Abstract
Lophiostomataceae species commonly occur as saprobes on twigs, stems or bark on diverse woody and herbaceous plants in aquatic and terrestrial habitats. In the present study, we introduce Lophiostoma khanzada-kirgizbaeva sp. nov. and Paucispora xishanensis sp. nov. as new species isolated from woody substrates from Uzbekistan and China, respectively. Species delimitation was conducted using morphological and molecular phylogenetic approaches. Maximum likelihood and Bayesian inference analyses of a combined SSU, ITS, LSU, tef1-α and rpb2 sequence matrix revealed that L. khanzada-kirgizbaev and P. xishanensis are distinct from other Lophiostoma and Paucispora species in Lophiostomataceae, respectively.
References
<p>Andreasen, M., Skrede, I., Jaklitsch, W.M., Voglmayr, H. & Nordén, B. (2021) Multi-locus phylogenetic analysis of lophiostomatoid fungi motivates a broad concept of <em>Lophiostoma</em> and reveals nine new species. <em>Persoonia </em>46: 240–271. https://doi.org/10.3767/persoonia.2021.46.09</p>
<p>Capella-gutiérrez, S., Silla-martínez, J.M. & Gabaldón, T. (2009) trimAl : a tool for automated alignment trimming in large-scale phylogenetic analyses. <em>Bioinformatics</em> 25: 1972–1973. https://doi.org/https://doi.org/10.1093/bioinformatics/btp348</p>
<p>Crous, P.W., Carnegie, A.J., Wingfield, M.J., Sharma, R., Mughini, G., Noordeloos, M.E., Santini, A., Shouche, Y.S., Bezerra, J.D.P., Dima, B., Guarnaccia, V., Imrefi, I., Jurjević, Ž., Knapp, D.G., Kovács, G.M., Magistà, D., Perrone, G., Rämä, T., Rebriev, Y.A., Shivas, R.G., Singh, S.M., Souza-Motta, C.M., Thangavel, R., Adhapure, N.N., Alexandrova, A.V., Alfenas, A.C., Alfenas, R.F., Alvarado, P., Alves, A.L., Andrade, D.A., Andrade, J.P., Barbosa, R.N., Barili, A., Barnes, C.W., Baseia, I.G., Bellanger, J.-M., Berlanas, C., Bessette, A.E., Bessette, A.R., Biketova, A.Y., Bomfim, F.S., Brandrud, T.E., Bransgrove, K., Brito, A.C.Q., Cano-Lira, J.F., Cantillo, T., Cavalcanti, A.D., Cheewangkoon, R., Chikowski, R.S., Conforto, C., Cordeiro, T.R.L., Craine, J.D., Cruz, R., Damm, U., de Oliveira, R.J.V., de Souza, J.T., de Souza, H.G., Dearnaley, J.D.W., Dimitrov, R.A., Dovana, F., Erhard, A., Esteve-Raventós, F., Félix, C.R., Ferisin, G., Fernandes, R.A., Ferreira, R.J., Ferro, L.O., Figueiredo, C.N., Frank, J.L., Freire, K.T.L.S., García, D., Gené, J., Gęsiorska, A., Gibertoni, T.B., Gondra, R.A.G., Gouliamova, D.E., Gramaje, D., Guard, F., Gusmão, L.F.P., Haitook, S., Hirooka, Y., Houbraken, J., Hubka, V., Inamdar, A., Iturriaga, T., Iturrieta-González, I., Jadan, M., Jiang, N., Justo, A., Kachalkin, A.V., Kapitonov, V.I., Karadelev, M., Karakehian, J., Kasuya, T., Kautmanová, I., Kruse, J., Kušan, I., Kuznetsova, T.A., Landell, M.F., Larsson, K.-H., Lee, H.B., Lima, D.X., Lira, C.R.S., Machado, A.R., Madrid, H., Magalhães, O.M.C., Majerova, H., Malysheva, E.F., Mapperson, R.R., Marbach, P.A.S., Martín, M.P., Martín-Sanz, A., Matočec, N., McTaggart, A.R., Mello, J.F., Melo, R.F.R., Mešič, A., Michereff, S.J., Miller, A.N., Minoshima, A., Molinero-Ruiz, L., Morozova, O.V., Mosoh, D., Nabe, M., Naik, R., Nara, K., Nascimento, S.S., Neves, R.P., Olariaga, I., Oliveira, R.L., Oliveira, T.G.L., Ono, T., Ordoñez, M.E., de M. Ottoni, A., Paiva, L.M., Pancorbo, F., Pant, B., Pawłowska, J., Peterson, S.W., Raudabaugh, D.B., Rodríguez-Andrade, E., Rubio, E., Rusevska, K., Santiago, A.L.C.M.A., Santos, A.C.S., Santos, C., Sazanova, N.A., Shah, S., Sharma, J., Silva, B.D.B., Siquier, J.L., Sonawane, M.S., Stchigel, A.M., Svetasheva, T., Tamakeaw, N., Telleria, M.T., Tiago, P.V., Tian, C.M., Tkalčec, Z., Tomashevskaya, M.A., Truong, H.H., Vecherskii, M.V., Visagie, C.M., Vizzini, A., Yilmaz, N., Zmitrovich, I.V., Zvyagina, E.A., Boekhout, T., Kehlet, T., Lżss¸e, T. & Groenewald, J.Z. (2019) Fungal Planet description sheets: 868–950. <em>Persoonia</em> 42: 291–473. https://doi.org/10.3767/persoonia.2019.42.11</p>
<p>Edgar, R.C. (2004) MUSCLE: multiple sequence alignment with high accuracy and high throughput. <em>Nucleic Acids Research</em> 32: 1792–1797. https://doi.org/10.1093/nar/gkh340</p>
<p>Hall, T.A. (1999) BioEdit: A user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. <em>Nucleic Acids Symposium Series</em> 41: 95–98.</p>
<p>Hashimoto, A., Hirayama, K., Takahashi, H., Matsumura, M., Okada, G., Chen, C.Y., Huang, J.W., Kakishima, M., Ono, T. & Tanaka, K. (2018) Resolving the <em>Lophiostoma bipolare</em> complex: Generic delimitations within Lophiostomataceae. <em>Studies in Mycology</em> 90: 161–189. https://doi.org/10.1016/j.simyco.2018.03.001</p>
<p>Hawksworth, D.L. & Lücking, R. (2017) Fungal diversity revisited: 2.2 to 3.8 million species. <em>The Fungal Kingdom</em>: 79–95. https://doi.org/10.1128/9781555819583.ch4</p>
<p>Hirayama, K., Hashimoto, A. & Tanaka, K. (2014) A new species, <em>Lophiostoma versicolor</em>, from Japan (Pleosporales, Dothideomycetes). <em>Mycosphere</em> 5: 411–417. https://doi.org/10.5943/mycosphere/5/3/3</p>
<p>Hirayama, K. & Tanaka, K. (2011) Taxonomic revision of <em>Lophiostoma</em> and <em>Lophiotrema</em> based on reevaluation of morphological characters and molecular analyses. <em>Mycoscience</em> 52: 401–412. https://doi.org/10.1007/S10267-011-0126-3</p>
<p>Hongsanan, S., Hyde, K.D., Phookamsak, R., Wanasinghe, D.N., McKenzie, E.H.C., Sarma, V.V., Boonmee, S., Lücking, R., Bhat, D.J., Liu, N.G., Tennakoon, D.S., Pem, D., Karunarathna, A., Jiang, S.H., Jones, E.B.G., Phillips, A.J.L., Manawasinghe, I.S., Tibpromma, S., Jayasiri, S.C., Sandamali, D.S., Jayawardena, R.S., Wijayawardene, N.N., Ekanayaka, A.H., Jeewon, R., Lu, Y.Z., Dissanayake, A.J., Zeng, X.Y., Luo, Z.L., Tian, Q., Phukhamsakda, C., Thambugala, K.M., Dai, D.Q., Chethana, K.W.T., Samarakoon, M.C., Ertz, D., Bao, D.F., Doilom, M., Liu, J.K., PérezOrtega, S., Suija, A., Senwanna, C., Wijesinghe, S.N., Konta, S., Niranjan, M., Zhang, S.N., Ariyawansa, H.A., Jiang, H.B., Zhang, J.F., Norphanphoun, C., de Silva, N.I., Thiyagaraja, V., Zhang, H., Bezerra, J.D.P., Miranda-González, R., Aptroot, A., Kashiwadani, H., Harishchandra, D., Sérusiaux, E., Aluthmuhandiram, J.V.S., Abeywickrama, P.D., Devadatha, B., Wu, H.X., Moon, K.H., Gueidan, C., Schumm, F., Bundhun, D., Mapook, A., Monkai, J., Chomnunti, P., Suetrong, S., Chaiwan, N., Dayarathne, M., Yang, J., Rathnayaka, A.R., Bhunjun, C.S., Xu, J.C., Zheng, J.S., Liu, G., Feng, Y. & Xie, N. (2020) Refined families of Dothideomycetes: Dothideomycetidae and Pleosporomycetidae. <em>Mycosphere</em> 11: 1553–2107. https://doi.org/10.5943/mycosphere/11/1/13</p>
<p>Hyde, K.D., Jeewon, R., Chen, Y.-J., Bhunjun, C.S., Calabon, M.S., Jiang, H.-B., Lin, C.-G., Norphanphoun, C., Sysouphanthong, P., Pem, D., Tibpromma, S., Zhang, Q., Doilom, M., Jayawardena, R.S., Liu, J.-K., Maharachchikumbura, S.S.N., Phukhamsakda, C., Phookamsak, R., Al-Sadi, A.M., Thongklang, N., Wang, Y., Gafforov, Y., Gareth Jones, E.B. & Lumyong, S. (2020) The numbers of fungi: is the descriptive curve flattening? <em>Fungal Diversity</em> 103: 219–271. https://doi.org/10.1007/s13225-020-00458-2</p>
<p>Hyde, K.D., Jones, E.B.G., Liu, J.K., Ariyawansa, H., Boehm, E., Boonmee, S., Braun, U., Chomnunti, P., Crous, P.W., Dai, D.Q., Diederich, P., Dissanayake, A., Doilom, M., Doveri, F., Hongsanan, S., Jayawardena, R., Lawrey, J.D., Li, Y.M., Liu, Y.X., Lücking, R., Monkai, J., Muggia, L., Nelsen, M.P., Pang, K.L., Phookamsak, R., Senanayake, I.C., Shearer, C.A., Suetrong, S., Tanaka, K., Thambugala, K.M., Wijayawardene, N.N., Wikee, S., Wu, H.X., Zhang, Y., Aguirre-Hudson, B., Alias, S.A., Aptroot, A., Bahkali, A.H., Bezerra, J.L., Bhat, D.J., Camporesi, E., Chukeatirote, E., Gueidan, C., Hawksworth, D.L., Hirayama, K., De Hoog, S., Kang, J.C., Knudsen, K., Li, W.J., Li, X.H., Liu, Z.Y., Mapook, A., McKenzie, E.H.C., Miller, A.N., Mortimer, P.E., Phillips, A.J.L., Raja, H.A., Scheuer, C., Schumm, F., Taylor, J.E., Tian, Q., Tibpromma, S., Wanasinghe, D.N., Wang, Y., Xu, J.C., Yacharoen, S., Yan, J.Y. & Zhang, M. (2013) Families of Dothideomycetes. <em>Fungal Diversity</em> 63: 1–313. https://doi.org/10.1007/s13225-013-0263-4</p>
<p>Hyde, K.D., Norphanphoun, C., Abreu, V.P., Bazzicalupo, A., Thilini Chethana, K.W., Clericuzio, M., Dayarathne, M.C., Dissanayake, A.J., Ekanayaka, A.H., He, M.-Q., Hongsanan, S., Huang, S.-K., Jayasiri, S.C., Jayawardena, R.S., Karunarathna, A., Konta, S., Kušan, I., Lee, H., Li, J., Lin, C.-G., Liu, N.-G., Lu, Y.-Z., Luo, Z.-L., Manawasinghe, I.S., Mapook, A., Perera, R.H., Phookamsak, R., Phukhamsakda, C., Siedlecki, I., Soares, A.M., Tennakoon, D.S., Tian, Q., Tibpromma, S., Wanasinghe, D.N., Xiao, Y.-P., Yang, J., Zeng, X.-Y., Abdel-Aziz, F.A., Li, W.-J., Senanayake, I.C., Shang, Q.-J., Daranagama, D.A., de Silva, N.I., Thambugala, K.M., Abdel-Wahab, M.A., Bahkali, A.H., Berbee, M.L., Boonmee, S., Bhat, D.J., Bulgakov, T.S., Buyck, B., Camporesi, E., Castañeda-Ruiz, R.F., Chomnunti, P., Doilom, M., Dovana, F., Gibertoni, T.B., Jadan, M., Jeewon, R., Jones, E.B.G., Kang, J.-C., Karunarathna, S.C., Lim, Y.W., Liu, J.-K., Liu, Z.-Y., Plautz, H.L., Lumyong, S., Maharachchikumbura, S.S.N., Matočec, N., McKenzie, E.H.C., Mešić, A., Miller, D., Pawłowska, J., Pereira, O.L., Promputtha, I., Romero, A.I., Ryvarden, L., Su, H.-Y., Suetrong, S., Tkalčec, Z., Vizzini, A., Wen, T.-C., Wisitrassameewong, K., Wrzosek, M., Xu, J.-C., Zhao, Q., Zhao, R.-L. & Mortimer, P.E. (2017) Fungal diversity notes 603–708: taxonomic and phylogenetic notes on genera and species. <em>Fungal Diversity</em> 87: 1–235. https://doi.org/10.1007/s13225-017-0391-3</p>
<p>Jayasiri, S.C., Hyde, K.D., Ariyawansa, H.A., Bhat, J., Buyck, B., Cai, L., Dai, Y.-C., Abd-Elsalam, K.A., Ertz, D., Hidayat, I., Jeewon, R., Jones, E.B.G., Bahkali, A.H., Karunarathna, S.C., Liu, J.-K., Luangsa-ard, J.J., Lumbsch, H.T., Maharachchikumbura, S.S.N., McKenzie, E.H.C., Moncalvo, J.-M., Ghobad-Nejhad, M., Nilsson, H., Pang, K.-L., Pereira, O.L., Phillips, A.J.L., Raspé, O., Rollins, A.W., Romero, A.I., Etayo, J., Selçuk, F., Stephenson, S.L., Suetrong, S., Taylor, J.E., Tsui, C.K.M., Vizzini, A., Abdel-Wahab, M.A., Wen, T.-C., Boonmee, S., Dai, D.Q., Daranagama, D.A., Dissanayake, A.J., Ekanayaka, A.H., Fryar, S.C., Hongsanan, S., Jayawardena, R.S., Li, W.-J., Perera, R.H., Phookamsak, R., de Silva, N.I., Thambugala, K.M., Tian, Q., Wijayawardene, N.N., Zhao, R.-L., Zhao, Q., Kang, J.-C. & Promputtha, I. (2015) The Faces of Fungi database: fungal names linked with morphology, phylogeny and human impacts. <em>Fungal Diversity</em> 74: 3–18. https://doi.org/10.1007/s13225-015-0351-8</p>
<p>Liu, Y.J., Whelen, S. & Hall, B.D. (1999) Phylogenetic relationships among ascomycetes: evidence from an RNA polymerse II subunit. <em>Molecular Biology and Evolution</em> 16: 1799–1808. https://doi.org/10.1093/oxfordjournals.molbev.a026092</p>
<p>Mapook, A., Hyde, K.D., McKenzie, E.H.C., Jones, E.B.G., Bhat, D.J., Jeewon, R., Stadler, M., Samarakoon, M.C., Malaithong, M., Tanunchai, B., Buscot, F., Wubet, T. & Purahong, W. (2020) Taxonomic and phylogenetic contributions to fungi associated with the invasive weed <em>Chromolaena odorata</em> (Siam weed). <em>Fungal Diversity</em> 101: 1–175. https://doi.org/10.1007/s13225-020-00444-8</p>
<p>Miller, M.A., Pfeiffer, W. & Schwartz, T. (2010) Creating the CIPRES Science Gateway for inference of large phylogenetic trees. <em>In:</em> <em>2010 Gateway Computing Environments Workshop (GCE)</em>. IEEE, pp. 1–8.</p>
<p>Minh, B.Q., Schmidt, H.A., Chernomor, O., Schrempf, D., Woodhams, M.D., von Haeseler, A. & Lanfear, R. (2020) IQ-TREE 2: New Models and Efficient Methods for Phylogenetic Inference in the Genomic Era. <em>Molecular Biology and Evolution</em> 37: 1530–1534. https://doi.org/10.1093/molbev/msaa015</p>
<p>Nguyen, L.-T., Schmidt, H.A., von Haeseler, A. & Minh, B.Q. (2015) IQ-TREE: A Fast and Effective Stochastic Algorithm for Estimating Maximum-Likelihood Phylogenies. <em>Molecular Biology and Evolution</em> 32: 268–274. https://doi.org/10.1093/molbev/msu300</p>
<p>Rambaut, A. (2012) FigTree version 1.4.0. Available from: http://tree.bio.ed.ac.uk/software/figtree/ (accessed 24 August 2022)</p>
<p>Rannala, B. & Yang, Z. (1996) Probability distribution of molecular evolutionary trees: A new method of phylogenetic inference. <em>Journal of Molecular Evolution</em> 43: 304–311. https://doi.org/10.1007/BF02338839</p>
<p>Rehner, S.A. & Buckley, E. (2005) A Beauveria phylogeny inferred from nuclear ITS and EF1-α sequences: evidence for cryptic diversification and links to <em>Cordyceps</em> teleomorphs. <em>Mycologia</em> 97: 84–98. https://doi.org/10.1080/15572536.2006.11832842</p>
<p>Rehner, S.A. & Samuels, G.J. (1994) Taxonomy and phylogeny of <em>Gliocladium</em> analysed from nuclear large subunit ribosomal DNA sequences. <em>Mycological Research</em> 98: 625–634. https://doi.org/10.1016/S0953-7562(09)80409-7</p>
<p>Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: Efficient Bayesian Phylogenetic Inference and Model Choice Across a Large Model Space. <em>Systematic Biology</em> 61: 539–542. https://doi.org/10.1093/sysbio/sys029</p>
<p>Sung, G.-H., Sung, J.-M., Hywel-Jones, N.L. & Spatafora, J.W. (2007) A multi-gene phylogeny of Clavicipitaceae (Ascomycota, Fungi): Identification of localized incongruence using a combinational bootstrap approach. <em>Molecular Phylogenetics and Evolution</em> 44: 1204–1223. https://doi.org/10.1016/j.ympev.2007.03.011</p>
<p>Swofford, D.L. (2002) PAUP: Phylogenetic analysis using parsimony. <em>Options</em> 42: 294–307. https://doi.org/10.1007/BF02198856</p>
<p>Tanaka, K. & Harada, Y. (2003) Pleosporales in Japan (1): the genus <em>Lophiostoma</em>. <em>Mycoscience</em> 44: 85–96. https://doi.org/10.1007/S10267-002-0085-9</p>
<p>Tennakoon, D.S., Kuo, C.H., Jeewon, R., Thambugala, K.M. & Hyde, K.D. (2018) Saprobic lophiostomataceae (Dothideomycetes): <em>Pseudolophiostoma mangiferae</em> sp. nov. and <em>Neovaginatispora fuckelii</em>, a new record from <em>Mangifera indica</em>. <em>Phytotaxa</em> 364: 157–171. https://doi.org/10.11646/phytotaxa.364.2.3</p>
<p>Thambugala, K.M., Hyde, K.D., Tanaka, K., Tian, Q., Wanasinghe, D.N., Ariyawansa, H.A., Jayasiri, S.C., Boonmee, S., Camporesi, E., Hashimoto, A., Hirayama, K., Schumacher, R.K., Promputtha, I. & Liu, Z.Y. (2015) Towards a natural classification and backbone tree for Lophiostomataceae, Floricolaceae, and Amorosiaceae fam. nov. <em>Fungal Diversity</em> 74: 199–266. https://doi.org/10.1007/s13225-015-0348-3</p>
<p>Vilgalys, R. & Hester, M. (1990) Rapid genetic identification and mapping of enzymatically amplified ribosomal DNA from several <em>Cryptococcus</em> species. <em>Journal of Bacteriology</em> 172: 4238–4246. https://doi.org/10.1128/jb.172.8.4238-4246.1990</p>
<p>Wanasinghe, D.N., Phukhamsakda, C., Hyde, K.D., Jeewon, R., Lee, H.B., Gareth Jones, E.B., Tibpromma, S., Tennakoon, D.S., Dissanayake, A.J., Jayasiri, S.C., Gafforov, Y., Camporesi, E., Bulgakov, T.S., Ekanayake, A.H., Perera, R.H., Samarakoon, M.C., Goonasekara, I.D., Mapook, A., Li, W.-J., Senanayake, I.C., Li, J., Norphanphoun, C., Doilom, M., Bahkali, A.H., Xu, J., Mortimer, P.E., Tibell, L., Tibell, S. & Karunarathna, S.C. (2018) Fungal diversity notes 709–839: taxonomic and phylogenetic contributions to fungal taxa with an emphasis on fungi on <em>Rosaceae</em>. <em>Fungal Diversity</em> 89: 1–236. https://doi.org/10.1007/s13225-018-0395-7</p>
<p>White, T.J., Bruns, T., Lee, S. & Taylor, J. (1990) Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. <em>PCR Protocols and Applications</em> 18: 315–322. https://doi.org/10.1016/B978-0-12-372180-8.50042-1</p>
<p>Wijayawardene, N.N., Hyde, K.D., Al-Ani, L.K.T., Tedersoo, L., Haelewaters, D., Rajeshkumar, K.C., Zhao, R.L., Aptroot, A., Leontyev, D.V., Saxena, R.K., Tokarev, Y.S., Dai, D.Q., Letcher, P.M., Stephenson, S.L., Ertz, D., Lumbsch, H.T., Kukwa, M., Issi, I.V., Madrid, H., Phillips, A.J.L., Selbmann, L., Pfliegler, W.P., Horváth, E., Bensch, K., Kirk, P.M., Kolaríková, K., Raja, H.A., Radek, R., Papp, V., Dima, B., Ma, J., Malosso, E., Takamatsu, S., Rambold, G., Gannibal, P.B., Triebel, D., Gautam, A.K., Avasthi, S., Suetrong, S., Timdal, E., Fryar, S.C., Delgado, G., Réblová, M., Doilom, M., Dolatabadi, S., Pawlowska, J.Z., Humber, R.A., Kodsueb, R., Sánchez-Castro, I., Goto, B.T., Silva, D.K.A., de Souza, F.A., Oehl, F., da Silva, G.A., Silva, I.R., Blaszkowski, J., Jobim, K., Maia, L.C., Barbosa, F.R., Fiuza, P.O., Divakar, P.K., Shenoy, B.D., Castañeda-Ruiz, R.F., Somrithipol, S., Lateef, A.A., Karunarathna, S.C., Tibpromma, S., Mortimer, P.E., Wanasinghe, D.N., Phookamsak, R., Xu, J., Wang, Y., Tian, F., Alvarado, P., Li, D.W., Kušan, I., Matocec, N., Mešic, A., Tkalcec, Z., Maharachchikumbura, S.S.N., Papizadeh, M., Heredia, G., Wartchow, F., Bakhshi, M., Boehm, E., Youssef, N., Hustad, V.P., Lawrey, J.D., Santiago, A.L.C.M.A., Bezerra, J.D.P., Souza-Motta, C.M., Firmino, A.L., Tian, Q., Houbraken, J., Hongsanan, S., Tanaka, K., Dissanayake, A.J., Monteiro, J.S., Grossart, H.P., Suija, A., Weerakoon, G., Etayo, J., Tsurykau, A., Vázquez, V., Mungai, P., Damm, U., Li, Q.R., Zhang, H., Boonmee, S., Lu, Y.Z., Becerra, A.G., Kendrick, B., Brearley, F.Q., Motiejunaite, J., Sharma, B., Khare, R., Gaikwad, S., Wijesundara, D.S.A., Tang, L.Z., He, M.Q., Flakus, A., Rodriguez-Flakus, P., Zhurbenko, M.P., McKenzie, E.H.C., Stadler, M., Bhat, D.J., Liu, J.K., Raza, M., Jeewon, R., Nassonova, E.S., Prieto, M., Jayalal, R.G.U., Erdogdu, M., Yurkov, A., Schnittler, M., Shchepin, O.N., Novozhilov, Y.K., Silva-Filho, A.G.S., Gentekaki, E., Liu, P., Cavender, J.C., Kang, Y., Mohammad, S., Zhang, L.F., Xu, R.F., Li, Y.M., Dayarathne, M.C., Ekanayaka, A.H., Wen, T.C., Deng, C.Y., Pereira, O.L., Navathe, S., Hawksworth, D.L., Fan, X.L., Dissanayake, L.S., Kuhnert, E., Grossart, H.P. & Thines, M. (2020) Outline of Fungi and fungus-like taxa. <em>Mycosphere</em> 11: 1060–1456. https://doi.org/10.5943/mycosphere/11/1/8</p>
<p>Zhang, Y., Crous, P.W., Schoch, C.L. & Hyde, K.D. (2012) Pleosporales. <em>Fungal Diversity</em> 53: 1–221. https://doi.org/10.1007/s13225-011-0117-x</p>
<p>Zhang, Y., Wang, H.K., Fournier, J., Crous, P.W., Jeewon, R., Pointing, S.B. & Hyde, K.D. (2009) Towards a phylogenetic clarification of <em>Lophiostoma/Massarina</em> and morphologically similar genera in the Pleosporales. <em>Fungal Diversity</em> 38: 225–251.</p>
<p>Capella-gutiérrez, S., Silla-martínez, J.M. & Gabaldón, T. (2009) trimAl : a tool for automated alignment trimming in large-scale phylogenetic analyses. <em>Bioinformatics</em> 25: 1972–1973. https://doi.org/https://doi.org/10.1093/bioinformatics/btp348</p>
<p>Crous, P.W., Carnegie, A.J., Wingfield, M.J., Sharma, R., Mughini, G., Noordeloos, M.E., Santini, A., Shouche, Y.S., Bezerra, J.D.P., Dima, B., Guarnaccia, V., Imrefi, I., Jurjević, Ž., Knapp, D.G., Kovács, G.M., Magistà, D., Perrone, G., Rämä, T., Rebriev, Y.A., Shivas, R.G., Singh, S.M., Souza-Motta, C.M., Thangavel, R., Adhapure, N.N., Alexandrova, A.V., Alfenas, A.C., Alfenas, R.F., Alvarado, P., Alves, A.L., Andrade, D.A., Andrade, J.P., Barbosa, R.N., Barili, A., Barnes, C.W., Baseia, I.G., Bellanger, J.-M., Berlanas, C., Bessette, A.E., Bessette, A.R., Biketova, A.Y., Bomfim, F.S., Brandrud, T.E., Bransgrove, K., Brito, A.C.Q., Cano-Lira, J.F., Cantillo, T., Cavalcanti, A.D., Cheewangkoon, R., Chikowski, R.S., Conforto, C., Cordeiro, T.R.L., Craine, J.D., Cruz, R., Damm, U., de Oliveira, R.J.V., de Souza, J.T., de Souza, H.G., Dearnaley, J.D.W., Dimitrov, R.A., Dovana, F., Erhard, A., Esteve-Raventós, F., Félix, C.R., Ferisin, G., Fernandes, R.A., Ferreira, R.J., Ferro, L.O., Figueiredo, C.N., Frank, J.L., Freire, K.T.L.S., García, D., Gené, J., Gęsiorska, A., Gibertoni, T.B., Gondra, R.A.G., Gouliamova, D.E., Gramaje, D., Guard, F., Gusmão, L.F.P., Haitook, S., Hirooka, Y., Houbraken, J., Hubka, V., Inamdar, A., Iturriaga, T., Iturrieta-González, I., Jadan, M., Jiang, N., Justo, A., Kachalkin, A.V., Kapitonov, V.I., Karadelev, M., Karakehian, J., Kasuya, T., Kautmanová, I., Kruse, J., Kušan, I., Kuznetsova, T.A., Landell, M.F., Larsson, K.-H., Lee, H.B., Lima, D.X., Lira, C.R.S., Machado, A.R., Madrid, H., Magalhães, O.M.C., Majerova, H., Malysheva, E.F., Mapperson, R.R., Marbach, P.A.S., Martín, M.P., Martín-Sanz, A., Matočec, N., McTaggart, A.R., Mello, J.F., Melo, R.F.R., Mešič, A., Michereff, S.J., Miller, A.N., Minoshima, A., Molinero-Ruiz, L., Morozova, O.V., Mosoh, D., Nabe, M., Naik, R., Nara, K., Nascimento, S.S., Neves, R.P., Olariaga, I., Oliveira, R.L., Oliveira, T.G.L., Ono, T., Ordoñez, M.E., de M. Ottoni, A., Paiva, L.M., Pancorbo, F., Pant, B., Pawłowska, J., Peterson, S.W., Raudabaugh, D.B., Rodríguez-Andrade, E., Rubio, E., Rusevska, K., Santiago, A.L.C.M.A., Santos, A.C.S., Santos, C., Sazanova, N.A., Shah, S., Sharma, J., Silva, B.D.B., Siquier, J.L., Sonawane, M.S., Stchigel, A.M., Svetasheva, T., Tamakeaw, N., Telleria, M.T., Tiago, P.V., Tian, C.M., Tkalčec, Z., Tomashevskaya, M.A., Truong, H.H., Vecherskii, M.V., Visagie, C.M., Vizzini, A., Yilmaz, N., Zmitrovich, I.V., Zvyagina, E.A., Boekhout, T., Kehlet, T., Lżss¸e, T. & Groenewald, J.Z. (2019) Fungal Planet description sheets: 868–950. <em>Persoonia</em> 42: 291–473. https://doi.org/10.3767/persoonia.2019.42.11</p>
<p>Edgar, R.C. (2004) MUSCLE: multiple sequence alignment with high accuracy and high throughput. <em>Nucleic Acids Research</em> 32: 1792–1797. https://doi.org/10.1093/nar/gkh340</p>
<p>Hall, T.A. (1999) BioEdit: A user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. <em>Nucleic Acids Symposium Series</em> 41: 95–98.</p>
<p>Hashimoto, A., Hirayama, K., Takahashi, H., Matsumura, M., Okada, G., Chen, C.Y., Huang, J.W., Kakishima, M., Ono, T. & Tanaka, K. (2018) Resolving the <em>Lophiostoma bipolare</em> complex: Generic delimitations within Lophiostomataceae. <em>Studies in Mycology</em> 90: 161–189. https://doi.org/10.1016/j.simyco.2018.03.001</p>
<p>Hawksworth, D.L. & Lücking, R. (2017) Fungal diversity revisited: 2.2 to 3.8 million species. <em>The Fungal Kingdom</em>: 79–95. https://doi.org/10.1128/9781555819583.ch4</p>
<p>Hirayama, K., Hashimoto, A. & Tanaka, K. (2014) A new species, <em>Lophiostoma versicolor</em>, from Japan (Pleosporales, Dothideomycetes). <em>Mycosphere</em> 5: 411–417. https://doi.org/10.5943/mycosphere/5/3/3</p>
<p>Hirayama, K. & Tanaka, K. (2011) Taxonomic revision of <em>Lophiostoma</em> and <em>Lophiotrema</em> based on reevaluation of morphological characters and molecular analyses. <em>Mycoscience</em> 52: 401–412. https://doi.org/10.1007/S10267-011-0126-3</p>
<p>Hongsanan, S., Hyde, K.D., Phookamsak, R., Wanasinghe, D.N., McKenzie, E.H.C., Sarma, V.V., Boonmee, S., Lücking, R., Bhat, D.J., Liu, N.G., Tennakoon, D.S., Pem, D., Karunarathna, A., Jiang, S.H., Jones, E.B.G., Phillips, A.J.L., Manawasinghe, I.S., Tibpromma, S., Jayasiri, S.C., Sandamali, D.S., Jayawardena, R.S., Wijayawardene, N.N., Ekanayaka, A.H., Jeewon, R., Lu, Y.Z., Dissanayake, A.J., Zeng, X.Y., Luo, Z.L., Tian, Q., Phukhamsakda, C., Thambugala, K.M., Dai, D.Q., Chethana, K.W.T., Samarakoon, M.C., Ertz, D., Bao, D.F., Doilom, M., Liu, J.K., PérezOrtega, S., Suija, A., Senwanna, C., Wijesinghe, S.N., Konta, S., Niranjan, M., Zhang, S.N., Ariyawansa, H.A., Jiang, H.B., Zhang, J.F., Norphanphoun, C., de Silva, N.I., Thiyagaraja, V., Zhang, H., Bezerra, J.D.P., Miranda-González, R., Aptroot, A., Kashiwadani, H., Harishchandra, D., Sérusiaux, E., Aluthmuhandiram, J.V.S., Abeywickrama, P.D., Devadatha, B., Wu, H.X., Moon, K.H., Gueidan, C., Schumm, F., Bundhun, D., Mapook, A., Monkai, J., Chomnunti, P., Suetrong, S., Chaiwan, N., Dayarathne, M., Yang, J., Rathnayaka, A.R., Bhunjun, C.S., Xu, J.C., Zheng, J.S., Liu, G., Feng, Y. & Xie, N. (2020) Refined families of Dothideomycetes: Dothideomycetidae and Pleosporomycetidae. <em>Mycosphere</em> 11: 1553–2107. https://doi.org/10.5943/mycosphere/11/1/13</p>
<p>Hyde, K.D., Jeewon, R., Chen, Y.-J., Bhunjun, C.S., Calabon, M.S., Jiang, H.-B., Lin, C.-G., Norphanphoun, C., Sysouphanthong, P., Pem, D., Tibpromma, S., Zhang, Q., Doilom, M., Jayawardena, R.S., Liu, J.-K., Maharachchikumbura, S.S.N., Phukhamsakda, C., Phookamsak, R., Al-Sadi, A.M., Thongklang, N., Wang, Y., Gafforov, Y., Gareth Jones, E.B. & Lumyong, S. (2020) The numbers of fungi: is the descriptive curve flattening? <em>Fungal Diversity</em> 103: 219–271. https://doi.org/10.1007/s13225-020-00458-2</p>
<p>Hyde, K.D., Jones, E.B.G., Liu, J.K., Ariyawansa, H., Boehm, E., Boonmee, S., Braun, U., Chomnunti, P., Crous, P.W., Dai, D.Q., Diederich, P., Dissanayake, A., Doilom, M., Doveri, F., Hongsanan, S., Jayawardena, R., Lawrey, J.D., Li, Y.M., Liu, Y.X., Lücking, R., Monkai, J., Muggia, L., Nelsen, M.P., Pang, K.L., Phookamsak, R., Senanayake, I.C., Shearer, C.A., Suetrong, S., Tanaka, K., Thambugala, K.M., Wijayawardene, N.N., Wikee, S., Wu, H.X., Zhang, Y., Aguirre-Hudson, B., Alias, S.A., Aptroot, A., Bahkali, A.H., Bezerra, J.L., Bhat, D.J., Camporesi, E., Chukeatirote, E., Gueidan, C., Hawksworth, D.L., Hirayama, K., De Hoog, S., Kang, J.C., Knudsen, K., Li, W.J., Li, X.H., Liu, Z.Y., Mapook, A., McKenzie, E.H.C., Miller, A.N., Mortimer, P.E., Phillips, A.J.L., Raja, H.A., Scheuer, C., Schumm, F., Taylor, J.E., Tian, Q., Tibpromma, S., Wanasinghe, D.N., Wang, Y., Xu, J.C., Yacharoen, S., Yan, J.Y. & Zhang, M. (2013) Families of Dothideomycetes. <em>Fungal Diversity</em> 63: 1–313. https://doi.org/10.1007/s13225-013-0263-4</p>
<p>Hyde, K.D., Norphanphoun, C., Abreu, V.P., Bazzicalupo, A., Thilini Chethana, K.W., Clericuzio, M., Dayarathne, M.C., Dissanayake, A.J., Ekanayaka, A.H., He, M.-Q., Hongsanan, S., Huang, S.-K., Jayasiri, S.C., Jayawardena, R.S., Karunarathna, A., Konta, S., Kušan, I., Lee, H., Li, J., Lin, C.-G., Liu, N.-G., Lu, Y.-Z., Luo, Z.-L., Manawasinghe, I.S., Mapook, A., Perera, R.H., Phookamsak, R., Phukhamsakda, C., Siedlecki, I., Soares, A.M., Tennakoon, D.S., Tian, Q., Tibpromma, S., Wanasinghe, D.N., Xiao, Y.-P., Yang, J., Zeng, X.-Y., Abdel-Aziz, F.A., Li, W.-J., Senanayake, I.C., Shang, Q.-J., Daranagama, D.A., de Silva, N.I., Thambugala, K.M., Abdel-Wahab, M.A., Bahkali, A.H., Berbee, M.L., Boonmee, S., Bhat, D.J., Bulgakov, T.S., Buyck, B., Camporesi, E., Castañeda-Ruiz, R.F., Chomnunti, P., Doilom, M., Dovana, F., Gibertoni, T.B., Jadan, M., Jeewon, R., Jones, E.B.G., Kang, J.-C., Karunarathna, S.C., Lim, Y.W., Liu, J.-K., Liu, Z.-Y., Plautz, H.L., Lumyong, S., Maharachchikumbura, S.S.N., Matočec, N., McKenzie, E.H.C., Mešić, A., Miller, D., Pawłowska, J., Pereira, O.L., Promputtha, I., Romero, A.I., Ryvarden, L., Su, H.-Y., Suetrong, S., Tkalčec, Z., Vizzini, A., Wen, T.-C., Wisitrassameewong, K., Wrzosek, M., Xu, J.-C., Zhao, Q., Zhao, R.-L. & Mortimer, P.E. (2017) Fungal diversity notes 603–708: taxonomic and phylogenetic notes on genera and species. <em>Fungal Diversity</em> 87: 1–235. https://doi.org/10.1007/s13225-017-0391-3</p>
<p>Jayasiri, S.C., Hyde, K.D., Ariyawansa, H.A., Bhat, J., Buyck, B., Cai, L., Dai, Y.-C., Abd-Elsalam, K.A., Ertz, D., Hidayat, I., Jeewon, R., Jones, E.B.G., Bahkali, A.H., Karunarathna, S.C., Liu, J.-K., Luangsa-ard, J.J., Lumbsch, H.T., Maharachchikumbura, S.S.N., McKenzie, E.H.C., Moncalvo, J.-M., Ghobad-Nejhad, M., Nilsson, H., Pang, K.-L., Pereira, O.L., Phillips, A.J.L., Raspé, O., Rollins, A.W., Romero, A.I., Etayo, J., Selçuk, F., Stephenson, S.L., Suetrong, S., Taylor, J.E., Tsui, C.K.M., Vizzini, A., Abdel-Wahab, M.A., Wen, T.-C., Boonmee, S., Dai, D.Q., Daranagama, D.A., Dissanayake, A.J., Ekanayaka, A.H., Fryar, S.C., Hongsanan, S., Jayawardena, R.S., Li, W.-J., Perera, R.H., Phookamsak, R., de Silva, N.I., Thambugala, K.M., Tian, Q., Wijayawardene, N.N., Zhao, R.-L., Zhao, Q., Kang, J.-C. & Promputtha, I. (2015) The Faces of Fungi database: fungal names linked with morphology, phylogeny and human impacts. <em>Fungal Diversity</em> 74: 3–18. https://doi.org/10.1007/s13225-015-0351-8</p>
<p>Liu, Y.J., Whelen, S. & Hall, B.D. (1999) Phylogenetic relationships among ascomycetes: evidence from an RNA polymerse II subunit. <em>Molecular Biology and Evolution</em> 16: 1799–1808. https://doi.org/10.1093/oxfordjournals.molbev.a026092</p>
<p>Mapook, A., Hyde, K.D., McKenzie, E.H.C., Jones, E.B.G., Bhat, D.J., Jeewon, R., Stadler, M., Samarakoon, M.C., Malaithong, M., Tanunchai, B., Buscot, F., Wubet, T. & Purahong, W. (2020) Taxonomic and phylogenetic contributions to fungi associated with the invasive weed <em>Chromolaena odorata</em> (Siam weed). <em>Fungal Diversity</em> 101: 1–175. https://doi.org/10.1007/s13225-020-00444-8</p>
<p>Miller, M.A., Pfeiffer, W. & Schwartz, T. (2010) Creating the CIPRES Science Gateway for inference of large phylogenetic trees. <em>In:</em> <em>2010 Gateway Computing Environments Workshop (GCE)</em>. IEEE, pp. 1–8.</p>
<p>Minh, B.Q., Schmidt, H.A., Chernomor, O., Schrempf, D., Woodhams, M.D., von Haeseler, A. & Lanfear, R. (2020) IQ-TREE 2: New Models and Efficient Methods for Phylogenetic Inference in the Genomic Era. <em>Molecular Biology and Evolution</em> 37: 1530–1534. https://doi.org/10.1093/molbev/msaa015</p>
<p>Nguyen, L.-T., Schmidt, H.A., von Haeseler, A. & Minh, B.Q. (2015) IQ-TREE: A Fast and Effective Stochastic Algorithm for Estimating Maximum-Likelihood Phylogenies. <em>Molecular Biology and Evolution</em> 32: 268–274. https://doi.org/10.1093/molbev/msu300</p>
<p>Rambaut, A. (2012) FigTree version 1.4.0. Available from: http://tree.bio.ed.ac.uk/software/figtree/ (accessed 24 August 2022)</p>
<p>Rannala, B. & Yang, Z. (1996) Probability distribution of molecular evolutionary trees: A new method of phylogenetic inference. <em>Journal of Molecular Evolution</em> 43: 304–311. https://doi.org/10.1007/BF02338839</p>
<p>Rehner, S.A. & Buckley, E. (2005) A Beauveria phylogeny inferred from nuclear ITS and EF1-α sequences: evidence for cryptic diversification and links to <em>Cordyceps</em> teleomorphs. <em>Mycologia</em> 97: 84–98. https://doi.org/10.1080/15572536.2006.11832842</p>
<p>Rehner, S.A. & Samuels, G.J. (1994) Taxonomy and phylogeny of <em>Gliocladium</em> analysed from nuclear large subunit ribosomal DNA sequences. <em>Mycological Research</em> 98: 625–634. https://doi.org/10.1016/S0953-7562(09)80409-7</p>
<p>Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: Efficient Bayesian Phylogenetic Inference and Model Choice Across a Large Model Space. <em>Systematic Biology</em> 61: 539–542. https://doi.org/10.1093/sysbio/sys029</p>
<p>Sung, G.-H., Sung, J.-M., Hywel-Jones, N.L. & Spatafora, J.W. (2007) A multi-gene phylogeny of Clavicipitaceae (Ascomycota, Fungi): Identification of localized incongruence using a combinational bootstrap approach. <em>Molecular Phylogenetics and Evolution</em> 44: 1204–1223. https://doi.org/10.1016/j.ympev.2007.03.011</p>
<p>Swofford, D.L. (2002) PAUP: Phylogenetic analysis using parsimony. <em>Options</em> 42: 294–307. https://doi.org/10.1007/BF02198856</p>
<p>Tanaka, K. & Harada, Y. (2003) Pleosporales in Japan (1): the genus <em>Lophiostoma</em>. <em>Mycoscience</em> 44: 85–96. https://doi.org/10.1007/S10267-002-0085-9</p>
<p>Tennakoon, D.S., Kuo, C.H., Jeewon, R., Thambugala, K.M. & Hyde, K.D. (2018) Saprobic lophiostomataceae (Dothideomycetes): <em>Pseudolophiostoma mangiferae</em> sp. nov. and <em>Neovaginatispora fuckelii</em>, a new record from <em>Mangifera indica</em>. <em>Phytotaxa</em> 364: 157–171. https://doi.org/10.11646/phytotaxa.364.2.3</p>
<p>Thambugala, K.M., Hyde, K.D., Tanaka, K., Tian, Q., Wanasinghe, D.N., Ariyawansa, H.A., Jayasiri, S.C., Boonmee, S., Camporesi, E., Hashimoto, A., Hirayama, K., Schumacher, R.K., Promputtha, I. & Liu, Z.Y. (2015) Towards a natural classification and backbone tree for Lophiostomataceae, Floricolaceae, and Amorosiaceae fam. nov. <em>Fungal Diversity</em> 74: 199–266. https://doi.org/10.1007/s13225-015-0348-3</p>
<p>Vilgalys, R. & Hester, M. (1990) Rapid genetic identification and mapping of enzymatically amplified ribosomal DNA from several <em>Cryptococcus</em> species. <em>Journal of Bacteriology</em> 172: 4238–4246. https://doi.org/10.1128/jb.172.8.4238-4246.1990</p>
<p>Wanasinghe, D.N., Phukhamsakda, C., Hyde, K.D., Jeewon, R., Lee, H.B., Gareth Jones, E.B., Tibpromma, S., Tennakoon, D.S., Dissanayake, A.J., Jayasiri, S.C., Gafforov, Y., Camporesi, E., Bulgakov, T.S., Ekanayake, A.H., Perera, R.H., Samarakoon, M.C., Goonasekara, I.D., Mapook, A., Li, W.-J., Senanayake, I.C., Li, J., Norphanphoun, C., Doilom, M., Bahkali, A.H., Xu, J., Mortimer, P.E., Tibell, L., Tibell, S. & Karunarathna, S.C. (2018) Fungal diversity notes 709–839: taxonomic and phylogenetic contributions to fungal taxa with an emphasis on fungi on <em>Rosaceae</em>. <em>Fungal Diversity</em> 89: 1–236. https://doi.org/10.1007/s13225-018-0395-7</p>
<p>White, T.J., Bruns, T., Lee, S. & Taylor, J. (1990) Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics. <em>PCR Protocols and Applications</em> 18: 315–322. https://doi.org/10.1016/B978-0-12-372180-8.50042-1</p>
<p>Wijayawardene, N.N., Hyde, K.D., Al-Ani, L.K.T., Tedersoo, L., Haelewaters, D., Rajeshkumar, K.C., Zhao, R.L., Aptroot, A., Leontyev, D.V., Saxena, R.K., Tokarev, Y.S., Dai, D.Q., Letcher, P.M., Stephenson, S.L., Ertz, D., Lumbsch, H.T., Kukwa, M., Issi, I.V., Madrid, H., Phillips, A.J.L., Selbmann, L., Pfliegler, W.P., Horváth, E., Bensch, K., Kirk, P.M., Kolaríková, K., Raja, H.A., Radek, R., Papp, V., Dima, B., Ma, J., Malosso, E., Takamatsu, S., Rambold, G., Gannibal, P.B., Triebel, D., Gautam, A.K., Avasthi, S., Suetrong, S., Timdal, E., Fryar, S.C., Delgado, G., Réblová, M., Doilom, M., Dolatabadi, S., Pawlowska, J.Z., Humber, R.A., Kodsueb, R., Sánchez-Castro, I., Goto, B.T., Silva, D.K.A., de Souza, F.A., Oehl, F., da Silva, G.A., Silva, I.R., Blaszkowski, J., Jobim, K., Maia, L.C., Barbosa, F.R., Fiuza, P.O., Divakar, P.K., Shenoy, B.D., Castañeda-Ruiz, R.F., Somrithipol, S., Lateef, A.A., Karunarathna, S.C., Tibpromma, S., Mortimer, P.E., Wanasinghe, D.N., Phookamsak, R., Xu, J., Wang, Y., Tian, F., Alvarado, P., Li, D.W., Kušan, I., Matocec, N., Mešic, A., Tkalcec, Z., Maharachchikumbura, S.S.N., Papizadeh, M., Heredia, G., Wartchow, F., Bakhshi, M., Boehm, E., Youssef, N., Hustad, V.P., Lawrey, J.D., Santiago, A.L.C.M.A., Bezerra, J.D.P., Souza-Motta, C.M., Firmino, A.L., Tian, Q., Houbraken, J., Hongsanan, S., Tanaka, K., Dissanayake, A.J., Monteiro, J.S., Grossart, H.P., Suija, A., Weerakoon, G., Etayo, J., Tsurykau, A., Vázquez, V., Mungai, P., Damm, U., Li, Q.R., Zhang, H., Boonmee, S., Lu, Y.Z., Becerra, A.G., Kendrick, B., Brearley, F.Q., Motiejunaite, J., Sharma, B., Khare, R., Gaikwad, S., Wijesundara, D.S.A., Tang, L.Z., He, M.Q., Flakus, A., Rodriguez-Flakus, P., Zhurbenko, M.P., McKenzie, E.H.C., Stadler, M., Bhat, D.J., Liu, J.K., Raza, M., Jeewon, R., Nassonova, E.S., Prieto, M., Jayalal, R.G.U., Erdogdu, M., Yurkov, A., Schnittler, M., Shchepin, O.N., Novozhilov, Y.K., Silva-Filho, A.G.S., Gentekaki, E., Liu, P., Cavender, J.C., Kang, Y., Mohammad, S., Zhang, L.F., Xu, R.F., Li, Y.M., Dayarathne, M.C., Ekanayaka, A.H., Wen, T.C., Deng, C.Y., Pereira, O.L., Navathe, S., Hawksworth, D.L., Fan, X.L., Dissanayake, L.S., Kuhnert, E., Grossart, H.P. & Thines, M. (2020) Outline of Fungi and fungus-like taxa. <em>Mycosphere</em> 11: 1060–1456. https://doi.org/10.5943/mycosphere/11/1/8</p>
<p>Zhang, Y., Crous, P.W., Schoch, C.L. & Hyde, K.D. (2012) Pleosporales. <em>Fungal Diversity</em> 53: 1–221. https://doi.org/10.1007/s13225-011-0117-x</p>
<p>Zhang, Y., Wang, H.K., Fournier, J., Crous, P.W., Jeewon, R., Pointing, S.B. & Hyde, K.D. (2009) Towards a phylogenetic clarification of <em>Lophiostoma/Massarina</em> and morphologically similar genera in the Pleosporales. <em>Fungal Diversity</em> 38: 225–251.</p>