nuclear DNA exchanges among Begonia sect. Baryandra species (Begoniaceae) from Palawan Island, Philippines, and descriptions of five new species. PLoS ONE 13: e0194877. https://doi.org/10.1371/journal.pone.0194877</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">IUCN Standards and Petitions Subcommittee. (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Available from: http://www.iucnredlist.org/documents/RedListGuidelines.pdf (accessed 15 December 2020).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Klotzsch, J.F. (1854) Begoniaceen-Gattungen und Arten. Monatsberichte der Königlich preussischen Akademie der Wissenschaften zu Berlin 1854: 119–128.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lillo, E.P., Fernando, E.S. & Lillo, M.J.R. (2019) Plant diversity and structure of forest habitat types on Dinagat Island, Philippines. Journal of Asia-Pacific Biodiversity 12: 83–105. https://doi.org/10.1016/j.japb.2018.07.003</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Lin, C.-W., Chung, S.-W. & Peng, C.-I. (2017) Eleven new species of Begonia (Begoniaceae) from Sarawak, Borneo. Taiwania 62: 219–251. https://doi.org/10.6165/tai.2017.62.219 </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Merrill, E.D. (1912 [“1911”]) The Philippine species of Begonia. Philippine Journal of Science 6: 396–406.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Naive, M.A.K., Alejandro, G.J.D. & Hughes, M. (2018) Taxonomic notes on the Philippine endemic Begonia colorata (Begoniaceae, section Petermannia). Phytotaxa 381: 127–131. https://doi.org/10.11646/phytotaxa.381.1.16</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Naive, M.A.K. & Cababan, M.A. (2021) Taxonomy of the genus Begonia (Begoniaceae) in Mindanao, Philippines I: Begonia olganunezae, a new Begonia section Petermannia species discovered in the province of Bukidnon. Taiwania 66: 433–438. https://doi.org/10.6165/tai.2021.66.433</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Naive, M.A.K., Mazo, K.R. & Hughes, M. (2022) Taxonomy of the genus Begonia (Begoniaceae) in Mindanao, Philippines II: Lectotypification of two Begonia section Petermannia species from the Zamboanga Peninsula. Kew Bulletin 77: 139–146. https://doi.org/10.1007/s12225-022-10008-5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Nakamura, K., Rubite, R.R., Kono, Y., Callado, J.R.C. & Peng, C.-I. (2013) Begonia tandangii (Begoniaceae, section Baryandra), a new species from Luzon Island, the Philippines. Phytotaxa 145: 27–37. https://doi.org/10.11646/phytotaxa.145.1.3</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pasion, O.P., Barrias, C.D.P., Asuncion, M.P., Angadol, A.H., Pabiling, R.R., Pasion Jr., A., Braulio, A.A. & Baysa Jr., A.M. (2021) Assessing tree diversity and carbon density of a riparian zone within a protected area in southern Philippines. Journal of Asia-Pacific Biodiversity 14: 78–86. https://doi.org/10.1016/j.japb.2020.10.006</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Pelser, P.B., Barcelona, J.F. & Nickrent, D.L. (Eds.) (2011) Co’s Digital Flora of the Philippines. Available from: www.philippineplants.org (accessed 22 January 2021).</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Peng, C.-I., Rubite, R.R., Lin, C.-W., Hughes, M., Kono, Y. & Chung, K.-F. (2017) Three new species of Begonia sect. Baryandra from Panay Island, Philippines. Botanical Studies 58: 28. https://doi.org/10.1186/s40529-017-0182-x</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R. & Madulid, D.A. (2009) The rediscovery of Philippine Begonia. Blumea 54: 267–268. https://doi.org/10.3767/000651909X476256</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R., Hughes, M., Alejandro, G.J.D. & Peng, C.-I. (2013) Recircumscription of Begonia sect. Baryandra (Begoniaceae): evidence from molecular data. Botanical Studies 54: 38. https://doi.org/10.1186/1999-3110-54-38</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R. (2013) Begonia section Petermannia of Luzon Island, The Philippines. Philippine Journal of Science 142: 183–197.</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R., Hughes, M., Blanc, P., Chung, K.-F., Ang, H.-A., Kono, Y., Alejandro, G.J.D., De Layola, L.B., Virata, A.G.N. & Peng, C.-I. (2015) Three new species of Begonia endemic to the Puerto Princesa Subterranean River National Park, Palawan. Botanical Studies 56: 19. https://doi.org/10.1186/s40529-015-0099-1</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rubite, R.R., Brillantes, R.Y., Tandang, D.N., Moran, C.B., Rule, M.G. & Lin, C.-W. (2021) Begonoa benitotanii (section Petermannia, Begoniaceae), a new species endemic to the Philippine island of Bucas Grande. Phytotaxa 513: 257–264. https://doi.org/10.11646/phytotaxa.513.3.5</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Rule, M.G.Q., Ang, Y.P., Rubite, R.R., Docot, R.V.A., Bustamante, R.A.A. & Robinson, A.S. (2020) Begonia makuruyot (Begoniaceae, section Baryandra), a new species from Surigao del Norte Province, Philippines. Phytotaxa 470: 226–234. https://doi.org/10.11646/phytotaxa.470.3.2</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Sangvirotjanapat, S., Denduangboriphant, J. & Newman, M.F. (2019) A taxonomic revision of Globba sect. Nudae subsect. Mediocalcaratae (Zingiberaceae). Edinburgh Journal of Botany 77 (1): 1–63. https://doi.org/10.1017/S0960428619000258</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Tandang, D.N., Rubite, R.R., Angeles Jr., R.T. & de Guzman, M.C. (2016) Begonia titoevangelistae (sect. Baryandra, Begoniaceae) a new species from Catanduanes Island, the Philippines. Phytotaxa 282: 273–281. https://doi.org/10.11646/phytotaxa.282.4.4</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Thiers, B.M. (2021) Index Herbariorum: a global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available from: http://sweetgum.nybg.org/science/ih/ (accessed 20 December 2020)</span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Turland, N.J., Wiersema, J.H., Barrie, F.R., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Kusber, W.-H., Li, D.-Z., Marhold, K., May, T.W., McNeill, J., Monro, A.M., Prado, J., Price, M.J. & Smith, G.F. (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017). Regnum Vegetabile 159. Koeltz Botanical Books, Glashu?tten. </span></span></span>
<p align="justify"><span style="color: #000000;"><span style="font-family: Times New Roman, serif;"><span style="font-size: small;">Warburg, O. (1904) Begoniaceae. In: Perkins, J.R. Fragmenta Florae Philippinae. Contributions to the Flora of the Philipines Islands. Borntraeger, Leipzig.</span></span></span>